Tree retention as a conservation measure in clear-cut forests of northern Europe: A review of ecological consequences

Scandinavian Journal of Forest Research

Volumn 25, Issue 4, 2010, Pages 295-308

  Gustafsson, Lena ^a   Kouki, Jari ^b   Sverdrup Thygeson, Anne ^c  

^a SWEDISH UNIVERSITY OF AGRICULTURAL SCIENCES   (Sweden)

^b UNIVERSITY OF EASTERN FINLAND   (Finland)

^c NORWEGIAN INSTITUTE FOR NATURE RESEARCH   (Norway)

Author keywords

Biodiversity; Conservation concern; Dead wood; Green tree retention; High stump; Multiscaled; Variable retention

Indexed keywords

BIODIVERSITY CONSERVATION; CERTIFICATION STANDARDS; CONSERVATION MEASURES; DEAD WOOD; EARLY-SUCCESSIONAL; ECOLOGICAL CONSEQUENCES; FINLAND; GREEN-TREE RETENTION; LONG TERM DYNAMICS; MATURE FOREST; NATIONAL LEGISLATION; PROTECTED AREAS; SPATIAL ARRANGEMENTS; SUBSTRATE TYPES; TREE RETENTION;

BIODIVERSITY; LAWS AND LEGISLATION;

FORESTRY;

BIODIVERSITY; CONSERVATION; DEAD WOOD; FORESTRY; LEGISLATION; RETENTION;

COLEOPTERA;

EID: 77955060132     PISSN: 02827581     EISSN: 16511891     Source Type: Journal    
DOI: 10.1080/02827581.2010.497495     Document Type: Review

References (113)

1. Abrahamsson, M., Jonsell, M., Niklasson, M. & Lindbladh, M. (2009). Saproxylic beetle assemblages in artificially created high-stumps of spruce (Picea abies) and birch (Betula pendula/pubescens)-Does the surrounding landscape matter? Insect Conservation and Diversity, 2, 284-294.
2. Abrahamsson, M. & Lindbladh, M. (2006). A comparison of saproxylic beetle occurrence between man-made high- and low-stumps of spruce (Picea abies). Forest Ecology and Management, 226, 230-237.
3. Abrahamsson, M., Lindbladh, M. & Ronnberg, J. (2008). Influence of butt rot on beetle diversity in artificially created high-stumps of Norway spruce. Forest Ecology and Management, 255, 3396-3403.
4. Ahti, T., Hamet-Ahti, L. & Jalas, J. (1968). Vegetation zones and their sections in north-western Europe. Annales Zoologici Fennici, 5, 169-211.
5. Aubry, K. B., Halpern, C. B. & Maguire, D. A. (2004). Ecological effects of variable-retention harvests in the northwestern United States: The DEMO study. Forest Snow and Landscape Research, 78, 119-137.
6. Baker, S. C., Grove, S. J., Forster, L., Bonham, K. J. & Bashford, D. (2009). Short-term responses of ground-active beetles to alternative silvicultural systems in the Warra Silvicultural Systems Trial, Tasmania, Australia. Forest Ecology and Management, 258, 444-459.
7. Brunet, J. & Isacsson, G. (2009a). Influence of snag characteristics on saproxylic beetle assemblages in a south Swedish beech forest. Journal of Insect Conservation, 13, 515-528.
8. Brunet, J. & Isacsson, G. (2009b). Restoration of beech forest for saproxylic beetles-Effects of habitat fragmentation and substrate density on species diversity and distribution. Biodiversity and Conservation, 18, 2387-2404.
9. Bryant, D., Nielsen, D. & Tangley, L. (1997). The last frontier forests. Ecosystems & Economies on the edge. What is the status of the world's remaining large, natural forest ecosystems?. Washington, DC: World Resources Institute.
10. den Herder, M., Kouki, J. & Ruusila, V. (2009). The effects of timber harvest, forest fire, and herbivores on regeneration of deciduous trees in boreal pine-dominated forests. Canadian Journal of Forest Research, 39, 712-722.
11. Djupstrom, L. B., Weslien, J. & Schroeder, L. M. (2008). Dead wood and saproxylic beetles in set-aside and non set-aside forests in a boreal region. Forest Ecology and Management, 255,3340-3350.
12. Eckerberg, K. (1986). Implementation of environmental protection in Swedish forestry: A policy perspective. Forest Ecology and Management, 17, 61-72.
13. Ekbom, B., Schroeder, L. M. & Larsson, S. (2006). Stand specific occurrence of coarse woody debris in a managed boreal forest landscape in central Sweden. Forest Ecology and Management, 221, 2-12.
14. Elfving, B. & Jakobsson, R. (2006). Effects of retained trees on tree growth and field vegetation in Pinus sylvestris stands in Sweden. Scandinavian Journal of Forest Research, 21, 29-36.
15. Esseen, P. A. (1994). Tree mortality patterns after experimental fragmentation of an old-growth conifer forest. Biological Conservation, 68, 19-28.
16. Esseen, P.-A., Ehnstrom, B., Ericson, L. & Sjoberg, K. (1997). Boreal forests. Ecological Bulletins, 46, 16-47.
17. FAO (2006). Global Forest Resources Assessment 2005. Progress towards sustainable forest management (No. 147). Rome. Food and Agriculture Organization of the United Nations.
18. Fossestøl, K. O. & Sverdrup-Thygeson, A. (2009). Saproxylic beetles in high stumps and residual downed wood on clearcuts and in forest edges. Scandinavian Journal of Forest Research, 24, 403-416.
19. Franklin, J. F., Berg, D. R., Thornburgh, D. A. & Tappeiner, J. C. (1997). Alternative silvicultural approaches to timber harvesting: Variable retention systems. In K. A. Kohm, & J. F. Franklin (Eds.), Creating a forestry for the 21st century. The science of forest management (pp. 111-139). Washington, DC: Island Press.
20. Fraver, S., Jonsson, B. G., Jonsson, M. & Esseen, P. A. (2008). Demographics and disturbance history of a boreal old-growth Picea abies forest. Journal of Vegetation Science, 19, 789-798.
21. Fridman, J. (2000). Conservation of forest in Sweden: A strategic ecological analysis. Biological Conservation, 96, 95-103.
22. Gardenfors, U. (2010). Rodlistade arter i Sverige 2005-The 2010 Redlist of Swedish species. Uppsala: ArtDatabanken, SLU.
23. Gibb, H., Pettersson, R. B., Hjalten, J., Hilszczanski, J., Ball, J. P., Johansson, T., Atlegrim, O. & Danell, K. (2006). Conservation-oriented forestry and early successional saproxylic beetles: Responses of functional groups to manipulated dead wood substrates. Biological Conservation, 129, 437-450.
24. Hågvar, S., Nygaard, P. & Bækken, B. T. (2004). Retention of forest strips for bird-life adjacent to water and bogs in Norway: Effect of different widths and habitat variables. Scandinavian Journal of Forest Research, 19, 452-465.
25. Hannerz, M. & Hånell, B. (1997). Effects on the flora in Norway spruce forests following clearcutting and shelterwood cutting. Forest Ecology and Management, 90, 29-49.
26. Hautala, H., Jalonen, J., Laaka-Lindberg, S. & Vanha-Majamaa, I. (2004). Impacts of retention felling on coarse woody debris (CWD) in mature boreal spruce forests in Finland. Biodiversity and Conservation, 13, 1541-1554.
27. Hautala, H. & Vanha-Majamaa, I. (2006). Immediate tree uprooting after retention-felling in a coniferous boreal forest in Fennoscandia. Canadian Journal of Forest Research, 36, 3167-3172.
28. Hazell, P. & Gustafsson, L. (1999). Retention of trees at final harvest-Evaluation of a conservation technique using epiphytic bryophyte and lichen transplants. Biological Conservation, 90, 133-142.
29. Hedenås, H., Blomberg, P. & Ericson, L. (2007). Significance of old aspen (Populus tremula) trees for the occurrence of lichen photobionts. Biological Conservation, 135, 380-387.
30. Hedenås, H. & Ericson, L. (2003). Response of epiphytic lichens on Populus tremula in a selective cutting experiment. Ecological Applications, 13, 1124-1134.
31. Hedenås, H. & Hedstrom, P. (2007). Conservation of epiphytic lichens: Significance of remnant aspen (Populus tremula) trees in clear-cuts. Biological Conservation, 135, 388-395.
32. Hedgren, P. O. (2007). Early arriving saproxylic beetles (Coleoptera) and parasitoids (Hymenoptera) in low and high stumps of Norway spruce. Forest Ecology and Management, 241, 155-161.
33. Hjalten, J., Johansson, T., Alinvi, O., Danell, K., Ball, J. P., Pettersson, R., et al. (2007). The importance of substrate type, shading and scorching for the attractiveness of dead wood to saproxylic beetles. Basic and Applied Ecology, 8, 364-376.
34. Hofgaard, A. (1993). 50 years of change in a Swedish boreal old-growth Picea abies forest. Journal of Vegetation Science, 4, 773-782.
35. Hylander, K. (2005). Aspect modifies the magnitude of edge effects on bryophyte growth in boreal forests. Journal of Applied Ecology, 42, 518-525.
36. Hylander, K. & Dynesius, M. (2006). Causes of the large variation in bryophyte species richness and composition among boreal streamside forests. Journal of Vegetation Science, 17, 333-346.
37. Hylander, K., Dynesius, M., Jonsson, B. G. & Nilsson, C. (2005). Substrate form determines the fate of bryophytes in riparian buffer strips. Ecological Applications, 15, 674-688.
38. Hylander, K., Jonsson, B. G. & Nilsson, C. (2002). Evaluating buffer strips along boreal streams using bryophytes as indicators. Ecological Applications, 12, 797-806.
39. Hylander, K., Nilsson, C. & Gothner, T. (2004). Effects of bufferstrip retention and clearcutting on land snails in boreal riparian forests. Conservation Biology, 18, 1052-1062.
40. Hyvärinen, E., Kouki, J., Martikainen, P. & Lappalainen, H. (2005). Short-term effects of controlled burning and green-tree retention on beetle (Coleoptera) assemblages in managed boreal forests. Forest Ecology and Management, 212, 315-332.
41. Hyvarinen, E., Kouki, J. & Martikainen, P. (2006). Fire and green-tree retention in conservation of red-listed and rare deadwood-dependent beetles in Finnish boreal forests. Con-servation Biology, 20, 1711-1719.
42. Jacks, G. & Norrstrom, A. C. (2004). Hydrochemistry and hydrology of forest riparian wetlands. Forest Ecology and Management, 196, 187-197.
43. Jakobsson, R. & Elfving, B. (2004). Development of an 80-year-old mixed stand with retained Pinus sylvestris in northern Sweden. Forest Ecology and Management, 194, 249-258.
44. Jalonen, J. & Vanha-Majamaa, I. (2001). Immediate effects of four different fellings methods on mature boreal spruce forest understorey vegetation in southern Finland. Forest Ecology and Management, 146, 25-34.
45. Johansson, T., Gibb, H., Hilszczanski, J., Pettersson, R. B., Hjalten, J., Atlegrim, O., et al. (2006). Conservation-oriented manipulations of coarse woody debris affect its value as habitat for spruce-infesting bark and ambrosia beetles (Coleoptera: Scolytinae) in northern Sweden. Canadian Journal of Forest Research, 36, 174-185.
46. Jonsell, M., Nitterus, K. & Stighall, K. (2004). Saproxylic beetles in natural and man-made deciduous high stumps retained for conservation. Biological Conservation, 118, 163-173.
47. Jonsell, M., Schroeder, M. & Weslien, J. (2005). Saproxylic beetles in high stumps of spruce: Fungal flora important for determining the species composition. Scandinavian Journal of Forest Research, 20, 54-62.
48. Jonsell, M. & Weslien, J. (2003). Felled or standing retained wood-It makes a difference for saproxylic beetles. Forest Ecology and Management, 175, 425-435.
49. Jonsson, M., Ranius, T., Ekvall, H. & Bostedt, G. (2010). Cost-effectiveness of silvicultural measures to increase substrate availability for wood-dwelling species: A comparison among boreal tree species. Scandinavian Journal of Forest Research, 25,46-60.
50. Jonsson, M., Ranius, T., Ekvall, H., Bostedt, G., Dahlberg, A., Ehnstrom, B., et al. (2006). Cost-effectiveness of silvicultural measures to increase substrate availability for red-listed wood-living organisms in Norway spruce forests. Biological Conservation, 127, 443-462.
51. Jonsson, M. T., Fraver, S. & Jonsson, B. G. (2009). Forest history and the development of old-growth characteristics in fragmented boreal forests. Journal of Vegetation Science, 20, 91-106.
52. Jonsson, M.T., Fraver, S., Jonsson, B. G., Dynesius, M., Rydgard, M. & Esseen, P. A. (2007). Eighteen years of tree mortality and structural change in an experimentally fragmented Norway spruce forest. Forest Ecology and Management, 242, 306-313.
53. Junninen, K., Penttilä, R. & Martikainen, P. (2007). Fallen retention aspen trees on clear-cuts can be important habitats for red-listed polypores: A case study in Finland. Biodiversity and Conservation, 16, 475-490.
54. Kaila, L., Martikainen, P. & Punttila, P. (1997). Dead trees left in clear-cuts benefit saproxylic Coleoptera adapted to natural disturbances in boreal forest. Biodiversity and Conservation, 6, 1-18.
55. Kålås, J. A., Viken, Å. & Bakken, T. (2006). Norwegian Red List pp. 416. Trondheim: Norwegian Biodiversity Information Centre. (In Norwegian and English.).
56. Koivula, M. (2002). Alternative harvesting methods and boreal carabid beetles (Coleoptera, Carabidae). Forest Ecology and Management, 167, 103-121.
57. Kouki, J., Arnold, K. & Martikainen, P. (2004). Long-term persistence of aspen-a key host for many threatened species-is endangered in old-growth conservation areas in Finland. Journal for Nature Conservation, 12, 41-52.
58. Kouki, J., Lofman, S., Martikainen, P., Rouvinen, S. & Uotila, A. (2001). Forest fragmentation in Fennoscandia: Linking habitat requirements of wood-associated threatened species to landscape and habitat changes. Scandinavian Journal of Forest Research, 16 (Suppl. 3), 27-37.
59. Kuuluvainen, T. (2009). Forest management and biodiversity conservation based on natural ecosystem dynamics in northern Europe. The complexity challenge. AMBIO, 38, 309-315.
60. Langvall, O. & Ottosson Lofvenius, M. (2002). Effect of shelter-wood density on nocturnal near-ground temperature, frost injury risk and budburst day of Norway spruce. Forest Ecology and Management, 168, 149-161.
61. Lauren, A., Finer, L., Koivusalo, H., Kokkonen, T., Karvonen, T., Kellomadki, S., et al. (2005). Water and nitrogen processes along a typical water flowpath and streamwater exports from a forested catchment and changes after clear-cutting: A modelling study. Hydrology and Earth System Sciences, 9, 657-674.
62. Lie, M. H., Arup, U., Grytnes, J. A. & Ohlson, M. (2009). The importance of host tree age, size and growth rate as determinants of epiphytic lichen diversity in boreal spruce forests. Biodiversity and Conservation, 18, 3579-3596.
63. Lindbladh, M., Abrahamsson, M., Seedre, M. & Jonsell, M. (2007). Saproxylic beetles in artificially created high-stumps of spruce and birch within and outside hotspot areas. Biodiversity and Conservation, 16, 3213-3226.
64. Lindenmayer, D. B. & Franklin, J. F. (2003). Towards forest sustainability. Collingwood: CSIRO Publishing.
65. Lindenmayer, D. B., Franklin, J. F. & Fischer, J. (2006). General management principles and a checklist of strategies to guide forest biodiversity conservation. Biological Conservation, 131, 433-445.
66. Lindhe, A., Asenblad, N. & Toresson, H. G. (2004). Cut logs and high stumps of spruce, birch, aspen and oak-Nine years of saproxylic fungi succession. Biological Conservation, 119, 443-454.
67. Lindhe, A. & Lindelow, Å. (2004). Cut high stumps of spruce, birch, aspen and oak as breeding substrates for saproxylic beetles. Forest Ecology and Management, 203, 1-20.
68. Lindhe, A., Lindelow, Å. & Åsenblad, N. (2005). Saproxylic beetles in standing dead wood density in relation to substrate sun-exposure and diameter. Biodiversity and Conservation, 14, 3033-3053.
69. Lofgren, S., Ring, E., von Bromssen, C., Sørensen, R. & Hogbom, L. (2009). Short-term effects of clear-cutting on the water chemistry of two boreal streams in northern Sweden: A paired catchment study. AMBIO, 38, 347-356.
70. McGeoch, M. A., Schroeder, M., Ekbom, B. & Larsson, S. (2007). Saproxylic beetle diversity in a managed boreal forest: Importance of stand characteristics and forestry conservation measures. Diversity and Distributions, 13, 418-429.
71. Martikainen, P. (2001). Conservation of threatened saproxylic beetles: Significance of retained aspen Populus tremula on clearcut areas. Ecological Bulletins, 49, 205-218.
72. Martikainen, P., Kouki, J. & Heikkala, O. (2006a). The effects of green tree retention and subsequent prescribed burning on ground beetles (Coleoptera: Carabidae) in boreal pine-dominated forests. Ecography, 29, 659-670.
73. Martikainen, P., Kouki, J., Heikkala, O., Hyvarinen, E. & Lappalainen, H. (2006b). Effects of green tree retention and prescribed burning on the crown damage caused by the pine shoot beetles (Tomicus spp.) in pine-dominated timber harvest areas. Journal of Applied Entomology, 130, 37-44.
74. Martinez Pastur, G., Lencinas, M. V., Cellini, J. M., Peri, P. L. & Esteban, R. S. (2009). Timber management with variable retention in Nothofagus pumilio forests of southern Patagonia. Forest Ecology and Management, 258, 436-443.
75. Mather, A. (1992). The forest transition. Area, 24, 367-379.
76. Matveinen-Huju, K., Koivula, M., Niemela, J. & Rauha, A. M. (2009). Short-term effects of retention felling at mire sites on boreal spiders and carabid beetles. Forest Ecology and Management, 258, 2388-2398.
77. Matveinen-Huju, K., Niemela, J., Rita, H. & O'Hara, R. B. (2006). Retention-tree groups in clear-cuts: Do they constitute "life-boats" for spiders and carabids? Forest Ecology and Management, 230, 119-135.
78. MCPFE. (2007). State of Europe's forests 2007. The MCPFE report on sustainable forest management in Europe.Warsaw: MCPFE.
79. Millennium Ecosystem Assessment (2005). Ecosystems and human well-being: Biodiversity synthesis. Washington, DC: World Resources Institute.
80. Monkkonen, M. & Mutanen, M. (2003). Occurrence of moths in boreal forest corridors. Conservation Biology, 17, 468-475.
81. Perhans, K., Appelgren, L., Jonsson, F., Nordin, U., Soderstrom, B. & Gustafsson, L. (2009). Retention patches as potential refugia for bryophytes and lichens in managed forest land-scapes. Biological Conservation, 142, 1125-1133.
82. Perhans, K., Gustafsson, L., Jonsson, F., Nordin, U. & Weibull, H. (2007). Bryophytes and lichens in different types of forest set-asides in boreal Sweden. Forest Ecology and Management, 242,374-390.
83. Pitkänen, A., Kouki, J., Viiri, H. & Martikainen, P. (2008). Effects of controlled forest burning and intensity of timber harvesting on the occurrence of pine weevils, Hylobius spp., in regeneration areas. Forest Ecology and Management, 255, 522-529.
84. Pitkänen, A., Tormanen, K., Kouki, J., Järvinen, E. & Viiri, H. (2005). Effects of green tree retention, prescribed burning and soil treatment on pine weevil (Hylobius abietis and Hylobius pinastri) damage to planted Scots pine seedlings. Agricultural and Forest Entomology, 7, 1-12.
85. Pullin, A. S. & Stewart, G. B. (2006). Guidelines for systematic review in conservation and environmental management. Conservation Biology, 20, 1647-1656.
86. Ranius, T., Ekvall, H., Jonsson, M. & Bostedt, G. (2005). Cost-efficiency of measures to increase the amount of coarse woody debris in managed Norway spruce forests. Forest Ecology and Management, 206, 119-133.
87. Ranius, T. & Kindvall, O. (2004). Modelling the amount of coarse woody debris produced by the new biodiversity-oriented silvicultural practices in Sweden. Biological Conservation, 119, 51-59.
88. Ranius, T., Kindvall, O., Kruys, N. & Jonsson, B. G. (2003). Modelling dead wood in Norway spruce stands subject to different management regimes. Forest Ecology and Management, 182, 13-29.
89. Rassi, P., Alanen, A., Kanerva, T. & Mannerkoski, I. (2001). Suomen lajien uhanalaisuus 2000 [Threatened species in Finland in 2000] pp. 432. Helsinki: Ymparistoministerio & Suomen Ymparistokeskus.
90. Rosenvald, R. & Lõmus, A. (2008). For what, when, and where is green-tree retention better than clear-cutting? A review of the biodiversity aspects. Biological Conservation, 255, 1-15.
91. Sahlin, E. & Ranius, T. (2009). Habitat availability in forests and clearcuts for saproxylic beetles associated with aspen. Biodiversity and Conservation, 18, 621-638.
92. Schmitt, C. B., Burgess, N. D., Coad, L., Belokurov, A., Besançon, C., Boisrobert, L., et al. (2009). Global analysis of the protection status of the world's forests. Biological Conservation, 142, 2122-2130.
93. Schroeder, L. M., Ranius, T., Ekbom, B. & Larsson, S. (2006). Recruitment of saproxylic beetles in high stumps created for maintaining biodiversity in a boreal forest landscape. Canadian Journal of Forest Research, 36, 2168-2178.
94. Schroeder, L. M., Weslien, J., Lindelow, Å. & Lindhe, A. (1999). Attacks by bark-and wood-boring Coleoptera on mechanically created high stumps of Norway spruce in the two years following cutting. Forest Ecology and Management, 123, 21-30.
95. Siira-Pietikainen, A. & Haimi, J. (2009). Changes in soil fauna 10 years after forest harvestings: Comparison between clear felling and green-tree retention methods. Forest Ecology and Management, 258, 332-338.
96. Siira-Pietikainen, A., Pietikainen, J., Fritze, H. & Haimi, J. (2001). Short-term responses of soil decomposer communities to forest management: Clear felling versus alternative forest harvesting methods. Canadian Journal of Forest Research, 31, 88-99.
97. Siira-Pietikainen, A., Haimi, J. & Siitonen, J. (2003). Short-term responses of soil macroarthropod community to clear felling and alternative forest regeneration methods. Forest Ecology and Management, 172, 339-353.
98. Siitonen, J. (2001). Forest management, coarse woody debris and saproxylic organisms: Fennoscandian boreal forest as an example. Ecological Bulletins, 49, 11-41.
99. Silvennoinen, H., Pukkala, T. & Tahvanainen, L. (2002). Effect of cuttings on the scenic beauty of a tree stand. Scandinavian Journal of Forest Research, 17, 263-273.
100. Sippola, A. L., Siitonen, J. & Punttila, P. (2002). Beetle diversity in timberline forests: A comparison between old-growth and regeneration areas in Finnish Lapland. Annales Zoologici Fennici, 39, 69-86.
101. Soderstrom, B. (2009). Effects of different levels of green-and dead-tree retention on hemi-boreal forest bird communities in Sweden. Forest Ecology and Management, 257, 215-222.
102. Stokland, J. N., Eriksen, R., Tomter, S. M., Korhonen, K. E. T., Rajaniemi, S., Soderberg, U., et al. (2003). Forest biodiversity indicators in the Nordic countries. Status based on national forest inventories, pp. 514 (TemaNord 2003). Copenhagen: Nordic Council of Ministers.
103. Sverdrup-Thygeson, A. & Birkemoe, T. (2009). What window traps can tell us: Effect of placement, forest openness and beetle reproduction in retention trees. Journal of Insect Conservation, 13, 183-191.
104. Sverdrup-Thygeson, A. & Ims, R. A. (2002). The effect of forest clearcutting in Norway on the community of saproxylic beetles on aspen. Biological Conservation, 106, 347-357.
105. Tikkanen, O.-P., Heinonen, T., Kouki, J. & Matero, J. (2007). Habitat suitability models of saproxylic red-listed boreal forest species in long-term matrix management: Cost-effective measures for multi-species conservation. Biological Conservation, 140, 359-372.
106. Timonen, J., Siitonen, J., Gustafsson, L., Kotiaho, J. S., Stokland, J. N., Sverdrup-Thygeson, A. & Monkkonen, M. (2010). Woodland key habitats in northern Europe: Concepts, inventory and protection. Scandinavian Journal of Forest Research, 25, 309-324.
107. Tonnes, S., Karjalainen, E., Lofstrom, I. & Neuvonen, M. (2004). Scenic impacts of retention trees in clear-cutting areas. Scandinavian Journal of Forest Research, 19, 348-357.
108. Uliczka, H. (2003). Nature conservation efforts by forest own-ers-Intentions and practice in a Swedish case study. Silva Fennica, 37, 459-475.
109. Vanha-Majamaa, I. & Jalonen, J. (2001). Green tree retention in Fennoscandian forestry. Scandinavian Journal of Forest Research, 16(Suppl. 3), 79-90.
110. Virkkala, R. & Rajasärkka, A. (2007). Uneven regional distribution of protected areas in Finland: Consequences for boreal forest bird populations. Biological Conservation, 134, 361-371.
111. Wikars, L. O., Sahlin, E. & Ranius, T. (2005). A comparison of three methods to estimate species richness of saproxylic beetles (Coleoptera) in logs and high stumps of Norway spruce. Canadian Entomologist, 137, 304-324.
112. Wikberg, S., Perhans, K., Kindstrand, C., Djupstrom, L. B., Boman, M., Mattsson, L., et al. (2009). Cost-effectiveness of conservation strategies implemented in boreal forests: The area selection process. Biological Conservation, 142, 614-624.
113. Work, T. T., Spence, J. R., Volney, W. J. A., Morgantini, L. E. & Innes, J. L. (2003). Integrating biodiversity and forestry practices in western Canada. Forestry Chronicle, 79, 906-916.