Redescription and reassessment of the phylogenetic affinities of euhelopus zdanskyi (dinosauria: Sauropoda) from the early cretaceous of China

Journal of Systematic Palaeontology

Volumn 7, Issue 2, 2009, Pages 199-239

  Wilson, Jeffrey A ^a   Upchurch, Paul ^b  

^a UNIVERSITY OF MICHIGAN   (United States)

^b UNIVERSITY COLLEGE LONDON   (United Kingdom)

Author keywords

Asia; Palaeobiogeography; Palaeontology; Phylogeny; Sauropod

Indexed keywords

ANATOMY; CRANIUM; CRETACEOUS; EXTINCTION; GEOGRAPHICAL DISTRIBUTION; JURASSIC; PALEOBIOGEOGRAPHY; PHYLOGENETICS; PHYLOGENY; REPTILE; VERTEBRATE;

ASIA; CHINA; EURASIA; FAR EAST;

DINOSAURIA; EUHELOPODIDAE; MAMENCHISAURUS; SAUROPODA; TITANOSAURIA; TITANOSAURIFORMES;

EID: 67650273647     PISSN: 14772019     EISSN: 14780941     Source Type: Journal    
DOI: 10.1017/S1477201908002691     Document Type: Article

References (197)

1. Allain, R., Tarquet, P., Battail, B., Dejax, J., Richir, P., Veran, M., Limon-Duparcmeur, F., Vacant, R., Mateus, O., Sayarath, P., Khenthavong, B. & Phouyavong, S. 1999. Un nouveau genere de dinosaure sauropode de la formation des Orés supérieurs (Aptien- Albien) du Laos. Comptes Rendus de I'Academie des Sciences Paris, Sciences de la Terre et des Planetes 329: 609-616.
2. Alexander, J. H. 1857. Universal dictionary of weights and measures, ancient and modern; reduced to the standards ofthe United States of America. W. Minifie, Baltimore, 158 pp.
3. Andrews, R. C. 1932. The New Conquest of Central Asia. A Narrative of the Explorations ofthe Central Asiatic Expeditions in Mongolia and China, 1921-1930. The American Museum of Natural History, New York, 678 pp.
4. Astibia, H., Buffetaut, E., Buscalioni, A. D., Cappetta, H., Corral, C., Estes, R., Garcia-Garmilla, F., Jaeger, J. J., Jimenez-Fuentes, E., Le Loeuff, J., Mazin, J. M., Orue-Etxebarria, X., Pereda-Suberbiola, J., Powell, J. E., Rage, J.-C., Rodriguez-Lazaro, J., Sanz, J. L. & Tong, H. 1990. The fossil vertebrates of Lano (Basque Country, Spain); new evidence on the composition and affinities of the Late Cretaceous continental faunas of Europe. Terra Nova 2: 460-466.
5. Averianov, A. O. & Skutschas, P. 2000. A eutherian mammal from the Early Cretaceous of Russia and biostratigraphy of the Asian Early Cretaceous vertebrate assemblages. Lethaia 33: 330-340.
6. -, Voronkevich, A. V., Maschenko, E. N., Leshchinskiy, S. V. & Faygertz, A. V. 2002. A sauropod foot from the Early Cretaceous of Western Siberia, Russia. Acta Palaeontologica Polonica 47: 117-124.
7. Azuma, Y. 2003. Early Cretaceous vertebrate remains from Katsuyama City, Fukui Prefecture, Japan. Memoir of the Fukui Prefectural Dinosaur Museum 2: 17-21.
8. Barrett, P. M. & Batten, D. J. 2007. Evolution and Palaeobiology of Early Sauropodomorph Dinosaurs. Special Papers in Palaeontology 77: 1-288.
9. - & Wang, X.-L. 2007. Basal titanosauriform (Dinosauria, Sauro- poda) teeth from the Lower Cretaceous Yixian Formation of Liaoning Province, China. Palaeoworld 16: 265-271.
10. -, Hasegawa, Y., Manabe, M., Isaji, S. & Matsouka, H. 2002. Sauropod dinosaurs from the Lower Cretaceous of eastern Asia: taxonomic and biogeographical implications. Palaeontology 45: 1197-1217.
11. Benson, R. 2008. New information on Stokesosaurus, a tyrannosauroid (Dinosauria: Theropoda) from North America and the United Kingdom. Journal of Vertebrate Paleontology 28: 732-750.
12. Blakey, R.C. 2006. Olobal paleogeographic views of earth history - late Precambrian to Recent. Available at: http://jan.ucc.nau.edu/~rcb/ globaltext2.html.
13. Bohlin, B. 1953. Reports from the Scientific Expedition to the Northwestern Provinces of China Under Leadership of Dr. Sven Hedin. VI. Vertebrate Palaeontology 6: Fossil Reptiles from Mongolia and Kansu. Statens Etnografiska Museum, Stockholm, 113 pp.
14. Bonnan, M. 2000. The presence of a calcaneum in a diplodocid sauropod. Journal of Vertebrate Paleontology 20: 317-323.
15. Bonaparte, J. F. 1984. Late Cretaceous faunal interchange of terrestrial vertebrates between the Americas. Pp. 19-24 in Reif,W.E.& Westphal, F. (eds) Third Symposium on Mesozoic Terrestrial Ecosystems, Short Papers. Attempto Verlag, Tübingen.
16. - 1986. The early radiation and phylogenetic relationships of sauropod dinosaurs, based on vertebral anatomy. Pp. 247-258 in Padian, K. (ed.) The Beginning of the Age of Dinosaurs. Cambridge University Press, Cambridge.
17. - 1999. Tetrapod faunas from South America and India: a palaeobiogeo- graphic interpretation. Proceedings of the Indian National Science Academy 65 A: 427-437.
18. - & Kielan-Jaworowska, Z. 1987. Late Cretaceous dinosaur and mammal faunas of Laurasia and Oondwana. Pp. 22-29 in Currie, P. & Koster, E. H. (eds) Fourth Symposium on Mesozoic Terrestrial Ecosystems, Short Papers. Tyrell Museum of Paleontology, Drumheller.
19. Borsuk-Bialynicka, M. 1977. A new camarasaurid sauropod Opis- thocoelicaudia skarzynskii gen. n., sp. n. from the Upper Cretaceous of Mongolia. Palaeontologica Polonica 37: 6-63.
20. Britt, B. B. 1993. Pneumatic postcranial bones in dinosaurs and other archosaurs. Unpublished Ph.D. Dissertation: University of Calgary, Alberta.
21. Brooks, D. R. & McLennan, D. A. 2002. The Nature ofDiversity: an Evolutionary Voyage ofDiscovery. University of Chicago Press, Chicago, 676 pp.
22. Buffetaut, E. & Suteethorn, V. 1999. The dinosaur fauna of the Sao Khua Formation of Thailand and the beginning of the Cretaceous radiation of dinosaurs in Asia. Palaeogeography, Palaeoclimatology, Palaeoecology 150: 13-23.
23. Buffetaut, E., Suteethorn, V., Tong, H.-Y., Chaimanee, Y. & Khansubha, S. 1997. New dinosaur discoveries in the Jurassic and Cretaceous of northeastern Thailand. Pp. 177-187 in Dheeradilok, P. (ed.) International Conference on Stratigraphy and Tectonic Evolution of Southeast Asia and the South Pacific, Department of Mineral Resources, Bangkok.
24. Buffetaut, E., Suteethorn, V., Le Loeuff, J., Cuny, G., Tong, H. & Khansubha, S. 2002. A review of the sauropod dinosaurs of Thailand. Pp. 95-101 in Mantajit, N. (ed.) The Symposium on Geology of Thailand. Department of Mineral Resources, Bangkok.
25. Calvo, J. O. & Salgado, L. 1995. Rebbachisaurus tessonei sp. nov. a new Sauropoda from the Albian-Cenomanian of Argentina; new evidence on the origin of the Diplodocidae. Gaia 11: 13-33.
26. Canudo, J. I. & Royo-Torres, R. 2004. El fémur saurópodo (Dinosauria) de Peñaroya de Tastavins (Teruel, Aptiense inferior, Cretácico inferior): Implicaciones paleobiogeograf́icas para los Titanosauriformes. Geo-Temas 6: 19-21.
27. -, Ruiz-Omeñaca, J. I., Barco, J. L. & Royo-Torres, R. 2002. iSaurópodos asiáticos en el Barremiense inferior (Cretácico Inferior) de Espafĩa? Ameghiniana 39: 443-452.
28. Chen, P., Li, W., Chen, J., Ye, C., Wang, Z., Shen, Y. & Sun, D. 1982. Stratigraphical classification of Jurassic and Cretaceous in China. Sci- entia Sinica (series B) 25: 1227-1248.
29. Cogné, J.-P., Kravchinsky, V. A., Halim, N. & Hankard, F. 2005. Late Jurassic-Early Cretaceous closure of the Mongol-Okhotsk Ocean demonstrated by new Mesozoic palaeomagnetic results from the Trans- Baikal area (SE Siberia). Geophysical Journal International 163: 813-832.
30. Croizat, L., Nelson, G. & Rosen, D. E. 1974. Centers of origin and related concepts. Systematic Zoology 23: 265-287.
31. Currie, P. J. 1995. Wandering dragons, the dinosaurs of Canada and China. Pp.101 in Sun, A. & Wang, Y. (eds) Sixth Symposium on Terrestrial Mesozoic Ecosystems and Biotas, Short Papers. China Ocean Press, Beijing.
32. Curry Rogers, K. A. 2005. Titanosauria: a phylogenetic overview. Pp. 50-103 in Curry Rogers, K. A. & Wilson, J. A. (eds) The Saur-opods: Evolution and Paleobiology. University of California Press, Berkeley and Los Angeles.
33. Day, J. J., Upchurch, P., Norman, D. B., Gale, A. S. & Powell, H. P. 2002. Sauropod trackways, evolution and behavior. Science 296: 1659.
34. -, Norman, D. B., Gale, A. S., Upchurch, P. & Powell, H. P. 2004. New Middle Jurassic dinosaur trackways from Oxfordshire, U.K. Palaeontology 47: 319-348.
35. Depéret, C. 1896. Note sur les Dinosauriens Sauropodes & Théropodes du Crétacé Supérieur de Madagascar. Bulletin de la Société Géologique de France 24: 176-194.
36. Dong, Z.-M. 1977. [On the dinosaurian remains from Turpan, Xinjiang]. Vertebrata Palasiatica 15: 59-65. [In Chinese with English abstract]
37. - 1992. Dinosaurs of China. China Ocean Press, Beijing, 188 pp.
38. - 1995. The dinosaur complexes of China and their biochronology. Pp. 91-96 in Sun, A. & Wang, Y. (eds) Sixth Symposium ofMeso- zoic Terrestrial Ecosystems and Biota, Short Papers. China Ocean Press, Beijing.
39. - 1997. On the sauropods from the Mazongshan area, Oansu Province, China. Pp. 19-23 in Dong, Z. (ed.) Sino-Japanese Silk Road Dinosaur Expedition. China Ocean Press, Beijing.
40. - 1998. On Asiatic sauropods and their phylogenetic relationships. Pp. 79-88 in Oalopim, A. M. C., Andrade, A. M. & Silva, C. M. (eds) I Encontro Internacional sobre Paleobiologia dos Dinossaáurios. Lisbon.
41. -, Zhou, S. & Zhang, Y. 1983. The dinosaurian remains from Sichuan Basin, China. Palaeontologia Sinica (New Series C) 23: 1-145.
42. -,Paik, I.S. &Kim, H.J. 2001. A preliminary report on a sauropod from the Hasandong Formation (Lower Cretaceous), Korea. Pp. 41-53 inDeng, T. & Wang, Y. (eds) Proceedings ofthe Eighth Annual Meeting ofthe Chinese Society ofVertebrate Paleontology. China Ocean Press, Beijing.
43. Enkin, R. J., Zhentu, Y., Yan, C. & Courtillot, V. 1992. Paleomagnetic constraints on the geodynamic history of the major blocks of China from the Permian to the present. Journal ofGeophysical Research 97: 13953-13989.
44. Falconer, H. 1868. Notes on fossil remains found in the Valley Indus below Attok, and at Jubbulpoor. Pp. 414-419 in Murchinson, C. (ed.) Pa- laeontological Memoirs and Notes ofthe Late Hugh Falconer. Robert Hardwicke, London.
45. Gauthier, J. A. 1986. Saurischian monophyly and the origin of birds. Pp. 1-55 in Padian, K. (ed.) The Origin ofBirds and the Evolution of Flight. Memoirs of the California Academy of Sciences 8. California Academy of Sciences, San Francisco.
46. Gilmore, C. W. 1933. Two new dinosaurian reptiles from Mongolia with notes on some fragmentary specimens. American Museum Novitates 679: 1-20.
47. - 1936. Osteology ofApatosaurus with special reference to specimens in the Carnegie Museum. Memoirs ofthe Carnegie Museum 11: 175-300.
48. Granger, W. & Gregory, W. K. 1923. Protoceratops andrewsi,apre- ceratopsian dinosaur from Mongolia. American Museum Novitates 72: 1-9.
49. Grant, T., Faivovich, J. & Pol, D. 2003. The perils of 'point-and-click' systematics. Cladistics 19: 276-285.
50. Halas, D., Zampar, D. & Brooks, D. R. 2005. A historical biogeograph- ical protocol for studying biotic diversification by taxon pulses. Journal ofBiogeography 32: 249-260.
51. Harris, J. D. 2006. The significance of Suuwassea emiliae (Dino- sauria: Sauropoda) for flagellicaudatan interrelationships and evolution. Journal ofSystematic Palaeontology 4: 185-198.
52. He, H.-Y., Wang, X.-L., Zhou, Z.-H., Wang, F., Boven, A., Shi, G.-H. & Zhu, R.-X. 2004. Timing of the Jiufotang Formation (Jehol Oroup) in Liaoning, northeastern China, and its implications. Geophysical Research Letters 31, L12605.
53. He, X., Li, K. &Cai, K. 1988. [The Middle Jurassic Dinosaur Fauna from Dashanpu, Zigong, Sichuan, Vol. IV. Sauropod Dinosaurs (2), Omeisaurus Tianfuensis]. Sichuan Scientific and Technological Publishing House, Chengdu, 144 pp. [In Chinese with English abstract].
54. Holtz Jr., T. R., Chapman, R. E. & Lamanna, M. C. 2004. Meso- zoic biogeography of Dinosauria. Pp. 627-642 in Weishampel, D. B., Dodson, P. & Osmolska, H. (eds) The Dinosauria, 2nd Edition.Uni- versity of California Press, Berkeley.
55. Hou, L.-H., Yeh, H.-K. & Zhao, X.-J. 1975. [Fossil reptiles from Fu- sui, Kwangshi]. Vertebrata PalAsiatica 13: 24-33. [In Chinese with English abstract].
56. Huene, F. von 1932. Die fossil Reptil-Ordnung Saurischia, ihre En- twicklung und Oeschichte. Monographen zur Geologie und Palae- ontologie 4: 1-361.
57. - 1956. PalHontologie und Phylogenie der niederen Tetrapoden.VEB Oustav Fisher Verlag, Jena.
58. Hunn, C. A. & Upchurch, P. 2001. The importance of time/space in diagnosing the causality ofphylogenetic events: towards a 'chronobi- ogeographical' paradigm? Systematic Biology 50: 391-407.
59. Ikejiri, T., Tidwell, V. & Trexler, D. L. 2005. New adult specimens of Camarasaurus lentus highlight ontogenetic variation within the species. Pp. 154-186 in Tidwell, V. & Carpenter, K. (eds) Thunder- lizards: the Sauropodomorph Dinosaurs. Indiana University Press, Bloomington.
60. Jacobs, L. L., Winkler, D. A., Downs, W. R. & Gomani, E. M. 1993. New material of an Early Cretaceous titanosaurid sauropod dinosaur from Malawi. Palaeontology 36: 523-534.
61. Janensch, W. 1929. Material und Formengehalt der Sauropoden in der Ausbeute der Tendaguru Expedition. Palaeontographica (Supplement 7) 2: 1-34.
62. - 1935-1936. Die Schadel der Sauropoden Brachiosaurus, Barosaurus, und Dicraeosaurus aus den Tendaguru-Schichten Deutsch-Ostafrikas. Palaeontographica (Supplement 7) 3: 147-298.
63. - 1950. Die Wirbelsaule von Brachiosaurus brancai. Palaeontographica (Supplement 7) 2: 27-93.
64. Jenner, R. A. 2001. Bilateral phylogeny and uncritical recycling of morphological data sets. Systematic Biology 50: 730-742.
65. Kim, H. M. 1983. Cretaceous dinosaurs from Korea. Journal of the Geological Society of Korea 19: 115-122.
66. Kluge, A. G. 1991. Boine snake phylogeny and research cycles. Miscellaneous Publications of the Museum of Zoology, University of Michigan 178: 1-58.
67. Ksepka, D. T. & Norell, M. A. 2006. Erketu ellisoni, a long-necked sauropod from Bor Guvé (Dornogov Aimag, Mongolia). American Museum Novitates 3508: 1-16.
68. Kurzanov, S. M. & Bannikov, A. F. 1983. A new sauropod from the Upper Cretaceous of Mongolia. Paleontological Journal 2: 90-96.
69. Lapparent, A. F. & Lavocat, R. 1955. Dinosauriens. Pp. 785-962 in Piveteau, J. (ed.) Traité dePaléontologie. Masson et Cie, Paris.
70. Lee, Y.-N., Yang, S.-Y. & Park, E.-J. 1997. Sauropod dinosaur remains from the Gyeongsang Supergroup, Korea. Palaeontological Societyof Korea, Special Publication 2: 103-114.
71. -,Yu, K.-M. &Wood, C.B. 2001. A review of the vertebrate faunas from the Gyeongsang Supergroup (Cretaceous) in South Korea. Pa- laeogeography, Palaeoclimatology, Palaeoecology 165: 357-373.
72. Leidy, J. 1858. Hadrosaurus foulkii, a new saurian from the Cretaceous of New Jersey. Proceedings of the Academy of Natural Sciences of Philadelphia 8: 72-73.
73. Le Loeuff, J. 1991. The Campano-Maastrichtian vertebrate faunas from southern Europe and their relationships with other faunas in the world: paleobiogeographical implications. Cretaceous Research 12: 93-114.
74. - 1997. Biogeography. Pp. 51-56 in Currie, P. J. & Padian, K. (eds) Encyclopedia of Dinosaurs. Academic Press Inc., San Diego.
75. - & Buffetaut, E. 1995. The evolution of Late Cretaceous non-marine vertebrate faunas in Europe. Pp. 181-184 in Sun, A. & Wang, Y. (eds) Sixth Symposium on Terrestrial Mesozoic Ecosystems and Biotas, Short Papers. China Ocean Press, Beijing.
76. Lieberman, B. S. 2000. Paleobiogeography: using fossils to study global change, plate tectonics and evolution. Kluver Academic/Plenum Publishers, New York, 208 pp.
77. Lim, J.-D., Martin, L. D. & Baek, K.-S. 2001. The first discovery of a brachiosaurid from the Asian continent. Naturwissenschaften 88:82-84.
78. Lü, J., Azuma, Y., Chen, R., Zheng, W. & Jin, X. 2008. A new ti- tanosauriform sauropod from the early Late Cretaceous of Dongyang Zhejiang Province. Acta Geologica Sinica 82: 225-235.
79. Luo, Z. 1999. A refugium for relicts. Nature 400: 23-25.
80. Lucas, S. G. & Hunt, A. P. 1989. Alamosaurus and the sauropod hiatus in the Cretaceous of the North American western interior. Geological Society ofAmerica Special Paper 238: 75-85.
81. Lydekker, R. 1893. The dinosaurs of Patagonia. Anales del Museo de La Plata 2: 1-14.
82. Madsen Jr., J. H., Mcintosh, J. S. & Berman, D. S. 1995. Skull and atlas- axis complex of the Upper Jurassic sauropod Camarasaurus Cope (Reptilia: Saurischia). Bulletin of the Carnegie Museum of Natural History 31: 1-115.
83. Manabe, M. & Hasegawa, Y. 1995. Diapsid fauna and its palaeobiogeo- graphical implication; the Neocomian section of the Tetori Group, Japan. Pp. 179 in Sun, A. & Wang, Y. (eds) Sixth Symposium on Meso- zoic Terrestrial Ecosystems and Biota, Short Papers. China Ocean Press, Beijing.
84. Mantell, G. A. 1825. Notice on the Iguanodon, a newly discovered fossil reptile, from the sandstone of Tilgate Forest, in Sussex. Philosophical Transactions ofthe Royal Society ofLondon 115: 179-186.
85. Marsh, O. C. 1878. Principal characters of American Jurassic dinosaurs. Pt. I. American Journal of Science 16: 411-416.
86. - 1882. Classification of the Dinosauria. American Journal of Science 23: 81-86.
87. - 1895. On the affinities and classification of the dinosaurian reptiles. American Journal of Science (Series 3) 50: 413-423.
88. Martin, V., Buffetaut, E. & Suteethorn, V. 1994. A new genus of saur- opod dinosaur from the Sao Khua Formation (Late Jurassic or Early Cretaceous) of northeastern Thailand. Comptes Rendus de l'Academie des Sciences Paris (serie II) 319: 1085-1092.
89. Martin-Rolland, V. 1999. Les sauropodes chinois. Revue Paléobiologie, Geneve 18: 287-315.
90. Martínez, R. D., Giménez, O., Rodríguez, J., Luna, M. & Lamanna, M. C. 2004. An articulated specimen of the basal Titanosaurian (Dino- sauria: Sauropoda) Epachthosaurus sciuttoi from the early Late Cretaceous Bajo Barreal formation of Chubut Province, Argentina. Journal of Vertebrate Paleontology 24: 107-120.
91. Mateer, N. J. & Lucas, S. G. 1985. Swedish vertebrate palaeontology in China: a history of the Lagrelius collection. Bulletin ofthe Geological Institutions of the University of Uppsala, new series 11: 1-24.
92. - & Mcintosh, J. S. 1985. A new reconstruction of the skull of Euhelopus zdanskyi (Saurischia:Sauropoda). Bulletin of the Geological Institutions of the University of Uppsala, new series 11: 25-132.
93. Mayor, A. 2000. The First Fossil Hunters. Princeton University Press, Princeton, 361 pp.
94. McIntosh, J. S. 1990. Sauropoda. Pp. 345-401 in Weishampel, D. B., Dodson, P. & Osmólska, H. (eds) The Dinosauria. University of California Press, Berkeley.
95. -, Coombs Jr., W. P. & Russell, D. A. 1992. A new diplodocid sauropod (Dinosauria) from Wyoming, U.S.A. Journal of Vertebrate Paleontology 12: 158-167.
96. Meers, M. B. 2003. Crocodylian forelimb musculature and its relevance to Archosauria. The Anatomical Record Part A 274A: 891-916.
97. Milner, A. R. & Norman, D. B. 1984. The biogeography of advanced or- nithopod dinosaurs (Archosauria: Ornithischia) -acladistic- vicariance model. Pp. 145-150 in Reif, W.-E. & Westphal, F. (eds) Third Symposium on Mesozoic Terrestrial; Ecosystems, Short Papers. Attempto Verlag, Tübingen.
98. Mo, J., Wang, W., Huang, Z., Huang, X. & Xing, X. 2006. A basal titanosauriform from the Early Cretaceous of Guangxi, China. Acta Geologica Sinica 80: 486-489.
99. -, Huang, C.-L., Zhao, Z.-R., Wang, W. & Xing, X. 2008. [A new titanosaur (Dinosauria: Sauropoda) from the Late Cretaceous of Guangxi, China]. Vertebrata PalAsiatica 46:147-156 [In Chinese with English summary].
100. Needham, J. 1956. Science and Civilisation in China. Volume 2: History of Scientific Thought. Cambridge University Press, Cambridge,697 pp.
101. Nelson, G. & Platnick, N. 1981. Systematics and Biogeography: Cladist-ics and Vicariance. Columbia University Press, New York, 567 pp.
102. Norman, D.B. 1998. On Asian ornithopods (Dinosauria: Ornithischia). 3. A new species of iguanodontid dinosaur. Zoological Journal of the Linnean Society of London122:291-348.
103. Nowinski, A. 1971. Nemegtosaurus mongoliensis n. gen., s. sp. (Sauro-poda) from the Uppermost Cretaceous of Mongolia. Palaeontologia Polonica 25: 57-81.
104. Osborn, H. F. 1923. Two Lower Cretaceous dinosaurs from Mongolia. American Museum Novitates 93: 1-3.
105. - 1924a. Psittcosaurus and Protiguanodon: two Lower Cretaceous iguanodonts from Mongolia. American Museum Novitates 127: 1-16.
106. - 1924b. Sauropoda and Theropoda of the Lower Cretaceous of Mongolia. American Museum Novitates 128: 1-7.
107. Ostrom, J. H. & Mcintosh, J. S. 1966. Marsh's Dinosaurs. Yale University Press, New Haven, 388 pp.
108. Ouyang, H. & Ye, Y. 2002. [Thefirst mamenchisaurian skeleton with complete skull Mamenchisaurus youngi]. Sichuan Scientific and Technology Publications, Sichuan, 111 pp. [In Chinese with English abstract].
109. Owen, R. 1841. A description of a portion of the skeleton of Cetiosaurus,a gigantic extinct saurian occurring in the Oolitic Formation of different parts of England. Proceedings ofthe Geological Society of London 3: 457-462.
110. Page, R.D.M. 1991. Random dendrograms and null hypotheses in cladistic biogeography. Systematic Zoology40:54-62.
111. Pang, Q. & Cheng, Z. 2000. A new family of sauropod dinosaur from the Upper Cretaceous of Tianzhen, Shanxi Province, China. Acta Geolo-gica Sinica 74: 117-125.
112. Plot, R. 1677. The Natural History of Oxford-shire, being an essay towards the Natural History of England. The Theater, Oxford, 358 pp.
113. Rauhut, O. W. M. 2003. A dentary of Patagosaurus (Sauropoda) from the Middle Jurassic of Patagonia. Ameghiniana 40: 425-432.
114. - 2006. A brachiosaurid sauropod from the Late Jurassic Cañadón Calcáreo Formation of Chubut, Argentina. Fossil Record 9: 226-237.
115. Rich, T. H., Vickers-Rich, P., Gimenez, O., Cúneo, R., Puerta, P. & Vacca, R. 1999. A new sauropod dinosaur from Chubut Province, Argentina. Pp. 61-84 in Tomida, Y., Rich, T. H. & Vickers-Rich, P. (eds) Proceedings of the Second Gondwana Dinosaur Symposium. National Science Museum Monographs, Tokyo.
116. Rieppel, O. & Reisz, R. R. 1999. The origin and early evolution of turtles. Annual Review of Ecology and Systematics 30: 1-22.
117. Romer, A. S. 1956. Osteology of the Reptiles. University of Chicago Press, Chicago, 772 pp.
118. Ruiz-Omeiñaca, J. I., Canudo, J. I. & Cuenca Bescoós, G. 1997. Primera evidencia de un area de alimentacioón de dinosaurios herbivoros en el cretácico inferior de Espafia (Teruel). Monografias de la Academia de Ciencias Exactas Físicas Quimicas y Naturales de Zaragoza 10: 1-48.
119. Russell, D. A. 1993. The role of central Asia in dinosaurian biogeography. Canadian Journal ofEarth Science 30: 2002-2012.
120. - 1995. China and the lost worlds of the dinosaurian era. Historical Biology 10: 3-12.
121. -Zheng, Z. 1993. A large mamenchisaurid from the Junggar Basin, Xinjiang, People's Republic of China. Canadian Journal of Earth Sciences 30: 2082-2095.
122. Ryabinin, A. N. 1939. [Some results of the studies of the Upper Cretaceous dinosaurian fauna from the vicinity of the Station Sary-Agach, south Kazakhstan.] Problemy Palaeontologii 4: 175-184. [In Russian].
123. Salgado, L., Coria, R. & Calvo, J. O. 1997. Evolution of titanosaurid sauropods. I: phylogenetic analysis based on the postcranial evidence. Ameghiniana 34: 3-32.
124. -, Apesteguia, S. &Heredia, S. 2005. A new specimen of Neuquensaurus australis, a Late Cretaceous saltasaurine titanosaur from north Patagonia. Journal of Vertebrate Paleontology 25: 623-634.
125. -, Carvalho, I. S. & Garrido, A. C. 2006. Zapalasaurus bonapartei,un nuevo dinosaurio saurópodo de La Formacioón La Amarga (Cretácico Inferior), noroeste de Patagonia, Provincia de Neuquén, Argentina. Geobios 39: 695-707.
126. Sanz, J. L., Powell, J. E., Loeuff, J. L., Martinez, R. & Pereda Suberbiola, X. 1999. Sauropod remains from the Upper Cretaceous of Lafĩo (northcentral Spain). Titanosaur phylogenetic relationships. Estudios del Museo de Ciencias Naturales de Alava 14: 235-255.
127. Säve-Söderbergh, G. 1946. Carl Wiman 1867 10/3 - 1944 15/6 [obituary]. Avtryck ur Geologiska FÜreningens i Stockholm Förhandlingar 68: 393-404.
128. Scotese, C. R. 2004. Cenozoic and Mesozoic paleogeography: changing terrestrial pathways. Pp. 9-26 in Lomolino, M. V. & Heaney, L. R. (eds) Frontiers of Biogeography. Sinauer Associates, Sunderland, Massachusetts.
129. Seeley, H. G. 1887. The classification of the Dinosauria. Geological Magazine 3: 562.
130. Sereno, P. C. 1986. Phylogeny of the bird-hipped dinosaurs (order Or- nithischia). National Geographic Research 2: 234-256.
131. - 1999. Evolution of dinosaurs. Science 284: 2137-2147.
132. - 2007. Logical basis for morphological characters in phylogenetics. Cladistics 23: 1-23.
133. -, Wilson, J. A., Witmer, L. M., Whitlock, J. A., Maga, A., Ide, O. & Rowe, T. A. 2007. Structural extremes in a Cretaceous dinosaur. PLoS ONE 2: e1230. doi:10.1371/journal.pone.0001230.
134. Sikes, D. S. & Lewis, P. O.2001. PAUPRat: a tool to implement Parsimony Ratchet searches using PAUP. Department of Ecology and Evolutionary Biology, University of Connecticut, Storrs.
135. Simpson, G. G. 1952. Probabilities of dispersal in geologic time. Bulletin ofthe American Museum ofNatural History 99: 163-176.
136. Smith, A. G., Smith, D. G. & Funnell, B. M. 1994. Atlas of Mesozoic and Cenozoic coastlines. Cambridge University Press, Cambridge, 102 pp.
137. Solounias, N. 1999. The remarkable anatomy of the giraffe's neck. Journal of Zoology247: 257-268.
138. Stannius, H. 1846. Lehrbuch der vergleichenden Anatomie der Wirbelth-iere. Verlag von Veit & Comp., Berlin, 482 pp.
139. Steel, R. 1970. Saurischia. Handbuch der Palcï oherpetologie 13: 1-88.
140. Sues, H.-D. & Averianov, A. O. 2004. Dinosaurs from the Upper Cretaceous (Turonian) of Dzharakuduk, Kyzlkum Desert, Uzbekistan. Journal ofVertebrate Paleontology 24: 119A-120A.
141. Swinton, W. E. 1947. New discoveries of Titanosaurus indicus Lyd. Annals and Magazine of Natural History (series 11) 14: 112-123.
142. Swofford, D.L. 2002. PAUP: Phylogenetic Analysis Using Parsimony version 4.10b10. Sinauer Associates, Sunderland, Massachusetts.
143. Tang, F., Jin, X., Kang, X. & Zhang, G. 2001 a.[Omeisaurus maoianusa complete Sauropoda from Jingyan, Sichuan]. Research works of Natural History Museum of Zhejiang 10: 1-127. [In Chinese with English summary].
144. -, Kang, X.-M., Jin, X.-S., Feng, W. & Wu, W.-T. 2001b.A new uropod dinosaur of Cretaceous from Jiangshan, Zheijiang Province. Vertebrata PalAsiatica 39: 272-281.
145. Tanimoto, M. & Suzuki, C. 1998. Wear facet of teeth in the Nemegto-sauridae -a new knowlege ofSuzuki's sauropod specimen from Iwaki. Research Reports Iwaki Natural History 9: 1-3.
146. Tidwell, V., Carpenter, K. & Meyer, S.. 2001. New titanosauriform (Sauropoda) from the Poison Strip Member of the Cedar Mountain Formation (Lower Cretaceous), Utah. Pp. 139-165 in Tanke, D. & Carpenter, K. (eds) Mesozoic Vertebrate Life: New Research Inspired by the Paleontology of Phillip J. Currie. Indiana University Press, Bloomington.
147. -, Stadtman, K. L. & Shaw, A. 2005. Age-related characteristics found in a partial pelvis of Camarasaurus. Pp. 180-186 in Tidwell, V. & Carpenter, K. (eds) Thunder-Lizards: The Sauropodomorph Dinosaurs. Indiana University Press, Bloomington.
148. Tomida, Y. & Tsumura, Y. 2006. A partial skeleton of titanosaurian saur- opod from the Early Cretaceous of Toba City, central Japan. Journal ofthe Paleontological Society of Korea 22: 217-238.
149. Tsuihiji, T. 2004. The ligament system in the neck of Rhea americana and its implication for the bifurcated neural spines of sauropod dinosaurs. Journal ofVertebrate Paleontology 24: 165-172.
150. Upchurch, P. 1995. Evolutionary history of sauropod dinosaurs. Philosophical Transactions ofthe Royal SocietyofLondon B 349: 365-390.
151. -1998. The phylogenetic relationships of sauropod dinosaurs. Zoological Journal ofthe Linnean Society 124: 43-103.
152. - 1999. The phylogenetic relationships of the Nemegtosauridae. Journal ofVertebrate Paleontology 19: 106-125.
153. - & Barrett, P. M. 2000. The evolution of sauropod feeding. Pp. 79-122 in Sues, H.-D. (ed.) Evolution ofFeeding in Terrestrial Vertebrates. Cambridge University Press, Cambridge.
154. 2005. Phylogenetic and taxic perspectives on sauropod diversity. Pp. 104-124 in Curry Rogers, K. A. & Wilson, J. A. (eds) The Saur-opods: Evolution and Paleobiology. University of California Press, Berkeley and Los Angeles.
155. - & Hunn, C. A. 2002 'Time': the neglected dimension in cladistic biogeography? Geobios 24, 277-286.
156. - & Martin, J. 2002. The Rutland Cetiosaurus: the anatomy and relationships of a Middle Jurassic British sauropod dinosaur. Palaeontology 45: 1049-1074.
157. -, Hunn, C. A. & Norman, D. B. 2002. An analysis of dinosaurian biogeography: evidence for the existence of vicariance and dispersal patterns caused by geological events. Proceedings ofthe Royal Society of London B 269: 613-621.
158. -, Barrett, P. M. & Dodson, P. 2004a. Sauropoda. Pp. 259-324 in Weishampel, D. B., Dodson, P. & Osmólska, H. (eds) The Dinosauria. 2nd Edition. University of California Press, Berkeley.
159. -, Tomida, Y. & Barrett, P. M. 2004b. A new specimen of Apatosaurus ajax (Sauropoda: Diplodocidae) from the Morrison Formation (Upper Jurassic) of Wyoming, USA. National Science Museum Monographs 26: 1-108.
160. -, Barrett, P. M., Zhao, X. & Xing, X. 2007. A re-evaluation of Chin- shakiangosaurus chunghoensis Ye vide Dong 1992 (Dinosauria, Saur- opodomorpha): implications for cranial evolution in basal sauropod dinosaurs. Geological Magazine 144: 247-262.
161. Wagler, J. 1832. Neue Sippen und Oattungen der Vogel. Isis 11: 1224.
162. Wang, X., Wang, Y., Wang, Y., Xu, X., Tang, Z., Zhang, F. & Hu, Y. 1998. [Stratigraphic sequence and vertebrate-bearing beds of the lower part of the Yixian Formation in Sihetun and neighboring area, western Liaoning, China. Vertebrata Pal 4Asiatica 36: 81-101. [In Chinese with English abstract].
163. -, You, H., Meng, Q., Gao, C., Cheng, X. & Liu, J. 2007. Dongbeititan dongi, the first sauropod dinosaur from the Lower Cretaceous Jehol Oroup of western Liaoning Province, China. Acta Geologica Sinica 81: 911-916.
164. Wedel, M. J., Cifelli, R. L. & Sanders, R. K. 2000. Sauroposeidon proteles, a new sauropod from the Early Cretaceous of Oklahoma. Journal ofVertebrate Paleontology 20: 109-114.
165. Wedel, M. J. & Sanders, R. K. 2002. Osteological correlates of cervical musculature in Aves and Sauropoda (Dinosauria: Saurischia), with comments on the cervical ribs of Apatosaurus. PaleoBios 22: 1-6.
166. Weishampel, D. B. 1990. Dinosaurian distributions. Pp. 63-189 in Weishampel, D. B., Dodson, P. & Osmólska, H. (eds) The Dinosauria. University of California Press, Berkeley.
167. - & Bjork, P. R. 1989. The first indisputable remains of Iguanodon (Or- nithischia: Ornithopoda) from North America: Iguanodon lakotaensis, sp. nov. Journal ofVertebrate Paleontology 9: 56-66.
168. - & Jianu, C.-M. 1997. The importance of phylogeny in paleogeo- graphic analyses, with examples from North-American hadrosaurids and European titanosaurids. Sargetia, Scientia Naturae 17: 261-279.
169. -, Barrett, P. M., Coria, R. A., Le Loeuff, J., Xing, X., Sahni, A., Gomani, E.M. &Noto, C.R. 2004. Dinosaurian distributions. Pp. 517-606 in Weishampel, D. B., Dodson, P. & Osmoólska, H. (eds) The Dinosauria, 2nd Edition. University of California Press, Berkeley.
170. Wilkinson, M. 1994. Common cladistic information and its consensus representation: reduced Adams and reduced cladistic consensus trees and profiles. Systematic Biology 43: 343-368.
171. - 1995. Coping with abundant missing entries in phylogenetic inference using parsimony. Systematic Biology 44: 501-514.
172. Wilson, J.A. 1999. A nomenclature for vertebral laminae in sauropods and other saurischian dinosaurs. Journal ofVertebrate Paleontology19:639-653.
173. - 2002. Sauropod dinosaur phylogeny: critique and cladistic analysis. Zoological Journal ofthe Linnean Society 136: 217-276.
174. - 2005a. Redescription of the Mongolian sauropod Nemegtosaurus mon- goliensis Nowinski (Dinosauria: Saurischia) and comments on Late Cretaceous sauropod diversity. Journal of Systematic Palaeontology 3: 283-318.
175. - 2005b. Morphological patterns in sauropod evolution. Pp. 15-49 in Curry Rogers, K. A. & Wilson, J. A. (eds) The Sauropods: Evolution and Paleobiology. University of California Press, Berkeley.
176. - 2005c. Integrating ichnofossil and body fossil records to estimate lo- comotor posture and spatiotemporal distribution of early sauropod dinosaurs: a stratocladistic approach. Paleobiology 31: 400-423.
177. - 2006. Anatomical nomenclature of fossil vertebrates: standardized terms or lingua franca? Journal ofVertebrate Paleontology 26: 511-518.
178. - & Carrano, M. T. 1999. Titanosaurs and the origin of "wide-gauge" trackways: a biomechanical and systematic perspective on sauropod locomotion. Paleobiology 25: 252-267.
179. - & Sereno, P. C. 1998. Early evolution and higher-level phylogeny of sauropod dinosaurs. Journal ofVertebrate Paleontology 18: 1-68.
180. - & Upchurch, P. 2003. A revision of Titanosaurus Lydekker (Dinosauria - Sauropoda), the first dinosaur genus with a 'Oondwanan' distribution. Journal ofSystematic Palaeontology 1: 125-160.
181. -, Sereno, P. C., Srivastava, S., Bhatt, D. K., Khosla, A. & Sahni, A. 2003. A new abelisaurid (Dinosauria, Theropoda) from the Lameta Formation (Cretaceous, Maastrichtian) of India. Contributions from the Museum ofPaleontology, The University ofMichigan 31: 1-42.
182. Wiman, C. 1929. Die Kreide-Dinosaurier aus Shantung. Palaeontologia Sinica (series C) 6: 1-67.
183. Wu, W.-H., Dong, Z.-M., Sun, Y.-W., Li, C.-T. & Li, T. 2006. A new sauropod dinosaur from the Cretaceous of Jiutai, Jilin, China. Global Geology 25: 6-8.
184. Wu, X.-C., Brinkman, D. B. & Lu, J.-C. 1994. A new species ofShantun- gosuchus from the Lower Cretaceous of Inner Mongolia (China), with comments on S. chuhsienensis Young, 1961 and the phylogenetic position of the genus. Journal ofVertebrate Paleontology 14: 210229
185. Xu, X., Zhang, X., Tan, Q., Zhao, X. & Tan, L. 2006. A new titanosaurian sauropod from Late Cretaceous of Nei Mongol, China. Acta Geologica Sinica 80: 20-26.
186. Yates, A. & Kitching, J. W. 2003. The earliest known sauropod dinosaur and the first steps towards sauropod evolution. Proceedings of the Royal Society of London B 270: 1753-1758.
187. You, H., Tang, F. & Luo, Z. 2003. A new basal titanosaur (Dinosauria: Sauropoda) from the Early Cretaceous of China. Acta Geologica Sinica 77: 424-429.
188. -, Ji, Q., Lamanna, M. C., Li, J. &Li, Y. 2004. A titanosaurian sauropod dinosaur with opisthocoelous caudal vertebrae from the Early Late Cretaceous of Liaoning Provice, China. Acta Geologica Sinica 78: 907-911.
189. -, Li, D., Zhou, L. & Ji, Q. 2006. Huanghetitan luijiaxiaensis,anew sauropod dinosaur from the Lower Cretaceous Hekou Oroup of Lanzhou Basin, Oansu Province, China. Geological Review 52: 668-674.
190. Young, C. C. 1935. Dinosaurian remains from Mengyin, Shantung. Bulletin ofthe Geological Society ofChina 15: 519-533.
191. - 1937. Anew dinosaurian from Sinkiang. Palaeontologica Sinica (series C) 2: 1-25.
192. - 1939. On a new Sauropoda, with notes on other fragmentary reptiles from Szechuan. Bulletin ofthe Geological Society of China 19: 279-315.
193. - 1954. On a new sauropod from Yiping, Szechuan, China. Acta Scientia Sinica 3: 491-504.
194. - 1958. New sauropods from China. Vertebrata PalAsiatica 2: 1-27.
195. - & Zhao, X.-J. 1972. [Mamenchisaurus hochuanensis, sp. nov.]. Aca- demia Sinica 8: 1-30. [In Chinese with English abstract].
196. Zhang, Y. 1988. [The Middle Jurassic dinosaur fauna from Dashanpu, Zigong, Sichuan]. Journal ofthe Chengdu College of Geology 3:187. [In Chinese with English abstract].
197. Zhou, Z., Barrett, P. M. & Hilton, J. 2003. An exceptionally preserved Lower Cretaceous ecosystem. Nature 421: 807-814.