Integrating ichnofossil and body fossil records to estimate locomotor posture and spatiotemporal distribution of early sauropod dinosaurs: A stratocladistic approach

Paleobiology

Volumn 31, Issue 3, 2005, Pages 400-423

  Wilson, Jeffrey A ^a  

^a UNIVERSITY OF MICHIGAN   (United States)

Author keywords

[No Author keywords available]

Indexed keywords

DINOSAUR; FOSSIL RECORD; POSTURE; SPATIOTEMPORAL ANALYSIS;

DINOSAURIA; SAUROPODA; VERTEBRATA;

EID: 24344482368     PISSN: 00948373     EISSN: None     Source Type: Journal    
DOI: 10.1666/0094-8373(2005)031[0400:IIABFR]2.0.CO;2     Document Type: Article

References (122)

1. Alifanov, V., and A. O. Averianov. 2003. Ferganasaurus verzilini, gen. et sp. nov., a new neosauropod (Dinosauria, Saurischia, Sauropoda) from the Middle Jurassic of Fergana Valley, Kirghizia. Journal of Vertebrate Paleontology 23:358-372.
2. Allain, R., N. Aquesbi, J. Dejax, C. Meyer, M. Monbaron, C. Montenat, P. Richir, M. Rodidy, D. Russell, and P. Taquet. 2004. A basal sauropod dinosaur from the Early Cretaceous of Morocco. Comptes Rendus Palevol 3:199-208.
3. Baird, D. 1980. A prosauropod trackway from the Navajo Sandstone (Lower Jurassic). Pp. 219-230 in L. Jacobs, ed. Aspects of vertebrate history: essays in honor of Edwin Harris Colbert. Museum of Northern Arizona, Flagstaff.
4. Bakker, R. T. 1971. Ecology of the brontosaurs. Nature 229:172-174.
5. Bandyopadhyay, S., D. P. Sengupta, and D. D. Gillette. 2003. Dentition of Barapasaurus tagorei from the Kota Formation (Upper Jurassic) of India. Journal of Vertebrate Paleontology 23:31A-32A.
6. Barrett, P. M. 1999. A sauropod dinosaur from the Lower Lufeng Formation (Lower Jurassic) of Yunnan Province, People's Republic of China. Journal of Vertebrate Paleontology 19:785-787.
7. Benton, M. J. 1993. The fossil record 2. Chapman and Hall, London.
8. Benton, M. J., L. Juul, G. W. Storrs, and P. M. Galton. 2000. Anatomy and systematics of the prosauropod dinosaur Thecodontosaurus antiquus from the Upper Triassic of southwest England. Journal of Vertebrate Paleontology 20:77-108.
9. Bird, R. T. 1941. A dinosaur walks into the museum. Natural History 47:74-81.
10. Bird, R. T. 1944. Did Brontosaurus ever walk on land? Natural History 53:60-67.
11. Bloch, J. I., D. C. Fisher, K. D. Rose, and P. D. Gingerich. 2001. Stratocladistic analysis of Paleocene Carpolestidae (Mammalia, Plesiadapiformes) with description of a new late Tiffanian genus. Journal of Vertebrate Paleontology 21:119-131.
12. Bodenbender, B. E., and D. C. Fisher. 2001. Stratocladistic analysis of blastoid phylogeny. Journal of Paleontology 75:351-369.
13. Bonaparte, J. F. 1999. Evolución de las vértebras presacras en Sauropodomorpha. Ameghiniana 36:115-187.
14. Bonaparte, J. F., and J. A. Pumares. 1995. Notas sobre el primer craneo de Riojasaurus incertus (Dinosauria, Prosauropoda, Melanorosauridae) del Triásico superior de La Rioja, Argentina. Ameghiniana 32:341-349.
15. Bonnan, M. F. 2003. The evolution of manus shape in sauropod dinosaurs: implications for functional morphology, limb orientation, and phylogeny. Journal of Vertebrate Paleontology 23:595-613.
16. Buffetaut, E. 2003. A sauropod with prosauropod teeth from the Jurassic of Madagascar. P. 15 in C. A. Meyer, ed. First European Association of Vertebrate Paleontologists Meeting. Natural History Museum, Basel.
17. Buffetaut, E., V. Suteethorn, G. Cuny, H. Tong, J. LeLoeuff, S. Khansubha, and S. Jongautchariyakul. 2000. The earliest known sauropod dinosaur. Nature 407:72-74.
18. Buffetaut, E., V. Suteethorn, J. Le Loeuff, G. Cuny, H. Tong, and S. Khansubha. 2002. The first giant dinosaurs: a large sauropod from the Late Triassic of Thailand. Comptes Rendus Palevoll: 1:103-109.
19. Calvo, J. O., and L. Salgado. 1995. Rebbachisaurus tessonei, sp. nov. a new Sauropoda from the Albian-Cenomanian of Argentina: new evidence on the origin of Diplodocidae. Gaia 11:13-33.
20. Carrano, M. T. 1997. Morphological indicators of foot posture in mammals: a statistical and biomechanical analysis. Zoological Journal of the Linnean Society 121:77-104.
21. Carrano, M. T. 2000. Homoplasy and the evolution of dinosaur locomotion. Paleobiology 26:489-512.
22. Carrano, M. T., and J. A. Wilson. 2001. Taxon distributions and the tetrapod track record. Paleobiology 27:563-581.
23. Casamiquela, R. M. 1967. Un nuevo dinosaurio ornitisquio Triásico ( Pisanosaurus mertii; Ornithopoda) de la formación Ischigualasto, Argentina. Ameghiniana 4:47-64.
24. Charig, A. J., J. Attridge, and A. W. Crompton. 1965. On the origin of the sauropods and the classification of the Saurischia. Proceedings of the Linnean Society of London 176:197-221.
25. Chatterjee, S. 1984. A new ornithischian dinosaur from the Triassic of North America. Naturwissenschaften 71:630.
26. Conrad, K., M. G. Lockley, and N. K. Prince. 1987. Triassic and Jurassic vertebrate-dominated trace fossil assemblages of the Cimarron Valley region. In S. G. Lucas and A. P. Hunt, eds. Northeastern New Mexico. New Mexico Geological Society Guidebook 38:127-138.
27. Coombs, W. P., Jr. 1975. Sauropod habits and habitats. Palaeogeography, Palaeoclimatology, Palaeoecology 17:1-33.
28. Cooper, M. R. 1981. The prosauropod dinosaur Massospondylus carinatus Owen from Zimbabwe: its biology, mode of life and phylogenetic significance. Occasional Papers of the National Museum of Rhodesia B 6:689-840.
29. Dalla Vecchia, F. M. 1994. Jurassic and Cretaceous sauropod evidence in the Mesozoic carbonate platforms of the southern Alps and Dinards. Gaia 10:65-73.
30. Day, J. J., P. Upchurch, D. B. Norman, A. S. Gale, and H. P. Powell. 2002. Sauropod trackways, evolution, and behavior. Science 296:1659.
31. Demathieu, G. R. 1990. Appearance of the first dinosaur tracks in the French Middle Triassic and their probable significance. Pp. 201-207 in D. D. Gillette and M. G. Lockley, eds. Dinosaur tracks and traces. Cambridge University Press, Cambridge.
32. Dong, Z. M. 1992. The dinosaurian faunas of China. Springer, Berlin.
33. Dutuit, J.-M. 1972. Découverte d'un dinosaure ornithischien dans le Trias supérieur de l'Atlas occidental marocain. Comptes Rendus de l'Académie des Sciences, Paris, D 275:2841-2844.
34. Ellenberger, P. 1972. Contribution à la classification des piste de vertébrés du Trias: les types du Stormberg d'Afrique do Sud (1). Palaeovertebrata, Mémoire Extraordinaire 1:1-152.
35. Farlow, J. O. 1992. Sauropod tracks and trackmakers: integrating the ichnological and skeletal records. Zubía 10:89-138.
36. Farlow, J. O., J. G. Pittman, and M. J. Hawthorne. 1989. Brontopodus birdi, Lower Cretaceous sauropod footprints from the U.S. Gulf Coastal Plain. Pp. 371-394 in M. G. Lockley and D. D. Gillette, eds. Dinosaur tracks and traces. Cambridge University Press, Cambridge.
37. Fisher, D. C. 1992. Stratigraphic parsimony. Pp. 124-129 in W. P. Maddison and D. R. Maddison, eds. MacClade version 3: analysis of phylogeny and character evolution. Sinauer, Sunderland, Mass.
38. Fisher, D. C. 1994. Stratocladistics: morphological and temporal patterns and their relation to phylogenetic trees. Pp. 133-171 in L. Grande and O. Rieppel, eds. Interpreting the hierarchy of nature. Academic Press, Los Angeles.
39. Fisher, D. C., M. Foote, D. L. Fox, and L. R. Leighton. 2002. Stratigraphy in phylogeny reconstruction - comment on Smith (2000). Journal of Paleontology 76:585-586.
40. Flynn, J. J., J. M. Parrish, B. Rakotosamimanana, W. F. Simpson, R. L. Whatley, and A. R. Wyss. 1999. A Triassic fauna from Madagascar, including early dinosaurs. Science 286:763-765.
41. Fox, D. L., D. C. Fisher, and L. R. Leighton. 1999. Reconstructing phylogeny with and without temporal data. Science 284:1816-1819.
42. Galton, P. M. 1976. Prosauropod dinosaurs (Reptilia: Saurischia) of North America. Postilla 169:1-98.
43. Galton, P. M. 1985. Notes on the Melanorosauridae, a family of large prosauropod dinosaurs (Saurischia: Sauropodomorpha). Geobios 18:671-676.
44. Galton, P. M. 1990. Prosauropoda. Pp. 320-345 in D. B. Weishampel, P. Dodson, and H. Osmólska, eds. The Dinosauria. University of California Press, Berkeley.
45. Galton, P. M. 2000. The prosauropod dinosaur Plateosaurus Meyer, 1837 (Saurischia: Sauropodomorpha). I. The syntypes of P. engelhardti Meyer, 1837 (Upper Triassic, Germany), with notes on other European sauropods with "distally straight" femora. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 216:233-275.
46. Galton, P. M., and P. Upchurch. 2000. Prosauropod dinosaurs: homeotic transformations ("frame shift") with third sacral as a caudosacral or a dorsosacral. Journal of Vertebrate Paleontology 20:43A.
47. Galton, P. M., and P. Upchurch, 2004. Prosauropoda. Pp. 232-258 in D. B. Weishampel, P. Dodson, and H. Osmólska, eds. The Dinosauria, 2d ed. University of California Press, Berkeley.
48. Galton, P. M., and J. van Heerden. 1985. Partial hindlimb of Blikanasaurus cromptoni n. gen., and n. sp., representing a new family of prosauropod dinosaurs from the Upper Triassic of South Africa. Geobios 18:509-516.
49. Galton, P. M., and J. van Heerden. 1998. Anatomy of the prosauropod dinosaur Blikanasaurus cromptoni (Upper Triassic, South Africa), with notes on the other tetrapods from the lower Elliot Formation. Paläontologische Zeitschrift 72:163-177.
50. Gauthier, J. A. 1986. Saurischian monophyly and the origin of birds. Memoirs of the California Academy of Sciences 8:1-55.
51. Gierlinski, G. 1997. Sauropod tracks in the Early Jurassic of Poland. Acta Palaeontologica Polonica 42:533-538.
52. Gillette, D. D. 2003. The geographic and phylogenetic position of sauropod dinosaurs from the Kota Formation (Early Jurassic) of India. Journal of Asian Earth Sciences 23:683-689.
53. Gilmore, C. W. 1936. Osteology of Apatosaurus with special reference to specimens in the Carnegie Museum. Memoirs of the Carnegie Museum 11:175-300.
54. Gradstein, F. M., F. P. Agterberg, J. G. Ogg, J. Hardenbol, and S. Backstrom. 1999. On the Cretaceous time scale. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 212:3-14.
55. Hatcher, J. B. 1901. Diplodocus (Marsh): its osteology, taxonomy, and probable habits, with a restoration of the skeleton. Memoirs of the Carnegie Museum 1:1-63.
56. Hatcher, J. B. 1902. Structure of the forelimb and manus of Brontosaurus. Annals of the Carnegie Museum 1:356-376.
57. Haughton, S. H. 1924. The fauna and stratigraphy of the Stormberg Series. Annals of the South African Museum 12:323-497.
58. He, X.-L., K. Li, and K.-J. Cai. 1988. The Middle Jurassic dinosaur fauna from Dashanpu, Zigong, Sichuan, Vol. IV. Sauropod dinosaurs (2) Omeisaurus tianfuensis. Sichuan Scientific and Technological Publishing House, Chengdu. [In Chinese with English abstract.]
59. He, X., C. Wang, S. Liu, F. Zhou, T. Liu, K. Cai, and B. Dai. 1998. A new sauropod dinosaur from the Early Jurassic in Gongxian County, South Sichuan. Acta Geologica Sichuan 18:1-6. [In Chinese with English abstract.]
60. Huene, F. von. 1920. Bemerkungen zur Systematik und Stammesgeschicht einigen Reptilien. Zeitschrift für induktive Astammungs- und Vererbungslehre 22:209-212.
61. Huene, F. von. 1926. Vollständige Osteologie eines Plateosauriden aus dem schwäbischen Keuper. Geologie und Paläontologie Abhandlungen 15:129-179.
62. Ishigaki, S. 1988. Les empreintes de dinosaures du Jurassique inférieur du Haut Atlas central marocain. Notes du Service Géologique du Maroc 44:79-86.
63. Jain, S. L., T. S. Kutty, T. Roy-Chowdhury, and S. Chatterjee. 1975. The sauropod dinosaur from the Lower Jurassic Kota Formation of India. Proceedings of the Royal Society of London B 188:221-228.
64. Langer, M. C., F. Abdala, M. Richter, and M. J. Benton. 1999. A sauropodomorph dinosaur from the Upper Triassic (Carnian) of southern Brazil. Comptes Rendus de l'Académie des Sciences, Paris 329:511-517.
65. Lockley, M. G. 1986. Dinosaur tracksites: a field guide published in conjunction with the First International Symposium on Dinosaur Tracks and Traces. University of Colorado at Denver, Geology Department Magazine 1:1-56.
66. Lockley, M. G. 1991. Tracking dinosaurs: a new look at an ancient world. Cambridge University Press, Cambridge.
67. Lockley, M. G., and J. R. Foster. 2003. Late Cretaceous mammal tracks from North America. Ichnos 10:269-276.
68. Lockley, M. G., and A. G. Hunt. 1995. Dinosaur tracks and other fossil footprints of the western United States. Columbia University Press, New York.
69. Lockley, M. G., and C. A. Meyer. 2000. Dinosaur tracks and other fossil footprints of Europe. Columbia University Press, New York.
70. Lockley, M. G., A. Meyer, A. G. Hunt, and S. G. Lucas. 1994. The distribution of sauropod tracks and trackmakers. Gaia 10: 233-248.
71. Lockley, M. G., J. L. Wright, S. G. Lucas, and A. G. Hunt. 2001. The Late Triassic sauropod track record comes into focus: old legacies and new paradigms. In S. G. Lucas and D. Ulmer-Scholle, eds. Geology of the Llano Estacado. New Mexico Geological Society Guidebook 52:181-190.
72. Longman, H. A. 1926. A giant dinosaur from Durham Downs. Memoirs of the Queensland Museum 8:183-194.
73. Lucas, S. G. 2003. Triassic tetrapod footprint biostratigraphy and biochronology. Albertiana 28:75-84.
74. Maddison, W. P., and D. R. Maddison. 2000. MacClade: analysis of phylogeny and character evolution, version 4.0. Sinauer, Sunderland, Mass.
75. Mantell, G. A. 1850. On the Pelorosaurus: an undescribed gigantic terrestrial reptile whose remains are associated with those of the Iguanodon and other saurians in the strata of Tilgate Forest, in Sussex. Philosophical Transactions of the Royal Society of London 140:379-390.
76. Marsh, O. C. 1878. Principal characters of American Jurassic dinosaurs. Pt. I. American Journal of Science, series 3,16:411-416.
77. Marsicano, C. A., and S. P. Barredo. 2004. A Triassic tetrapod footprint assemblage from southern South America: palaeobiogeographical and evolutionary implications. Palaeogeography, Palaeoclimatology, Palaeoecology 203:313-335.
78. McIntosh, J. S. 1990. Sauropoda. Pp. 345-401 in D. B. Weishampel, P. Dodson, and H. Osmólska, eds. The Dinosauria. University of California Press, Berkeley.
79. e cycle. Université de Paris, Paris.
80. Olsen, P. E. 1995. A new approach for recognizing track makers. Geological Society of America Abstracts with Programs 27:72.
81. Ouyang, H., and Y. Ye. 2002. The first mamenchisaurian skeleton with complete skull, Mamenchisaurus youngi. Sichuan Science and Technology Publishing House, Chengdu. [In Chinese with extended English abstract.]
82. Phillips, J. 1871. Geology of Oxford and the Valley of the Thames. Clarendon, Oxford.
83. Polly, P. D. 1997. Ancestry and species definition in paleontology: a stratocladistic analysis of Paleocene-Eocene Viverradidae (Mammalia, Carnivora) from Wyoming. Contributions from the Museum of Paleontology, University of Michigan 30: 1-53.
84. Raath, M. 1972. Fossil vertebrate studies in Rhodesia: a new dinosaur (Reptilia, Saurischia) from the near the Trias-Jurassic boundary. Arnoldia 30:1-37.
85. Rainforth, E. C. 2003. Revision and re-evaluation of the Early Jurassic dinosaurian ichnogenus Otozoum. Palaeontology 46: 803-838.
86. Reig, O. 1963. La presencia de dinosaurios saurisquios en los "Estratos de Ischigualasto" (Mesotriasico superior) de las provincias de San Juan y La Rioja (Republico Argentina). Ameghiniana 3:3-20.
87. Riggs, E. S. 1904. Structure and relationships of opisthocoelian dinosaurs, Part II. The Brachiosauridae. Publications of the Field Columbian Museum, Geological Series 2:229-248.
88. Rogers, R. R., C. C. Swisher III, P. C. Sereno, A. M. Monetta, C. A. Forster, and R. N. Martinez. 1993. The Ischigualasto tetrapod assemblage (Late Triassic, Argentina) and 40Ar/39Ar dating of dinosaur origins. Science 260:794-797.
89. Russell, D. A., and Z. Zheng. 1993. A large mamenchisaurid from the Junggar Basin, Xinjiang, People's Republic of China. Canadian Journal of Earth Science 30:2082-2095.
90. Santos, V. F., M. G. Lockley, C. A. Meyer, J. Carvalho, A. M. Galopim, and J. J. Moratalla. 1994. A new sauropod tracksite from the Middle Jurassic of Portugal. Gaia 10:5-14.
91. Sereno, P. C. 1991. Lesothosaurus, "fabrosaurids" and the early evolution of Ornithischia. Journal of Vertebrate Paleontology 11:168-197.
92. Sereno, P. C. 1998. A rationale for phylogenetic definitions, with application to the higher-level phylogeny of Dinosauria. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 210: 41-83.
93. Sereno, P. C. 1999. The evolution of dinosaurs. Science 284:2137-2147.
94. Sereno, P. C., and A. B. Arcucci. 1993. Dinosaurian precursors from the Middle Triassic of Argentina: Largerpeton chanarensis. Journal of Vertebrate Paleontology 13:385-399.
95. Sereno, P. C., and A. B. Arcucci. 1994. Dinosaurian precursors from the Middle Triassic of Argentina: Marasuchus lilloensis, gen. nov. Journal of Vertebrate Paleontology 14:53-73.
96. Sereno, P. C., and F. E. Novas. 1992. The complete skull and skeleton of an early dinosaur. Science 258:1137-1140.
97. Serene, P. C., C. A. Forster, R. R. Rogers, and A. M. Monetta. 1993. Primitive dinosaur skeleton from Argentina and the early evolution of Dinosauria. Nature 361:64-66.
98. Smith, A. G., D. G. Smith, and B. M. Funnell. 1994. Atlas of Mesozoic and Cenozoic coastlines. Cambridge University Press, Cambridge.
99. Sullivan, C., F. A. Jenkins, S. M. Gatesy, and N. H. Shubin. 2003. A functional asessment of hind foot posture in the prosauropod dinosaur Plateosaurus. Journal of Vertebrate Paleontology 23:102A.
100. Swofford, D. L. 2001. PAUP*: Phylogenetic Analysis Using Parsimony (*and other methods), Version 4.07b. Sinauer, Sunderland, Mass.
101. Tang, F. X. Jin, X. Kang, and G. Zhang. 2001. Omeisaurus maoianus, a complete Sauropoda from Jingyan, Sichuan. Research works of Natural Museum of Zhejiang, China Ocean Press, Beijing. [In Chinese with extended English abstract.]
102. Thulborn, R. A. 1990. Dinosaur tracks. Chapman and Hall, London.
103. Upchurch, P. 1995. The evolutionary history of sauropod dinosaurs. Philosophical Transactions of the Royal Society of London B 349:365-390.
104. Upchurch, P. 1998. The phylogenetic relationships of sauropod dinosaurs. Zoological Journal of the Linnean Society 124:43-103.
105. Upchurch, P., C. A. Hunn, and D. B. Norman. 2002. An analysis of dinosaurian biogeography: evidence for the existence of vicariance and dispersal patterns caused by geological events. Proceedings of the Royal Society of London B 269:613-621.
106. Upchurch, P., P. M. Barrett, and P. Dodson. 2004. Sauropoda. Pp. 259-322 in D. B. Weishampel, P. Dodson, and H. Osmólska, eds. The Dinosauria, 2d ed. University of California Press, Berkeley.
107. van Heerden, J. 1979. The morphology and taxonomy of Euskelosaurus (Reptilia: Saurischia; Late Triassic) from South Africa. Navorsinge Nasionale Museum, Bloemfontein 4:21-84.
108. van Heerden, J. 1997. Prosauropods. Pp. 242-263 in J. O. Farlow and M. K. Brett-Surman, eds. The complete dinosaur. Indiana University Press, Bloomington.
109. van Heerden, J., and P. M. Galton. 1997. The affinities of Melanorosaurus - a Late Triassic prosauropod dinosaur from South Africa. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 1997:39-55.
110. Weems, R. E. 1987. A Late Triassic footprint fauna from the Culpepper Basin Northern Virginia (U.S.A.). Transactions of the American Philosophical Society 77:1-79.
111. Welman, J. 1999. The basicranium of a basal prosauropod from the Euskelosaurus range zone and thoughts on the origin of dinosaurs. Journal of African Earth Sciences 29:227-232.
112. Wilson, J. A. 2002. Sauropod dinosaur phylogeny: critique and cladistic analysis. Zoological Journal of the Linnean Society 136:217-276.
113. Wilson, J. A., and M. T. Carrano. 1999. Titanosaurs and the origin of "wide-gauge" trackways: a biomechanical and systematic perspective on sauropod locomotion. Paleobiology 25: 252-267.
114. Wilson, J. A., and P. C. Sereno. 1998. Early evolution and higher-level phylogeny of sauropod dinosaurs. Society of Vertebrate Paleontology Memoir 5:1-68 (Suppl. to Journal of Vertebrate Paleontology 18).
115. Wilson, J. A., and P. Upchurch. 2003. A revision of Titanosaurus Lydekker (Dinosauria - Sauropoda), the first dinosaur genus with a 'Gondwanan' distribution. Journal of Systematic Palaeontology 1:125-60.
116. Yadagiri, P. 2001. The osteology of Kotasaurus yamanpalliensis, a sauropod dinosaur from the Early Jurassic Kota Formation of India. Journal of Vertebrate Paleontology 21:242-252.
117. Yates, A. M. 2001. A new look at Thecodontosaurus and the origin of sauropod dinosaurs. Journal of Vertebrate Paleontology 21: 116A.
118. Yates, A. M. 2003. A new species of the primitive dinosaur, Thecodontosaurus (Saurischia: Sauropodomorpha) and its implications for the systematics of early dinosaurs. Journal of Systematic Palaeontology 1:1-42.
119. Yates, A. M. 2004. Anchisaurus polyzelus Hitchcock: the smallest known sauropod dinosaur and the evolution of gigantism amongst sauropodomorph dinosaurs. Postilla 230:1-58.
120. Yates, A. M., and J. W. Kitching. 2003. The earliest known sauropod dinosaur and the first steps towards sauropod evolution. Proceedings of the Royal Society of London B 270:1753-1758.
121. Yates, A. M., P. J. Hancox, and B. S. Rubidge. 2004. First record of a sauropod dinosaur from the upper Elliot Formation (Early Jurassic) of South Africa. South African Journal of Science 100:504-506.
122. Zhang, Y. 1988. The Middle Jurassic dinosaur fauna from Dashanpu, Zigong, China, Vol. III. Sauropod dinosaur (I). Shunosaurus. Sichuan Publishing House of Science and Technology, Chengdu. [In Chinese with extended English summary.]