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note
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We thank M. M. Poo, R. H. Masland, and W. R. Levick for insightful comments, and for going through the manuscript several times to improve style and grammar. S.H. is supported by a "Hundreds Talents Plan" of the Chinese Academy of Sciences, a MOST Major State Basic Research program grant (G2000077800), and an NSFC Outstanding Young Researcher Award (39925010).
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