End-Permian mass extinctions: A review

Special Paper of the Geological Society of America

Volumn 356, Issue , 2002, Pages 363-383

  Erwin, Douglas H ^a   Bowring, Samuel A ^b   Yugan, Jin ^c  

^a DEPARTMENT OF PALEOBIOLOGY   (United States)

^b MASSACHUSETTS INSTITUTE OF TECHNOLOGY   (United States)

^c NANJING INSTITUTE OF GEOLOGY AND PALAEONTOLOGY   (China)

Author keywords

[No Author keywords available]

Indexed keywords

ACID RAIN; FLOODS; GEOCHRONOLOGY; GLOBAL WARMING;

CLIMATIC EFFECTS; CONTINENTAL FLOOD BASALT; FILTER FEEDERS; FLOOD BASALT; MASS EXTINCTION; SHALLOW WATERS; SOUTHERN CHINA; SUSPENSION FEEDER;

BASALT;

ANOXIA; CARBONATE; CATASTROPHIC EVENT; FLOOD BASALT; GEOCHRONOLOGY; MASS EXTINCTION; PALEONTOLOGY; PALEOZOIC; PERMIAN;

CHINA;

EID: 84871940772     PISSN: 00721077     EISSN: None     Source Type: Book Series    
DOI: 10.1130/0-8137-2356-6.363     Document Type: Article

References (175)

1. Abbas, S., Abbas, A., and Mohanty, S., 2000, Anoxia during the Late Permian binary mass extinction: Current Science, v. 78, p. 1290-1292.
2. Alt, D., Sears, J.W., and Hyndman, D.W., 1988, Terrestrial maria: The origins of large basalt plateaus, hotspot tracks and spreading ridges: Journal of Geology, v. 96, p. 647-662.
3. Archibold, N.W., and Shi Guanrong, 1996, Western Pacific Permian marine invertebrate palaeobiogeography: Australian Journal of Earth Sciences, v. 43, p. 635-641.
4. Arndt, N., 2000, Hot heads and cold tails: Nature v. 407, p. 458-460.
5. Asaro, F., Alvarez, L.W., Alvarez, W., and Michel, H.V., 1982, Geochemical anomalies near the Eocene/Oligocene and Permian/Triassic boundaries, in Silver, L.T., and Schultz, P.H., eds., Geological implications of impact hypothesis of large asteroids and comets on the earth: Geological Society of America Special Paper 190, p. 517-528.
6. Basu, A.R., Poreda, R.J., Renne, P.R., Teichmann, F., Vasiliev, Y.R., Sobolev, N.V., and Turrin, B.D., 1995, High-3He plume origin and temporal-spatial evolution of the Siberian flood basalts: Nature, v. 269, p. 822-825.
7. Batten, R.L., 1973, The vicissitudes of the Gastropoda during the interval of Guadalupian-Ladinian time, in Logan, A., and Hills, L.V., eds., The Permian and Triassic systems and their mutual boundary: Calgary, Canadian Society of Petroleum Geologists, Memoir 2, p. 596-607.
8. Baud, A., Magaritz, M., and Holser,W.T., 1989, Permian-Triassic of the Tethys: Carbon isotope studies: Sonderdruck aus Geologische Rundschau, v. 78, p. 649-677.
9. Becker, L., Poreda, R.J., Hunt, A.G., Bunch, T.E., and Rampino, M., 2001, Impact event at the Permian-Triassic boundary: Evidence from extraterrestrial noble gases in fullerenes: Science, v. 291, p. 1530-1533.
10. Becker, L., and Poreda, R.J., 2001, An extraterrestrial impact at the Permian-Triassic Boundary?: Reply: Science, v. 293, p. 2343a.
11. Bowring, S.A., Erwin, D.H., Jin Yugan, Martin, M.W., Davidek, K.L., and Wang Wei, 1998, U/Pb zircon geochronology and tempo of the end-Permian mass extinction: Science, v. 280, p. 1039-1045.
12. Bowring, S.A., Erwin, D.H., and Isozaki, Y., 1999, The tempo of mass extinction and recovery: The end-Permian example: Proceedings of the National Academy of Sciences of the United States of America, v. 96, p. 8827-8828.
13. Brack, P., Mundil, R., Oberli, F., Meiser, M., and Rieber, H., 1996, Biostratigraphic and radiometric age data question the Milankovitch characteristics of the Latemar cycles (southern Alps, Italy): Geology, v. 24, p. 371-375.
14. Broecker, W.S., and Peacock, S., 1999, An ecologic explanation for the Permo-Triassic carbon and sulfur isotope shifts: Global Biogeochemical Cycles, v. 13, p. 1167-1172.
15. Campbell, I.H., Czamanske, G.K., Fedorenko, V.A., Hill, R.I., and Stepanov, V., 1992, Synchronism of the Siberian traps and the Permian-Triassic boundary: Science, v. 258, p. 1760-1763.
16. Chen Zhonqiang, Jin Yugan and Shi Guanrong, 1998, Permian transgressionregression sequences and sea-level changes of south China: Proceedings of the Royal Society of Victoria, v. 110, p. 345-367.
17. Chijiwa, T., Arai, T., Sugai, T., Shinohara, H., Kumazawa, M., Takano, M., and Kawakami, S., 1999, Fullerenes found in the Permo-Triassic mass extinction period: Geophysical Research Letters, v. 26, p. 767-770.
18. Claoue-Long, J.C., Zhang Zichao, Ma Guogan, and Du Shaohua, 1991, The age of the Permian-Triassic boundary: Earth and Planetary Science Letters, v. 105, p. 182-190.
19. Clark, D.J., Wang, C.Y., Orth, C.J., and Gilmore, J.S., 1986, Conodont survivaland low iridium abundances across the Permian-Triassic boundary in South China: Science, v. 233, p. 984-986.
20. Courtillot, V., Jaupart, C. Manighetti, I., Tapponnier, P., and Besse, J., 1999, On causal links between flood basalts and continental breakup: Earth and Planetary Science Letters, v. 166, p. 177-195.
21. Czamanske, G.K., Gurevitch, A.B., Fedorenko, V., and Simonov, O., 1998, Demise of the Siberian plume: Paleogeographic and paleotectonic reconstruction from the prevolcanic and volcanic record, north-central Siberia: International Geological Review, v. 40, p. 95-115.
22. Dahl, J., Maldowan, J.M., and Sundararaman, P., 1993, Relationship of biomarker distribution to depositional environment: Organic Geochemistry, v. 20, p. 1001-1017.
23. de Wit, M.J., Ghosh, J.G., de Villiers, S., Rakotosolofo, N., Alexander, J., Tripathi, A., and Looy, C., 2002. Multiple organic carbon isotope reversals across the Permo-Triassic boundary of Terrestrial Gondwanan sequences: Clues to extinction patterns and delayed ecosystem recovery. Journal of Geology, v. 110, p. 227-240.
24. Dickins, J.M., 1983, Permian to Triassic changes in life: Memoirs of the Association of Australasian Paleontologists, v. 1, p. 297-303.
25. Dickins, J.M., Yang Zunyi, Yin Hongfu, Lucas, S.G., and Acharyya, S.K., 1997, Late Palaeozoic and Early Mesozoic circum-Pacific events and their global correlation: Cambridge, Cambridge University Press, 245 p.
26. Erwin, D.H., 1990, Carboniferous-Triassic gastropod diversity patterns and the Permo-Triassic mass extinction: Paleobiology, v. 16, p. 187-203.
27. Erwin, D.H., 1993, The great Paleozoic crisis: New York, Columbia University Press, 327 p.
28. Erwin, D.H., 1994, The Permo-Triassic extinction: Nature, v. 367, p. 231-236.
29. Erwin, D.H., 1996, Understanding biotic recoveries: Extinction, survival and preservation during the End-Permian mass extinction, in Jablonski, D., Erwin, D.H., and Lipps, J.H., eds., Evolutionary paleobiology: Chicago, University of Chicago Press, p. 398-418.
30. Erwin, D.H., 1998, The end and the beginning: Recoveries from mass extinctions: Trends in Ecology and Evolution, v. 13, p. 344-349.
31. Fan Jiasong, Rigby, J.K, Qi Jingwen, 1990, The Permian reefs of South China and comparisons with the Permian Reef Complex of the Guadalupe Mountains, West Texas and New Mexico: Brigham Young University Geological Studies, v. 36, p. 15-55.
32. Farley, K.A., and Mukhopadhyay, S., 2001, An extraterrestrial impact at the Permian-Triassic Boundary?: Comment: Science, v. 293, p. 2343a.
33. Faure, K., de Wit, M.J., and Wills, J.P., 1995, Late Permian global coal hiatus linked to 13C-depleted CO2 flux into the atmosphere during the final consolidation of Pangea: Geology, v. 23, p. 507-510.
34. Fedorenko, V., Czamnske, G., Zen'ko, T., Budahn, J., and Siems, D., 2000, Field and geochemical studies of the melilite-bearing Arydzhangsky Suite, and an overall perspective on the Siberian alkaline-ultramafic flood basalt volcanic rocks: International Geology Review, v. 42, p. 769-804.
35. Flügel, E., 1994, Pangean shelf carbonates: Controls and paleoclimatic significance of Permian and Triassic reefs, in Klein, G.D., ed., Pangea: Paleoclimate, tectonics and sedimentation during accretion, zenith, and breakup of a supercontinent: Geological Society of America Special Paper 288, p. 247-266.
36. Flügel, E., and Reinhardt, J., 1989, Uppermost Permian reefs in Skyros (Greece) and Sichuan (China): Implications for the Late Permian mass extinction event: Palaios, v. 4, p. 502-518.
37. Foster, C.B., Logan, G.A., and Summons, R.E., 1998, The Permian-Triassic boundary in Australia: Where is it and how is it expressed?: Proceedings of the Royal Society of Victoria, v. 110, p. 247-266.
38. Gilkson, A.Y., 1999, Oceanic mega-impacts and crustal evolution: Geology, v. 27, p. 387-390.
39. Golonka, J., and Bocharova, N.Y., 2000, Hot spot activity and the break-up of Pangea: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 161, p. 49-69.
40. Grotzinger, J.P., and Knoll, A.H., 1995, Anomalous carbonate precipitates: Is the Precambrian the key to the past?: Palaios, v. 10, p. 578-596.
41. Gruszczynski, M., Halas, S. Hoffman, A., and Malkowski, K., 1989, A brachiopod calcite record of the oceanic carbon and oxygen level isotope shifts at the Permian/Triassic transition: Nature, v. 337, p. 64-68.
42. Hallam, A., 1991, Why was there a delayed radiation after the end-Palaeozoic extinctions?: Historical Biology, v. 5, p. 257-262.
43. Hallam, A., 1994, The earliest Triassic as an anoxic event, and its relationship to the end-Paleozoic mass extinction, in Embry, A.F., Beauchamp, B., and Glass, D.J., eds., Pangea: Global environments and research: Canadian Society of Petroleum Geologists Memoir 17, p. 797-804.
44. Hallam, A., and Wignall, P.B., 1997, Mass extinctions and their aftermath: Oxford, Oxford University Press, 320 p.
45. Hallam, A., and Wignall, P.B., 1999, Mass extinctions and sea-level changes: Earth-Science Reviews, v. 48, p. 217-250.
46. Holser,W.T., and Magaritz, M., 1987, Events near the Permian-Triassic boundary: Modern Geology, v. 11, p. 155-180.
47. Holser, W.T., Schönlaub, H.P., Attrep, M., Jr., Boeckelmann, K., Klein, P., Magaritz, M., and Orth, C.J., 1989, A unique geochemical record at the Permian/Triassic boundary: Nature, v. 337, p. 39-44.
48. Hotinski, R.M., Bice, K.L., Kump, L.R., Najjar, R.G., and Arthur, M.A., 2001, Ocean stagnation and end-Permian anoxia: Geology, v. 29, p. 7-10.
49. Hotton, N., III, 1967, Stratigraphy and sedimentation in the Beaufort Series (Permian-Triassic), South Africa, in Teichert, C., and Yochelson, E.L., eds., Essays in paleontology and stratigraphy, R.C. Moore Commemorative Volume: Lawrence, University of Kansas Press, p. 390-428.
50. Isbell, J.L. and Askin, R.A., 1999, Search for impact at the Permian-Triassic boundary in Antarctica and Australia: Comment: Geology, v. 27, p. 859.
51. Isozaki, Y., 1995, Superanoxia across the Permo-Triassic boundary: Record in accreted deep-sea pelagic chert in Japan, in Embry, A.F., Beauchamp, B., and Glass, D.J., eds., Pangea: Global environments and research: Canadian Society of Petroleum Geologists Memoir 17, p. 805-812.
52. Isozaki, Y., 1997, Permo-Triassic boundary superanoxia and stratified superocean: Records from lost deep sea: Science, v. 276, p. 235-238.
53. Isozaki, Y., 2001, An extraterrestrial impact at the Permian-Triassic Boundary?: Comment: Science, v. 293, p. 2344a.
54. Jablonski, D., 1986, Causes and consequences of mass extinctions, a comparative approach, in Elliot, D.K., ed., Dynamics of extinction: Chichester, Wiley, p. 183-229.
55. Jin Yugan, Wardlaw, B.R., Glenister, B.F., and Kotlyar, G.V., 1997, Permian chronostratigraphic subdivisions: Episodes, v. 20, no. 1., p. 6-10.
56. Jin Yugan, Wang Wei, Wang Yue, and Cao Changqun, 1998, Prospects for global correlation of Permian sequences: Proceedings of the Royal Society of Victoria, v. 111, p. 73-83.
57. Jin Yugan, Wang Yue, Wang Wei, Shang Qinghua, Cao Changqun, and Erwin, D.H., 2000a, Pattern of marine mass extinction near the Permian-Triassic boundary in South China: Science, v. 289, p. 432-436.
58. Jin Yugan, Shang Qinghua, and Cao Changqun, 2000b, Late Permian magnetostratigraphy and its global correlation: Chinese Science Bulletin, v. 45, p. 698-704.
59. Jin Yugan, Zhang, J., and Shang Qinghua, 1994, Two phases of the end-Permian mass extinction, in Embry, A.F., Beauchamp, B., and Glass, D.J., eds., Pangea: Global environments and resources: Canadian Society of Petroleum Geologists Memoir 17, p. 813-822.
60. Kaiho, K., Kajiwara, Y., Nakano, T., Miura, Y., Kawahata, H., Tazaki, K., Ueshima, M., Chen Z.Q., and Shi, G.R., 2001, End-Permian catastrophe by a bolide impact: Evidence of a gigantic release of sulfur from the mantle: Geology, v. 29, p. 815-818.
61. Kajiwara, Y., Yamakita, S., Ishida, K., Ishiga, H., and Imai, A., 1994, Development of a largely anoxic stratified ocean and its temporary mixing at the Permian/Triassic boundary supported by the sulfur isotope record: Paleaeogeography, Palaeoecology, Palaeoecology, v. 111, p. 367-379.
62. Kakuwa, Y., 1996, Permian-Triassic mass extinction event recorded in bedded chert sequence in southwest Japan: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 121, p. 35-51.
63. Kamo, S.L., Czamanske, G.K., and Krough, T.E., 1996, A minimum U-Pb agefor Siberian flood basalt volcanism: Geochimica et Cosmochimica Acta, v. 60, p. 3505-3511.
64. Kamo, S.L., Czamnske, G.K., Amelin, Y., Fedorenko, V.A., and Trofimov,V.R., 2000, U-Pb zircon and baddeleyite and U-Th-Pb perovskite ages from Siberian flood volcanism, Maymecha-Kotuy area, Siberia: Goldschmidt 2000, Oxford, UK, European Association for Geochemistry and the Geochemical Society, Journal of Conference Abstracts, v. 5, no. 2, p. 569.
65. Kershaw, S., Zhang Tingshan, and Lan Guangzhi, 1999, A ?microbialite carbonate crust at the Permian-Triassic boundary in South China, and its palaeoenvironmental significance: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 146, p. 1-18.
66. King, G.M., 1991, Terrestrial tetrapods and the end Permian event: A comparison of analyses: Historical Biology, v. 5, p. 239-255.
67. King, G.M., and Jenkins, I., 1997, The dicynodont Lystrosaurus from the Upper Permian of Zambia: Evolutionary and stratigraphic implications: Palaeontology, v. 40, p. 149-156.
68. Knoll, A.H., Bambach, R.K., Canfield, D.E., and Grotzinger, J.P., 1996, Comparative earth history and Late Permian mass extinction: Science, v. 273, p. 452-457.
69. Kozur, H.W., 1998, Some aspects of the Permian-Triassic boundary (PTB) and of the possible causes for the biotic crisis around this boundary: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 143, p. 227-272.
70. Krull, E.S., Retallack, G.J., Campbell, H.J., and Lyon G.L. 2000, 13Corg chemostratigraphy of the Permian-Triassic boundary in the Maitai Group, New Zealand: Evidence for high-latitudinal methane release: New Zealand Journal of Geology and Geophysics, v. 43, p. 21-32.
71. Krull, E.S., and Retallack, G.J., 2000, 13C depth profiles from paleosols across the Permian-Triassic boundary: Evidence for methane release: Geological Society of America Bulletin, v. 112, p. 1459-1472.
72. Lehrmann, D.J., 1999, Early Triassic calcimicrobial mounds and biostromes of the Nanpanjiang Basin, South China: Geology, v. 27, p. 359-362.
73. Lind, E.N., Kropotov, S.V., Czamanske, G.K., Gromme, S.C., and Fedorenko, V.A., 1994, Paleomagnetism of the Siberian flood basalts of the Noril'sk area: A constrain on eruption duration: International Geological Review, v. 36, p. 1139-1150.
74. Looy, C.V., Brygman,W.A., Dilcher, D.L., and Visscher, H., 1999, The delayed resurgence of equatorial forests after the Permian-Triassic ecologic crisis: Proceedings of the National Academy of Sciences of the United States of America, v. 96, p. 13857-13862.
75. Lozovsky, V.R., 1998, The Permian-Triassic boundary in the continental series of Eurasia: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 143, p. 273-283.
76. Lucas, S.G. and Yin, H.F., editors, 1998, The Permian-Triassic boundary and global correlations: Palaeogeography, Palaeoclimatology, Palaeoecology, Special Volume 143, no. 4, p. 195-384.
77. MacLeod, K.G., Smith, R.M.H., Koch, P.L., and Ward, P.D., 2000, Timing of mammal-like reptile extinctions across the Permian-Triassic boundary in South Africa: Geology, v. 28, p. 227-230.
78. Magaritz, M., Krishnamurthy, R.V., and Holser, W.T., 1992, Parallel trends in organic and inorganic isotopes across the Permian/Triassic boundary: American Journal of Science, v. 292, p. 727-739.
79. Marshall, C.R., 1994, Confidence intervals on stratigraphic ranges: Partial relaxation of the assumption of randomly distributed fossil horizons: Paleobiology, v. 20, p. 459-469.
80. Marshall, C.R., and Ward, P.D., 1996, Sudden and gradual molluscan extinctions in the latest Cretaceous of western European Tethys: Science, v. 274, p. 1360-1363.
81. Martin, E.E., and Macdougall, J.D., 1995, Sr and Nd isotopes at the Permian/ Triassic boundary: A record of climate change: Chemical Geology, v. 125, p. 73-95.
82. Marzoli, A.R., Renne, P., Piccirillo, E.M., Ernesto, M., Bellienni, G., and De Min, A., 1999, Extensive 200-million-year-old continental flood basalts of the Central Atlantic Magmatic Province: Science, v. 284, p. 616-618.
83. McLoughlin, S., Lindstrom, S., and Drinnan, A.N., 1997, Gondwanan floristicand sedimentological trends during the Permian-Triassic transition: New evidence from the Amery Group, northern Prince Charles Mountains, East Antarctica: Antarctic Science, v. 9, p. 281-298.
84. Mei, S.L., Zhang, K., and Wardlaw, B.R., 1998, A refined succession of Chanhsingian and Griesbachian neogondolellid conodonts from the Meishan section, candidate of the global stratotype section and point of the Permian-Triassic boundary: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 143, p. 213-226.
85. Meldahl, K.H., 1990, Sampling, species abundance, and the stratigraphic signature of mass extinction: A test using Holocene tidal flat Mollusca: Geology, v. 18, p. 890-893.
86. Melosh, H.J., 2000, Can impacts induce volcanic eruptions?, in Catastrophic events and mass extinctions: Impacts and beyond: Houston, Texas, Lunar and Planetary Institute, LPI Contribution No. 1053, p. 141-142.
87. Menning, M., 1995, A numerical time scale for the Permian and Triassic Periods: An integrated analysis, in Scholle, P.A., Peryt, T.M., and Ulmer-Scholle, D.S., eds., The Permian of northern Pangea: Berlin, Springer-Verlag, v. 1, p. 77-97.
88. Menning, M., and Jin Yugan, 1998, Permo-Triassic magnetostratigraphy in China: The type section near Taiyuan, Shanxi Province, North China: Comment: Geophysics Journal International, v. 133, p. 213-216.
89. Mertmann, D., 2000, Sequence stratigraphic subdivision of the Permian Zaluch Group (Salt Range and Trans Indus Ranges, Pakistan): International Geological Congress, 31st, Rio de Jainero, Brazil, Abstracts, CD-ROM.
90. Mii, H.S., Grossman, E.L., and Yancey, T.E., 1997, Stable carbon and oxygen isotopic shifts in Permian seas of West Spitsbergen: Global change or diagenetic artifact: Geology, v. 25, p. 227-230.
91. Min, K., Mundil, R., Renne, P.R., and Ludwig, K.R., 2000, A test for systematic errors in 40Ar/39Ar geochronology through comparison with U/Pb analysis of a 1.1-Ga rhyolite: Geochimica et Cosmochimica Acta, v. 64, p. 73-98.
92. Min, K., Renne, P.R., and Huff, W.D., 2001, 40Ar/39Ar dating of Ordovician K-bentonites in Laurentian and Baltoscandia: Earth and Planetary Science Letters, v. 185, p. 121-134.
93. Morante, R., 1996, Permian and early Triassic isotopic records of carbon and strontium in Australia and a scenario of events about the Permian-Triassic boundary: Historical Biology, v. 11, p. 289-310.
94. Morgan, W.J., 1981, Hot spot tracks and the opening of the Atlantic and Indian Oceans, in Emiliani, C., ed., The oceanic lithosphere: New York, Wiley, The Sea, v. 7, p. 443-487.
95. Mory, A.J., Lasky, R.P., Gilkson, A.Y., and Pirajno, F., 2000a, Woodleigh, Carnarvon Basin, Western Australia: A new 120 km diameter impact structure: Earth and Planetary Science Letters, v. 177, p. 119-128.
96. Mory, A.J., Lasky, R.P., Gilkson, A.Y., and Pirajno, F., 2000b, Response to "Critical comment on: A.J. Mory, et al., Woodleigh, Carnarvon Basin, Western Australia: A new 120 km diameter impact structure" by W.U. Reimold and C. Koeberl: Earth and Planetary Science Letters, v. 184, p. 359-365.
97. Mundil, R., Metacalfe, I., Ludwig, K.R. Renne, P.R. Oberli, F., and Nicoll, R.S., 2001, Timing of the Permian-Triassic biotic crisis: Implications from new zircon U/Pb age data (and their limitations): Earth and Planetary Science Letters, v. 187, p. 131-145.
98. Musashi, M., Isozaki, Y., Koike, T., and Krueulen, R., 2001, Stable carbon isotope signature in mid-Panthalassa shallow-water carbonates across the Permo-Triassic boundary: Evidence for 13C-depleted superocean: Earth and Planetary Science Letters, v. 191, p. 9-20.
99. Newell, A.J., Tverdokhlebov, V.P., and Benton, M.J., 1999, Interplay of tectonics and climate on a transverse fluvial system, Upper Permian, southern Uralian foreland basin, Russia: Sedimentary Geology, v. 127, p. 11-29.
100. Newell, N.D., 1967, Revolutions in the history of life, in Albritton, C.C., Jr., ed., Uniformity and simplicity: Geological Society of America Special Paper 89, p. 63-91.
101. Ogg, J.G., and Steiner, M.B., 1991, Early Triassic magnetic polarity time scaleintegration of magnetostratigraphy, ammonite zonation and sequence stratigraphy from stratotype sections (Canadian Arctic archipelago): Earth and Planetary Science Letters, v. 107, p. 69-89.
102. Orchard, M.J., and Krystyn, L., 1998, Conodonts of the lowermost Triassic of Spiti, and a new zonation based on Neogondolella successions: Rivista Italiana di Paleontologia e Stratigrafia, v. 104, p. 341-368.
103. Orth, C.J., 1989, Geochemistry of the bio-event horizons, in Donovan, S., ed., Mass extinction: Processes and evidence: London, Belkhaven Press, p. 37-72.
104. Orth, C.J., Attrep, M., Jr., and Quintana, L.R., 1990, Iridium anomalies: Abundance patterns across bio-event horizons in the fossil record, in Sharpton, V.L., and Ward, P.D., eds., Global catastrophes in Earth history: Geological Society of America Special Paper 247, p. 45-60.
105. Ouyang Shu, and Utting, J., 1990, Palynology of Upper Permian and Lower Triassic rocks, Meishan, Changxing County, Zhejiang Province, China: Review of Palaeobotany and Palynology, v. 66, p. 65-103.
106. Palfy, J., Mortensen, J.K., Carter, E.S., Smith, P.L., and Tipper, H.W., 2000, Timing the end-Triassic mass extinction: First on land, then in the sea?: Geology, v. 28, p. 39-42.
107. Parrish, J.M., 1995, Geologic evidence of Permian climate, in Scholle, P.A., Peryt, T.M., and Ulmer-Scholle, D.S., eds., The Permian of northern Pangea: Berlin, Springer-Verlag, v. 1, p. 53-61.
108. Puffer, J.H., 2001, Contrasting high field strength element contents of continental flood basalts from plume versus reactivated-arc sources: Geology, v. 29, p. 675-678.
109. Rampino, M.R., Prokoph, A., and Adler, A., 2000, Tempo of the end-Permian event: High-resolution cyclostratigraphy at the Permian-Triassic boundary: Geology, v. 28, p. 643-646.
110. Rampino, M.R., and Strothers, R.B., 1988, Flood basalt volcanism during the past 250 million years: Science, v. 241, p. 663-668.
111. Raup, D.M., 1979, Size of the Permo-Triassic bottleneck and its evolutionary implications: Science, v. 206, p. 217-218.
112. Reimold, W.U., and Koeberl, C., 2000, Critical comment on: A.J. Mory et al., Woodleigh, Carnarvon Basin,Western Australia: A new 120 km diameter impact structure: Earth and Planetary Science Letters, v. 184, p. 353-367.
113. Reinhardt, J.W., 1988, Uppermost Permian reefs and Permo-Triassic sedimentary facies from the southeastern margin of Sichuan Basin, China: Facies, v. 18, p. 231-288.
114. Renne, P.R., and Basu, A.R., 1991, Rapid eruption of the Siberian traps flood basalts at the Permo-Triassic boundary: Science, v. 253, p. 176-179.
115. Renne, P.R., Swisher, C.C., Deino, A.L., Karner, D.B., Owens, T.L., and DePaolo, D.J., 1998a, Intercalibration of standards, absolute ages and uncertainties in 40Ar/39Ar dating: Chemical Geology, v. 145, p. 117-152.
116. Renne, P.R., Zhang Zichao, Richards, M.A., Black, M.T., and Basu, A.R., 1995, Synchrony and causal relations between Permian-Triassic boundary crises and Siberian flood volcanism: Science, v. 269, p. 1413-1416.
117. Renne, P.R., Sharp, W.D., and Becker, T.A., 1998b, 40Ar/39Ar dating of langbeinite (K2Mg2(SO4)3) in late Permian evaporites of the Salado Formation, southeastern New Mexico, USA: Proceedings of the Goldschmidt Conference, Abstracts, p. 1253-1254.
118. Renne, P.R., Steiner, M.B., Sharp, W.D., and Ludwig, K.R., 1996, 40Ar/39Ar and U/Pb SHRIMP dating of latest Permian tephras in the Midland Basin, Texas: American Geophysical Union, Fall Meeting, Abstracts, p. F794.
119. Retallack, G.J., 1995, Permian-Triassic life crisis on land: Science, v. 267, p. 77-80.
120. Retallack, G.J., 1997, Earliest Triassic origin of Isoetes and quillwort evolutionary radiation: Journal of Paleontology, v. 71, p. 500-521.
121. Retallack, G.J., 1999, Postapocalyptic greenhouse paleoclimate revealed by earliest Triassic paleosols in the Sydney Basin, Australia: Geological Society of America Bulletin, v. 111, p. 52-70.
122. Retallack, G.J., and Krull, E.S., 1999, Landscape ecological shift at the Permian-Triassic boundary in Antarctica: Australian Journal of Earth Sciences, v. 46, p. 785-812.
123. Retallack, G.J., Seyedolai, A., Krull, E.S., Holser, W.T., Ambers, C.P., andKyte, F.T., 1998, Search for evidence of impact at the Permian-Triassic boundary in Antarctica and Australia: Geology, v. 26, p. 979-982.
124. Retallack, G.J., Veevers, J.J., and Morante, R., 1996, Global coal gap between Permian-Triassic extinction and Middle Triassic recovery of peat-forming plants: Geological Society of America Bulletin, v. 108, p. 195-207.
125. Riebesell, U., Zondervan, I., Rost, B., Tortell, P.D., Zeebe, R.E., and Morel, F.M.M., 2000, Reduced calcification of marine plankton in response to increased atmospheric CO2: Nature, v. 407, p. 364-367.
126. Ross, C.A., and Ross, J.R.P., 1987, Late Paleozoic sea levels and depositional sequences, in Ross, C.A., and Haman, D., eds., Timing and depositional history of eustatic sequences: Constraints on seismic stratigraphy: Cushman Foundation for Foraminiferal Research Special Publication No. 2, p. 137-149.
127. Rubidge, B.S., 1995, Biostratigraphy of the Beaufort Group (Karoo Supergroup): Geological Survey of South Africa, Biostratigraphic Series, no. 1, p. 1-46.
128. Schmitz, M.D., and Bowring, S.A., 2001, U-Pb zircon and titanite systematics of the Fish Canyon Tuff: An assessment of high-precision U-Pb geochronology and its application to young volcanic rocks: Geochimica et Cosmochimica Acta, v. 65, p. 2571-2587.
129. Scholger, R., Mauritsch, H.J., and Bradner, R., Permian-Triassic boundary magnetostratigraphy from the southern Alps (Italy): Earth and Planetary Science Letters, v. 176, p. 495-508.
130. Scholle, P.A., 1995, Carbon and sulfur isotope stratigraphy of the Permian and adjacent intervals, in Scholle, P.A., Peryt, T.M., and Ulmer-Scholle, D.S., eds., The Permian of northern Pangea: Berlin, Springer-Verlag, v. 1, p. 133-149.
131. Schubert, J.K., and Bottjer, D.J., 1995, Aftermath of the Permian-Triassic mass extinction event: Paleoecology of Lower Triassic carbonates in the western USA: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 116, p. 1-39.
132. Sepkoski, J.J., Jr., 1992, A compendium of fossil marine animal families (second edition): Milwaukee Public Museum Contributions in Biology and Geology, no. 83, p. 155.
133. Shao Longyi, Zhang Pengfei, Duo Jianwei, and Shen Shuzhong, 2000, Carbon isotope compositions of the Late Permian carbonate rocks in southern China: Their variations between the Wujiaping and Changxing formations: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 161, p. 179-192.
134. Shen, Jianwei, Kwamura, T., and Yang Wanrong, 1998, Upper Permian coral reef and colonia rugose carals in northwest Huwan, South China; Facies, v. 39, p. 35-65.
135. Shen Shuzhong, and Shi Guirong, 2000, Wuchaipingian (early Lopingian, Permain) global brachiopod palaeobiogeography: A quantitative approach: Paleaeogeography, Palaeoecology, Palaeoecology, v. 162, p. 299-318.
136. Shishkin, M.A., and Rubidge, B.S., 2000, A relict rhinesuchid (Amphibia: Temnospondyli) from the Lower Triassic of South Africa: Palaeontology, v. 43, p. 653-670.
137. Signor, P.W., III, and Lipps, J.H., 1982, Sampling bias, gradual extinction patterns, and catastrophes in the fossil record, in Silver, L.T., and Schultz, P.H., eds., Geological implications of impacts of large asteroids and comets on the Earth: Geological Society of America Special Paper 190, p. 291-296.
138. Smith, R.M.H., 1995, Changing fluvial environments across the Permian-Triassic boundary in the Karoo Basin, South Africa and possible causes of tetrapod extinctions: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 117, p. 81-95.
139. Stanley, S.M., and Yang, X., 1994, A double mass extinction at the end of the Paleozoic Era: Science, v. 266, p. 1340-1344.
140. Steiner, M., Ogg, J., Zhang, Z., and Sun, S., 1989, The late Permian/Early Triassic magnetic polarity time scale and plate motions of South China: Journal of Geophysical Research, v. 94, p. 7343-7363.
141. Sweet, W.C., Yang Zunyi, Dickins, J.M., and Yin Hongfu, eds., 1992, PermoTriassic events in the eastern Tethys: Cambridge, Cambridge University Press, 181 p.
142. Tanton, L.T.E., and Hager, B.H., 2000, Melt intrusion as a trigger for lithospheric foundering and the eruption of the Siberian flood basalts: Geophysical Research Letters, v. 27, p. 3937-3940.
143. Teichert, C., 1990, The Permian-Triassic boundary revisited, in Kauffman, E.G., and Walliser, O.H., eds., Extinction events in Earth history: Berlin, Springer-Verlag. p. 199-238.
144. Thackeray, J.F., van der Merwe, N.J., Lee-Thorp, J.A., Sillen, A., Lanham, J.L., Smith, R., Keyser, A., and Monteiro, P.M.S., 1990, Changes in carbon isotope ratios in the late Permian recorded in therapsid tooth apatite: Nature, v. 347, p. 751-753.
145. Tong Jinnan, Yin Hongfu, and Zhang Kexing, 1999, Permian and Triassic sequence stratigraphy and sea level changes of eastern Yangtze platform: Journal of China University of Geosciences, v. 10, p. 161-169.
146. Toomey, D.F., 1991, Late Permian reefs of southern Tunisia: Facies patterns and comparison with the Capitan Reef, southwestern United States: Facies, v. 25, p. 119-146.
147. Tosk, T.A., and Andersson, K.A., 1988, Late Early Triassic foraminifers from possible dysaerobic to anaerobic paleoenvironments of the Thaynes Formation, southeast Idaho: Journal of Foraminiferal Research, v. 18, p. 286-301.
148. Twitchett, R.J., and Wignall, P., 1996, Trace fossils and the aftermath of the Permo-Triassic mass extinction: Evidence from northern Italy: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 124, p. 137-151.
149. Twitchett, R.J., Looy, C.V., Morante, R., Visscher, H., and Wignall, P., 2001, Rapid and synchronous collapse of marine and terrestrial ecosystems during the end-Permian biotic crisis: Geology, v. 29, p. 351-354.
150. Veevers, J.J., Conaghan, P.J., and Shaw, S.E., 1994, Turning point in Pangean environmental history at the Permian/Triassic (P/Tr) boundary, in Klein, G.D., ed., Pangea: Paleoclimate, tectonics, and sedimentation during accretion, zenith, and breakup of a supercontinent: Geological Society of America Special Paper 288, p. 187-196.
151. Visscher, H., Brinkhuis, H., Dilcher, D.L., Elsik, W.C., Eshet, Y., Looy, C.V., Rampino, M.R., and Traverse, A., 1996, The terminal Paleozoic fungal event: Evidence of terrestrial ecosystem destabilization and collapse: Proceedings of the National Academy of Sciences of the United States of America, v. 93, p. 2155-2158.
152. Wang, K., Geldsetzer, H.H.J., and Krouse, H.R., 1994, Permian-Triassic extinction: Organic 13C evidence from British Columbia, Canada: Geology, v. 22, p. 580-584.
153. Wang Xiang-Dong, and Sugiyama, T., 2000, Diversity and extinction patterns of Permian coral faunas of China: Lethaia, v. 33, p. 285-294.
154. Wang Zhiqiang, 1993, Evolutionary ecosystem of Permian-Triassic red beds in N. China: A historical record of natural global desertification, in Lucas, S.G., and Morales, M., eds., The nonmarine Triassic: Bulletin of the New Mexico Museum of Natural History and Science, v. 3, p. 471-476.
155. Wang Zhiqiang, 2000, Vegetation of the eve of the P-T event in North China and plant survival strategies: An example of Upper Permian refugium in northwestern Shanxi, China: Acta Palaeontologica Sinica, v. 39 (supplement), p. 127-153.
156. Ward, P.D., Montgomery, D.R., and Smith, R., 2000, Altered river morphology in South Africa related to the Permian-Triassic extinction: Science, v. 289, p. 1740-1743.
157. Wignall, P.B., 2001, Large igneous provinces and mass extinction: Earth-Science Reviews, v. 53, p. 1-33.
158. Wignall, P.B., and Benton, M.J., 1999, Lazarus taxa and fossil abundance at times of biotic crisis: Journal of the Geological Society, London, v. 156, p. 453-456.
159. Wignall, P.B., and Hallam, A., 1992, Anoxia as a cause of the Permo-Triassic mass extinction: Facies evidence from northern Italy and the westernUnited States: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 93, p. 21-46.
160. Wignall, P.B., and Hallam, A., 1993, Griesbachian (earliest Triassic) paleoenvironmental changes in the Salt Range, Pakistan and southeast China and their bearing on the Permo-Triassic mass extinction: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 101, p. 215-237.
161. Wignall, P.B., Hallam, A., Lai Xulong, and Yang Fengqing, 1995, Palaeoenvironmental changes across the Permian/Triassic boundary at Shangsi (N. Sichuan, China): Historical Biology, v. 10, p. 175-189.
162. Wignall, P.B., Morante, R., and Newton, R., 1998, The Permo-Triassic transition in Spitsbergen: 13Corg chemostratigraphy, Fe and S geochemistry, facies, fauna and trace fossils: Geological Magazine, v. 135, p. 47-62.
163. Wignall, P.B., and Twitchett, R.J., 1996, Oceanic anoxia and the end Permian mass extinction: Science, v. 272, p. 1155-1158.
164. Wolf-Gladrow, D.A., Riebesell, U., Burkhardt, S., and Bijma, J., 1999, Direct effects of CO2 concentration of growth and isotopic composition of marine plankton: Tellus, v. 51B, p. 461-476.
165. Woods, A.D., and Bottjer, D.J., 2000, Distribution of ammonoids in the Lower Triassic Union Wash Formation (Eastern California): Evidence for paleoceanographic conditions during recovery from the end-Permian mass extinction: Palaios, v. 15, p. 535-545.
166. Woods, A.D., Bottjer, D.J., Mutti, M., and Morrison, J., 1999, Lower
167. Xu Daoyi, Ma Shulang, Chai Zhifang, Mao Xiuying, Sun Yeyin, Zhang Qingweng, and Yang Zunyi, 1985, Abundance variation of Iridium anomalies and trace elements at the Permian/Triassic boundary at Shangsi in China: Nature, v. 314, p. 154-156.
168. Xu Daoyi, Zhang Qinwen, Sun Yeyin, Yan Zheng, Chai Zhifang, and He Jinwen, 1989, Astrogeological events in China: New York, Van Nostrand Reinhold, 264 p.
169. Yang Fengqing andWang Hongmei, 2000, Ammonoid succession model across the Paleozoic-Mesozoic transition in South China, in Yin Hongfu, Dickins, J.M., Shi, G.R., and Tong Jinnan, eds., Permian-Triassic evolution of Tethys and western circum-Pacific: Amsterdam, Elsevier, p. 353-370.
170. Yang Zunyi, Wu Shunbao, Yin Hongfu, Xu Guirong, Zhang Kexing, and Bie Xianmei, 1993, Permo-Triassic events of South China: Geological Publishing House, Beijing, 153 p.
171. Yin Hongfu, Dickins, J.M., Shi Guanrong, and Tong Jinnan, eds., 2000, Permian-Triassic evolution of Tethys and western circum-Pacific: Amsterdam, Elsevier, 392 p.
172. Yin Hongfu and Tong Jinnan, 1998, Multidisciplinary high-resolution correlation of the Permian-Triassic boundary: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 143, p. 199-212.
173. Zhang Kexing, Tong Jinnan, Yin Hongfu, and Wu Shunbao, 1996, Sequence stratigraphy near the Permian-Triassic boundary at Meishan section, South China, in Yin Hongfu, ed., The Palaeozoic-Mesozoic boundary: Candidates of the global stratotype section and point (GSSP) of the Permian-Triassic boundary: Wuhan, China, University of Geosciences Press, p. 57-64.
174. Zhou Lei, and Kyte, F.T, 1988, The Permian-Triassic boundary event: A geochemical study of three Chinese sections: Earth and Planetary Science Letters, v. 90, p. 411-421.
175. Zhou Zuren, Glenister, B.F., Furnish, W.M., and Spinosa, C., 1996, Multiepisodal extinction and ecological differentiation of Permian ammonoids: Permophiles, v. 29, p. 195-212.