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Volumn 179, Issue 2, 2015, Pages 527-535

Concordance and discordance between taxonomic and functional homogenization: responses of soil mite assemblages to forest conversion

Author keywords

Community reassembly; Ecosystem functioning; Forest restoration; Oribatid mites; Soil arthropods; Taxonomic and functional homogenization

Indexed keywords

ARTHROPOD; COMMUNITY DYNAMICS; ECOSYSTEM FUNCTION; FOREST ECOSYSTEM; FUNCTIONAL CHANGE; HOMOGENEITY; LANDSCAPE; MITE; TAXONOMY;

EID: 84941879122     PISSN: 00298549     EISSN: None     Source Type: Journal    
DOI: 10.1007/s00442-015-3342-2     Document Type: Article
Times cited : (26)

References (61)
  • 1
    • 84862328335 scopus 로고    scopus 로고
    • Rethinking the relationship between nestedness and beta diversity: a comment on Baselga (2010)
    • Almeida-Neto M, Frensel DMB, Ulrich W (2012) Rethinking the relationship between nestedness and beta diversity: a comment on Baselga (2010). Glob Ecol Biogeogr 21:772–777
    • (2012) Glob Ecol Biogeogr , vol.21 , pp. 772-777
    • Almeida-Neto, M.1    Frensel, D.M.B.2    Ulrich, W.3
  • 2
    • 78650061319 scopus 로고    scopus 로고
    • Navigating the multiple meanings of beta diversity: a roadmap for the practicing ecologist
    • PID: 21070562
    • Anderson MJ et al (2010) Navigating the multiple meanings of beta diversity: a roadmap for the practicing ecologist. Ecol Lett 14:19–28
    • (2010) Ecol Lett , vol.14 , pp. 19-28
    • Anderson, M.J.1
  • 3
    • 1942476319 scopus 로고
    • Analysis of oribatid communities by relative abundance in the species and individual numbers of the three major groups (MGP analysis)
    • (In Japanese with an English abstract)
    • Aoki J (1983) Analysis of oribatid communities by relative abundance in the species and individual numbers of the three major groups (MGP analysis) (in Japanese with an English abstract). Yokohama Natl Univ Environ Res Rep 10:171–176
    • (1983) Yokohama Natl Univ Environ Res Rep , vol.10 , pp. 171-176
    • Aoki, J.1
  • 5
    • 84923094743 scopus 로고    scopus 로고
    • Belowground biodiversity and ecosystem functioning
    • COI: 1:CAS:528:DC%2BC2cXitFamsrnM, PID: 25428498
    • Bardgett RD, van der Putten WH (2014) Belowground biodiversity and ecosystem functioning. Nature 515:505–511
    • (2014) Nature , vol.515 , pp. 505-511
    • Bardgett, R.D.1    van der Putten, W.H.2
  • 7
    • 72449151245 scopus 로고    scopus 로고
    • Partitioning the turnover and nestedness components of beta diversity
    • Baselga A (2010) Partitioning the turnover and nestedness components of beta diversity. Glob Ecol Biogeogr 19:134–143
    • (2010) Glob Ecol Biogeogr , vol.19 , pp. 134-143
    • Baselga, A.1
  • 8
    • 84857566449 scopus 로고    scopus 로고
    • The relationship between species replacement, dissimilarity derived from nestedness, and nestedness
    • Baselga A (2012) The relationship between species replacement, dissimilarity derived from nestedness, and nestedness. Glob Ecol Biogeogr 21:1223–1232
    • (2012) Glob Ecol Biogeogr , vol.21 , pp. 1223-1232
    • Baselga, A.1
  • 9
    • 84857137957 scopus 로고    scopus 로고
    • Extreme winter warming events more negatively impact small rather than large soil fauna: shift in community composition explained by traits not taxa
    • Bokhorst S, Phoenix GK, Bjerke JW, Callaghan TV, Huyer-Brugman F, Berg MP (2012) Extreme winter warming events more negatively impact small rather than large soil fauna: shift in community composition explained by traits not taxa. Glob Change Biol 18:1152–1162
    • (2012) Glob Change Biol , vol.18 , pp. 1152-1162
    • Bokhorst, S.1    Phoenix, G.K.2    Bjerke, J.W.3    Callaghan, T.V.4    Huyer-Brugman, F.5    Berg, M.P.6
  • 10
    • 80054120424 scopus 로고    scopus 로고
    • Spread of common species results in local-scale floristic homogenization in grassland of Switzerland
    • Bühler C, Roth T (2011) Spread of common species results in local-scale floristic homogenization in grassland of Switzerland. Divers Distrib 17:1089–1098
    • (2011) Divers Distrib , vol.17 , pp. 1089-1098
    • Bühler, C.1    Roth, T.2
  • 11
    • 80052800716 scopus 로고    scopus 로고
    • Beyond species: functional diversity and the maintenance of ecological processes and services
    • Cadotte MW, Carscadden K, Mirotchnick N (2011) Beyond species: functional diversity and the maintenance of ecological processes and services. J Appl Ecol 48:1079–1087
    • (2011) J Appl Ecol , vol.48 , pp. 1079-1087
    • Cadotte, M.W.1    Carscadden, K.2    Mirotchnick, N.3
  • 12
    • 79955613233 scopus 로고    scopus 로고
    • Worldwide decline of specialist species: toward a global functional homogenization?
    • Clavel J, Julliard R, Devictor V (2011) Worldwide decline of specialist species: toward a global functional homogenization? Front Ecol Environ 9:222–228
    • (2011) Front Ecol Environ , vol.9 , pp. 222-228
    • Clavel, J.1    Julliard, R.2    Devictor, V.3
  • 13
    • 36049012711 scopus 로고    scopus 로고
    • From peds to paradoxes: linkages between soil biota and their influences on ecological processes
    • COI: 1:CAS:528:DC%2BD2sXhtlajs7bM
    • Coleman DC (2008) From peds to paradoxes: linkages between soil biota and their influences on ecological processes. Soil Biol Biochem 40:271–289
    • (2008) Soil Biol Biochem , vol.40 , pp. 271-289
    • Coleman, D.C.1
  • 14
    • 65349175908 scopus 로고    scopus 로고
    • Community assembly and shifts in plant trait distributions across an environmental gradient in coastal California
    • Cornwell WK, Ackerly DD (2009) Community assembly and shifts in plant trait distributions across an environmental gradient in coastal California. Ecol Monogr 79:109–126
    • (2009) Ecol Monogr , vol.79 , pp. 109-126
    • Cornwell, W.K.1    Ackerly, D.D.2
  • 15
    • 0035501320 scopus 로고    scopus 로고
    • Vive la différence: plant functional diversity matters to ecosystem processes
    • Díaz S, Cabido M (2001) Vive la différence: plant functional diversity matters to ecosystem processes. Trends Ecol Evol 16:646–655
    • (2001) Trends Ecol Evol , vol.16 , pp. 646-655
    • Díaz, S.1    Cabido, M.2
  • 16
    • 84870910545 scopus 로고    scopus 로고
    • Functional traits explain phytoplankton community structure and seasonal dynamics in a marine ecosystem
    • PID: 23033839
    • Edwards KF, Litchman E, Klausmeier CA, Elser J (2013) Functional traits explain phytoplankton community structure and seasonal dynamics in a marine ecosystem. Ecol Lett 16: 56–63
    • (2013) Ecol Lett , vol.16 , pp. 56-63
    • Edwards, K.F.1    Litchman, E.2    Klausmeier, C.A.3    Elser, J.4
  • 17
    • 84888424224 scopus 로고    scopus 로고
    • Management intensity affects traits of soil microarthropod community in montane spruce forest
    • Farská J, Prejzková K, Rusek J (2014) Management intensity affects traits of soil microarthropod community in montane spruce forest. Appl Soil Ecol 75:71–79
    • (2014) Appl Soil Ecol , vol.75 , pp. 71-79
    • Farská, J.1    Prejzková, K.2    Rusek, J.3
  • 18
    • 33646790192 scopus 로고    scopus 로고
    • Utility and limitations of species richness metrics for conservation planning
    • Fleishman E, Noss R, Noon B (2006) Utility and limitations of species richness metrics for conservation planning. Ecol Ind 6:543–553
    • (2006) Ecol Ind , vol.6 , pp. 543-553
    • Fleishman, E.1    Noss, R.2    Noon, B.3
  • 19
    • 5344221650 scopus 로고    scopus 로고
    • Plant functional markers capture ecosystem properties during secondary succession
    • Garnier E et al (2004) Plant functional markers capture ecosystem properties during secondary succession. Ecology 85:2630–2637
    • (2004) Ecology , vol.85 , pp. 2630-2637
    • Garnier, E.1
  • 20
    • 85003340092 scopus 로고    scopus 로고
    • Functional identity and diversity of animals predict ecosystem functioning better than species-based indices
    • Gagic V et al (2015) Functional identity and diversity of animals predict ecosystem functioning better than species-based indices. Proc Biol Sci 282:2014–2620
    • (2015) Proc Biol Sci , vol.282 , pp. 2014-2620
    • Gagic, V.1
  • 22
    • 0007896159 scopus 로고    scopus 로고
    • Diversity in the decomposing landscape
    • Coleman DC, Hendrix PF, (eds), CABI, New York
    • Hansen RA (2000a) Diversity in the decomposing landscape. In: Coleman DC, Hendrix PF (eds) Invertebrates as webmasters in ecosystems. CABI, New York, p 336
    • (2000) Invertebrates as webmasters in ecosystems , pp. 336
    • Hansen, R.A.1
  • 23
    • 0033938983 scopus 로고    scopus 로고
    • Effects of habitat complexity and composition on a diverse litter microarthropod assemblage
    • Hansen RA (2000b) Effects of habitat complexity and composition on a diverse litter microarthropod assemblage. Ecology 81:1120–1132
    • (2000) Ecology , vol.81 , pp. 1120-1132
    • Hansen, R.A.1
  • 24
    • 84910099809 scopus 로고    scopus 로고
    • The effects of mixed broad-leaved trees on the collembolan community in larch plantations of central Japan
    • Hasegawa M, Ota AT, Kabeya D, Okamoto T, Saitoh T, Nishiyama Y (2013) The effects of mixed broad-leaved trees on the collembolan community in larch plantations of central Japan. Appl Soil Ecol 83: 125–132
    • (2013) Appl Soil Ecol , vol.83 , pp. 125-132
    • Hasegawa, M.1    Ota, A.T.2    Kabeya, D.3    Okamoto, T.4    Saitoh, T.5    Nishiyama, Y.6
  • 25
    • 71749095781 scopus 로고    scopus 로고
    • Novel ecosystems: implications for conservation and restoration
    • PID: 19683830
    • Hobbs RJ, Higgs E, Harris JA (2009) Novel ecosystems: implications for conservation and restoration. Trends Ecol Evol 24:599–605
    • (2009) Trends Ecol Evol , vol.24 , pp. 599-605
    • Hobbs, R.J.1    Higgs, E.2    Harris, J.A.3
  • 26
    • 84980090975 scopus 로고
    • The distribution of the flora in the alpine zone
    • Jaccard P (1912) The distribution of the flora in the alpine zone. New Phytol 11:37–50
    • (1912) New Phytol , vol.11 , pp. 37-50
    • Jaccard, P.1
  • 27
    • 0024186336 scopus 로고
    • Feeding-habits and cheliceral size of oribatid mites in cool temperate forest soils in Japan
    • Kaneko N (1988) Feeding-habits and cheliceral size of oribatid mites in cool temperate forest soils in Japan. Rev Ecol Biol Sol 25:353–363
    • (1988) Rev Ecol Biol Sol , vol.25 , pp. 353-363
    • Kaneko, N.1
  • 28
    • 0033152520 scopus 로고    scopus 로고
    • Mixed leaf litter effects on decomposition rates and soil microarthropod communities in an oak-pine stand in Japan
    • Kaneko N, Salamanca EF (1999) Mixed leaf litter effects on decomposition rates and soil microarthropod communities in an oak-pine stand in Japan. Ecol Res 14:131–138
    • (1999) Ecol Res , vol.14 , pp. 131-138
    • Kaneko, N.1    Salamanca, E.F.2
  • 30
    • 84908473465 scopus 로고    scopus 로고
    • Interpreting the replacement and richness difference components of beta diversity
    • Legendre P (2014) Interpreting the replacement and richness difference components of beta diversity. Glob Ecol Biogeogr 23:1324–1334
    • (2014) Glob Ecol Biogeogr , vol.23 , pp. 1324-1334
    • Legendre, P.1
  • 31
    • 79952505107 scopus 로고    scopus 로고
    • Partitioning global patterns of freshwater fish beta diversity reveals contrasting signatures of past climate changes
    • PID: 21303436
    • Leprieur F et al (2011) Partitioning global patterns of freshwater fish beta diversity reveals contrasting signatures of past climate changes. Ecol Lett 14:325–334
    • (2011) Ecol Lett , vol.14 , pp. 325-334
    • Leprieur, F.1
  • 32
    • 84863632127 scopus 로고    scopus 로고
    • Traits explain community disassembly and trophic contraction following experimental environmental change
    • Lindo Z, Whiteley J, Gonzalez A (2012) Traits explain community disassembly and trophic contraction following experimental environmental change. Glob Change Biol 18:2448–2457
    • (2012) Glob Change Biol , vol.18 , pp. 2448-2457
    • Lindo, Z.1    Whiteley, J.2    Gonzalez, A.3
  • 33
    • 79951680551 scopus 로고    scopus 로고
    • Forest fragmentation drives Atlantic forest of northeastern Brazil to biotic homogenization
    • Lôbo D, Leão T, Melo FPL, Santos AMM, Tabarelli M (2011) Forest fragmentation drives Atlantic forest of northeastern Brazil to biotic homogenization. Divers Distrib 17:287–296
    • (2011) Divers Distrib , vol.17 , pp. 287-296
    • Lôbo, D.1    Leão, T.2    Melo, F.P.L.3    Santos, A.M.M.4    Tabarelli, M.5
  • 34
    • 0000289822 scopus 로고
    • Studies on the oribatid mites of a Danish beech wood soil. I. Nutritional biology
    • Luxton M (1972) Studies on the oribatid mites of a Danish beech wood soil. I. Nutritional biology. Pedobiologia 12:434–463
    • (1972) Pedobiologia , vol.12 , pp. 434-463
    • Luxton, M.1
  • 35
    • 78650676827 scopus 로고    scopus 로고
    • Traits explain the responses of a sub-arctic Collembola community to climate manipulation
    • COI: 1:CAS:528:DC%2BC3MXhtlGmtw%3D%3D
    • Makkonen M, Berg MP, van Hal JR, Callaghan TV, Press MC, Aerts R (2011) Traits explain the responses of a sub-arctic Collembola community to climate manipulation. Soil Biol Biochem 43:377–384
    • (2011) Soil Biol Biochem , vol.43 , pp. 377-384
    • Makkonen, M.1    Berg, M.P.2    van Hal, J.R.3    Callaghan, T.V.4    Press, M.C.5    Aerts, R.6
  • 36
    • 84885838816 scopus 로고    scopus 로고
    • Gains in native species promote biotic homogenization over four decades in a human-dominated landscape
    • McCune JL, Vellend M (2013) Gains in native species promote biotic homogenization over four decades in a human-dominated landscape. J Ecol 101:1542–1551
    • (2013) J Ecol , vol.101 , pp. 1542-1551
    • McCune, J.L.1    Vellend, M.2
  • 37
    • 0033231959 scopus 로고    scopus 로고
    • Biotic homogenization: a few winners replacing many losers in the next mass extinction
    • PID: 10511724
    • McKinney ML, Lockwood JL (1999) Biotic homogenization: a few winners replacing many losers in the next mass extinction. Trends Ecol Evol 14:450–453
    • (1999) Trends Ecol Evol , vol.14 , pp. 450-453
    • McKinney, M.L.1    Lockwood, J.L.2
  • 38
    • 84875810402 scopus 로고    scopus 로고
    • Response diversity determines the resilience of ecosystems to environmental change
    • PID: 23217173
    • Mori AS, Furukawa T, Sasaki T (2013a) Response diversity determines the resilience of ecosystems to environmental change. Biol Rev 88:349–364
    • (2013) Biol Rev , vol.88 , pp. 349-364
    • Mori, A.S.1    Furukawa, T.2    Sasaki, T.3
  • 40
    • 84922011348 scopus 로고    scopus 로고
    • Biotic homogenization and differentiation of soil faunal communities in the production forest landscape: taxonomic and functional perspectives
    • PID: 25322821
    • Mori AS et al (2015) Biotic homogenization and differentiation of soil faunal communities in the production forest landscape: taxonomic and functional perspectives. Oecologia 177:533–544
    • (2015) Oecologia , vol.177 , pp. 533-544
    • Mori, A.S.1
  • 41
    • 84875854174 scopus 로고    scopus 로고
    • Oribatid mites (Acari, Oribatida) from Tohoku (northeast Japan), collected after a tidal wave in 2011
    • Nakamura K, Nakamura YN, Fujikawa T (2013) Oribatid mites (Acari, Oribatida) from Tohoku (northeast Japan), collected after a tidal wave in 2011. Acarologia 53:41–76
    • (2013) Acarologia , vol.53 , pp. 41-76
    • Nakamura, K.1    Nakamura, Y.N.2    Fujikawa, T.3
  • 42
    • 84898458444 scopus 로고    scopus 로고
    • Size matters: implications of the loss of large individuals for ecosystem function
    • PID: 24025973
    • Norkko A, Villnas A, Norkko J, Valanko S, Pilditch C (2013) Size matters: implications of the loss of large individuals for ecosystem function. Sci Rep 3:2646
    • (2013) Sci Rep , vol.3 , pp. 2646
    • Norkko, A.1    Villnas, A.2    Norkko, J.3    Valanko, S.4    Pilditch, C.5
  • 43
    • 31144457908 scopus 로고    scopus 로고
    • On defining and quantifying biotic homogenization
    • Olden JD, Rooney TP (2006) On defining and quantifying biotic homogenization. Glob Ecol Biogeogr 15:113–120
    • (2006) Glob Ecol Biogeogr , vol.15 , pp. 113-120
    • Olden, J.D.1    Rooney, T.P.2
  • 45
    • 84879320466 scopus 로고    scopus 로고
    • Several scales of biodiversity affect ecosystem multifunctionality
    • COI: 1:CAS:528:DC%2BC3sXhtFOmsLzK, PID: 23733963
    • Pasari JR, Levi T, Zavaleta ES, Tilman D (2013) Several scales of biodiversity affect ecosystem multifunctionality. Proc Natl Acad Sci USA 110:10219–10222
    • (2013) Proc Natl Acad Sci USA , vol.110 , pp. 10219-10222
    • Pasari, J.R.1    Levi, T.2    Zavaleta, E.S.3    Tilman, D.4
  • 46
    • 84858063552 scopus 로고    scopus 로고
    • Taxonomic and functional homogenization of an endemic desert fish fauna
    • Pool TK, Olden JD (2012) Taxonomic and functional homogenization of an endemic desert fish fauna. Divers Distrib 18:366–376
    • (2012) Divers Distrib , vol.18 , pp. 366-376
    • Pool, T.K.1    Olden, J.D.2
  • 47
    • 84872572048 scopus 로고    scopus 로고
    • Conversion of the Amazon rainforest to agriculture results in biotic homogenization of soil bacterial communities
    • COI: 1:CAS:528:DC%2BC3sXhs1yhtro%3D, PID: 23271810
    • Rodrigues JL et al (2013) Conversion of the Amazon rainforest to agriculture results in biotic homogenization of soil bacterial communities. Proc Natl Acad Sci USA 110:988–993
    • (2013) Proc Natl Acad Sci USA , vol.110 , pp. 988-993
    • Rodrigues, J.L.1
  • 48
    • 84898463223 scopus 로고    scopus 로고
    • Body size as a predictor of species loss effect on ecosystem functioning
    • PID: 24714619
    • Seguin A, Harvey E, Archambault P, Nozais C, Gravel D (2014) Body size as a predictor of species loss effect on ecosystem functioning. Sci Rep 4:4616
    • (2014) Sci Rep , vol.4 , pp. 4616
    • Seguin, A.1    Harvey, E.2    Archambault, P.3    Nozais, C.4    Gravel, D.5
  • 49
    • 0027331723 scopus 로고
    • Feeding guilds of oribatid mites based on their carbohydrase activities
    • Siepel H, Deruiterdijkman EM (1993) Feeding guilds of oribatid mites based on their carbohydrase activities. Soil Biol Biochem 25:1491–1497
    • (1993) Soil Biol Biochem , vol.25 , pp. 1491-1497
    • Siepel, H.1    Deruiterdijkman, E.M.2
  • 50
    • 51649151268 scopus 로고
    • Dynamics and maintenance of collembolan community structure in a forest soil system
    • Takeda H (1987) Dynamics and maintenance of collembolan community structure in a forest soil system. Res Popul Ecol 29:291–346
    • (1987) Res Popul Ecol , vol.29 , pp. 291-346
    • Takeda, H.1
  • 51
    • 84857916002 scopus 로고    scopus 로고
    • Does taxonomic homogenization imply functional homogenization in temperate forest herb layer communities?
    • Tobias N, Monika W (2011) Does taxonomic homogenization imply functional homogenization in temperate forest herb layer communities? Plant Ecol 213:431–443
    • (2011) Plant Ecol , vol.213 , pp. 431-443
    • Tobias, N.1    Monika, W.2
  • 52
    • 84879148556 scopus 로고    scopus 로고
    • Biotic homogenization at the community scale: disentangling the roles of urbanization and plant invasion
    • Trentanovi G, von der Lippe M, Sitzia T, Ziechmann U, Kowarik I, Cierjacks A (2013) Biotic homogenization at the community scale: disentangling the roles of urbanization and plant invasion. Divers Distrib 19:738–748
    • (2013) Divers Distrib , vol.19 , pp. 738-748
    • Trentanovi, G.1    von der Lippe, M.2    Sitzia, T.3    Ziechmann, U.4    Kowarik, I.5    Cierjacks, A.6
  • 53
    • 0001617782 scopus 로고
    • Bias and confidence in not-quite large samples
    • Tukey JW (1958) Bias and confidence in not-quite large samples. Ann Math Sci 29:614
    • (1958) Ann Math Sci , vol.29 , pp. 614
    • Tukey, J.W.1
  • 54
    • 84887183430 scopus 로고    scopus 로고
    • Weighing in: size spectra as a standard tool in soil community analyses
    • COI: 1:CAS:528:DC%2BC3sXhvFOltbnL
    • Turnbull MS, George PBL, Lindo Z (2014) Weighing in: size spectra as a standard tool in soil community analyses. Soil Biol Biochem 68:366–372
    • (2014) Soil Biol Biochem , vol.68 , pp. 366-372
    • Turnbull, M.S.1    George, P.B.L.2    Lindo, Z.3
  • 55
    • 84925282317 scopus 로고    scopus 로고
    • Functional diversity enhances the resistance of ecosystem multifunctionality to aridity in Mediterranean drylands
    • PID: 25615801
    • Valencia E et al (2015) Functional diversity enhances the resistance of ecosystem multifunctionality to aridity in Mediterranean drylands. New Phytol 206:660–671
    • (2015) New Phytol , vol.206 , pp. 660-671
    • Valencia, E.1
  • 56
    • 84888341917 scopus 로고    scopus 로고
    • Global meta-analysis reveals no net change in local-scale plant biodiversity over time
    • COI: 1:CAS:528:DC%2BC3sXhvFemur3I, PID: 24167259
    • Vellend M et al (2013) Global meta-analysis reveals no net change in local-scale plant biodiversity over time. Proc Natl Acad Sci USA 110:19456–19459
    • (2013) Proc Natl Acad Sci USA , vol.110 , pp. 19456-19459
    • Vellend, M.1
  • 57
    • 84877631794 scopus 로고    scopus 로고
    • Decomposing functional β-diversity reveals that low functional β-diversity is driven by low functional turnover in European fish assemblages
    • Villéger S, Grenouillet G, Brosse S (2013) Decomposing functional β-diversity reveals that low functional β-diversity is driven by low functional turnover in European fish assemblages. Glob Ecol Biogeogr 22:671–681
    • (2013) Glob Ecol Biogeogr , vol.22 , pp. 671-681
    • Villéger, S.1    Grenouillet, G.2    Brosse, S.3
  • 60
    • 84892865765 scopus 로고    scopus 로고
    • Spruce forest conversion to a mixed beech-coniferous stand modifies oribatid community structure
    • Zaitsev AS, Chauvat M, Wolters V (2014) Spruce forest conversion to a mixed beech-coniferous stand modifies oribatid community structure. Appl Soil Ecol 76:60–67
    • (2014) Appl Soil Ecol , vol.76 , pp. 60-67
    • Zaitsev, A.S.1    Chauvat, M.2    Wolters, V.3


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