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Volumn 163, Issue 4, 2011, Pages 1003-1025

Systematics and spicule evolution in dictyonal sponges (Hexactinellida: Sceptrulophora) With description of two new species

Author keywords

Aspidoscopulia; Deep Down Under Expedition; Farreidae; Lonchiphora; Molecular systematics; Porifera; Psilocalyx; Sarostegia; Sceptrules

Indexed keywords

ANTHOZOA; EURETIDAE; FARREIDAE; HEXACTINELLIDA; HEXACTINOSIDA; PORIFERA; PSILOCALYX; TRETODICTYIDAE;

EID: 80255130530     PISSN: 00244082     EISSN: 10963642     Source Type: Journal    
DOI: 10.1111/j.1096-3642.2011.00753.x     Document Type: Article
Times cited : (24)

References (62)
  • 2
    • 65649131928 scopus 로고    scopus 로고
    • Gene tree discordance, phylogenetic inference and the multispecies coalescent
    • Degnan JH, Rosenberg NA. 2009. Gene tree discordance, phylogenetic inference and the multispecies coalescent. Trends in Ecology and Evolution 24: 332-340.
    • (2009) Trends in Ecology and Evolution , vol.24 , pp. 332-340
    • Degnan, J.H.1    Rosenberg, N.A.2
  • 3
    • 67349140912 scopus 로고    scopus 로고
    • New insights into the phylogeny of glass sponges (Porifera, Hexactinellida): monophyly of Lyssacinosida and Euplectellinae, and the phylogenetic position of Euretidae
    • Dohrmann M, Collins AG, Wörheide G. 2009. New insights into the phylogeny of glass sponges (Porifera, Hexactinellida): monophyly of Lyssacinosida and Euplectellinae, and the phylogenetic position of Euretidae. Molecular Phylogenetics and Evolution 52: 257-262.
    • (2009) Molecular Phylogenetics and Evolution , vol.52 , pp. 257-262
    • Dohrmann, M.1    Collins, A.G.2    Wörheide, G.3
  • 5
    • 33947732464 scopus 로고
    • Radiolaria and Porifera (spicula) from the Upper Triassic of Aghdarband (NE-Iran)
    • Donofrio DA. 1991. Radiolaria and Porifera (spicula) from the Upper Triassic of Aghdarband (NE-Iran). Abhandlungen der Geologischen Bundes-Anstalt 38: 205-222.
    • (1991) Abhandlungen der Geologischen Bundes-Anstalt , vol.38 , pp. 205-222
    • Donofrio, D.A.1
  • 6
    • 4344571592 scopus 로고    scopus 로고
    • Three new species and a new genus of Farreidae (Porifera: Hexactinellida: Hexactinosida)
    • Duplessis K, Reiswig HM. 2004. Three new species and a new genus of Farreidae (Porifera: Hexactinellida: Hexactinosida). Proceedings of the Biological Society of Washington 117: 199-212.
    • (2004) Proceedings of the Biological Society of Washington , vol.117 , pp. 199-212
    • Duplessis, K.1    Reiswig, H.M.2
  • 7
    • 77956342397 scopus 로고    scopus 로고
    • Aphyhly: a systematic designation for a taxonomic problem
    • Ebach MC, Williams DM. 2010. Aphyhly: a systematic designation for a taxonomic problem. Evolutionary Biology 37: 123-127.
    • (2010) Evolutionary Biology , vol.37 , pp. 123-127
    • Ebach, M.C.1    Williams, D.M.2
  • 8
    • 58249100073 scopus 로고    scopus 로고
    • Is a new and general theory of molecular systematics emerging?
    • Edwards SV. 2009. Is a new and general theory of molecular systematics emerging? Evolution 63: 1-19.
    • (2009) Evolution , vol.63 , pp. 1-19
    • Edwards, S.V.1
  • 9
    • 33646891565 scopus 로고    scopus 로고
    • CO1 phylogenies in diploblasts and the 'Barcoding of life'- are we sequencing a suboptimal partition?
    • Erpenbeck D, Hooper JNA, Wörheide G. 2006. CO1 phylogenies in diploblasts and the 'Barcoding of life'- are we sequencing a suboptimal partition? Molecular Ecology Notes 6: 550-553.
    • (2006) Molecular Ecology Notes , vol.6 , pp. 550-553
    • Erpenbeck, D.1    Hooper, J.N.A.2    Wörheide, G.3
  • 10
    • 38349149515 scopus 로고    scopus 로고
    • On the molecular phylogeny of sponges (Porifera)
    • Erpenbeck D, Wörheide G. 2007. On the molecular phylogeny of sponges (Porifera). Zootaxa 1668: 107-126.
    • (2007) Zootaxa , vol.1668 , pp. 107-126
    • Erpenbeck, D.1    Wörheide, G.2
  • 11
    • 0000461280 scopus 로고
    • Confidence limits on phylogenies: an approach using the bootstrap
    • Felsenstein J. 1985. Confidence limits on phylogenies: an approach using the bootstrap. Evolution 39: 783-791.
    • (1985) Evolution , vol.39 , pp. 783-791
    • Felsenstein, J.1
  • 12
    • 80052496438 scopus 로고    scopus 로고
    • ANT XXIV/2 (SYSTCO) Hexactinellida (Porifera) and bathymetric traits of Antarctic glass sponges (incorporating ANDEEP-material); including an emendation of the rediscovered genus Lonchiphora
    • Göcke C, Janussen D. 2011. ANT XXIV/2 (SYSTCO) Hexactinellida (Porifera) and bathymetric traits of Antarctic glass sponges (incorporating ANDEEP-material); including an emendation of the rediscovered genus Lonchiphora. Deep-Sea Research Part II 58: 2013-2021.
    • (2011) Deep-Sea Research Part II , vol.58 , pp. 2013-2021
    • Göcke, C.1    Janussen, D.2
  • 13
    • 74549125386 scopus 로고    scopus 로고
    • SeaView version 4: a multiplatform graphical user interface for sequence alignment and phylogenetic tree building
    • Gouy M, Guindon S, Gascuel O. 2010. SeaView version 4: a multiplatform graphical user interface for sequence alignment and phylogenetic tree building. Molecular Biology and Evolution 27: 221-224.
    • (2010) Molecular Biology and Evolution , vol.27 , pp. 221-224
    • Gouy, M.1    Guindon, S.2    Gascuel, O.3
  • 14
    • 34547697011 scopus 로고    scopus 로고
    • Glass sponges and bilaterian animals share derived mitochondrial genomic features: a common ancestry or parallel evolution?
    • Haen KM, Lang BF, Pomponi SA, Lavrov DV. 2007. Glass sponges and bilaterian animals share derived mitochondrial genomic features: a common ancestry or parallel evolution? Molecular Biology and Evolution 24: 1518-1527.
    • (2007) Molecular Biology and Evolution , vol.24 , pp. 1518-1527
    • Haen, K.M.1    Lang, B.F.2    Pomponi, S.A.3    Lavrov, D.V.4
  • 15
    • 12044253190 scopus 로고
    • An empirical test of bootstrapping as a method for assessing confidence in phylogenetic analysis
    • Hillis DM, Bull JJ. 1993. An empirical test of bootstrapping as a method for assessing confidence in phylogenetic analysis. Systematic Biology 42: 182-192.
    • (1993) Systematic Biology , vol.42 , pp. 182-192
    • Hillis, D.M.1    Bull, J.J.2
  • 18
    • 67650079798 scopus 로고    scopus 로고
    • Hexactinellida (Porifera) from the ANDEEP III expedition to the Weddell sea, Antarctica
    • Janussen D, Reiswig HM. 2009. Hexactinellida (Porifera) from the ANDEEP III expedition to the Weddell sea, Antarctica. Zootaxa 2136: 1-20.
    • (2009) Zootaxa , vol.2136 , pp. 1-20
    • Janussen, D.1    Reiswig, H.M.2
  • 19
    • 13444259388 scopus 로고    scopus 로고
    • TREEFINDER: a powerful graphical analysis environment for molecular phylogenetics
    • Jobb G, von Haeseler A, Strimmer K. 2004. TREEFINDER: a powerful graphical analysis environment for molecular phylogenetics. BMC Evolutionary Biology 4: 18.
    • (2004) BMC Evolutionary Biology , vol.4 , pp. 18
    • Jobb, G.1    von Haeseler, A.2    Strimmer, K.3
  • 20
    • 0034943199 scopus 로고    scopus 로고
    • Discovery of a 'living dinosaur': globally unique modern hexactinellid sponge reefs off British Columbia, Canada
    • Krautter M, Conway KW, Barrie JV, Neuweiler M. 2001. Discovery of a 'living dinosaur': globally unique modern hexactinellid sponge reefs off British Columbia, Canada. Facies 44: 265-282.
    • (2001) Facies , vol.44 , pp. 265-282
    • Krautter, M.1    Conway, K.W.2    Barrie, J.V.3    Neuweiler, M.4
  • 23
    • 79956371915 scopus 로고
    • Claviscopulia furcillata n. sp. et autres Hexactinellida (Porifera) des mers indonésiennes
    • Lévi C. 1990. Claviscopulia furcillata n. sp. et autres Hexactinellida (Porifera) des mers indonésiennes. Bulletin Du Muséum National d'Histoire Naturelle (4, A) 8: 277-290.
    • (1990) Bulletin Du Muséum National d'Histoire Naturelle (4, A) , vol.8 , pp. 277-290
    • Lévi, C.1
  • 26
    • 79952957490 scopus 로고    scopus 로고
    • Taxonomy of Farrea (Porifera, Hexactinellida, Hexactinosida) from the southwestern Atlantic, with description of a new species and a discussion on the recognition of subspecies in Porifera
    • Lopes DA, Hajdu E, Reiswig HM. 2011. Taxonomy of Farrea (Porifera, Hexactinellida, Hexactinosida) from the southwestern Atlantic, with description of a new species and a discussion on the recognition of subspecies in Porifera. Canadian Journal of Zoology 89: 169-189.
    • (2011) Canadian Journal of Zoology , vol.89 , pp. 169-189
    • Lopes, D.A.1    Hajdu, E.2    Reiswig, H.M.3
  • 27
    • 0033592605 scopus 로고    scopus 로고
    • Decline in Mesozoic reef-building sponges explained by silicon limitation
    • Maldonado M, Carmona MC, Uriz MJ, Cruzado A. 1999. Decline in Mesozoic reef-building sponges explained by silicon limitation. Nature 401: 785-788.
    • (1999) Nature , vol.401 , pp. 785-788
    • Maldonado, M.1    Carmona, M.C.2    Uriz, M.J.3    Cruzado, A.4
  • 28
    • 0002635796 scopus 로고
    • Die Entwicklung der Hexactinellida seit dem Mesozoikum. Paläobiologie, Phylogenie und Evolutionsökologie
    • Mehl D. 1992. Die Entwicklung der Hexactinellida seit dem Mesozoikum. Paläobiologie, Phylogenie und Evolutionsökologie. Berliner Geowissenschaftliche Abhandlungen Reihe E: Paläobiologie 2: 1-164.
    • (1992) Berliner Geowissenschaftliche Abhandlungen Reihe E: Paläobiologie , vol.2 , pp. 1-164
    • Mehl, D.1
  • 29
    • 42549144579 scopus 로고    scopus 로고
    • On the origins of novelty in development and evolution
    • Moczek AP. 2008. On the origins of novelty in development and evolution. BioEssays 30: 432-447.
    • (2008) BioEssays , vol.30 , pp. 432-447
    • Moczek, A.P.1
  • 30
    • 0342964707 scopus 로고
    • Hexactinellide Poriferen aus pelagischen Kieselkalken (Unterer Lias, Nördliche Kalkalpen)
    • Mostler H. 1990. Hexactinellide Poriferen aus pelagischen Kieselkalken (Unterer Lias, Nördliche Kalkalpen). Geologisch-Paläontologische Mitteilungen der Universität Innsbruck 17: 143-178.
    • (1990) Geologisch-Paläontologische Mitteilungen der Universität Innsbruck , vol.17 , pp. 143-178
    • Mostler, H.1
  • 32
    • 0033510946 scopus 로고    scopus 로고
    • Postpaleozoic history of the siliceous sponges with rigid skeletons
    • Pisera A. 1999. Postpaleozoic history of the siliceous sponges with rigid skeletons. Memoirs of the Queensland Museum 44: 463-472.
    • (1999) Memoirs of the Queensland Museum , vol.44 , pp. 463-472
    • Pisera, A.1
  • 33
    • 0042917313 scopus 로고
    • A monograph of the upper Cretaceous Hexactinellida of Great Britain and Northern Ireland. Part II
    • Reid REH. 1958. A monograph of the upper Cretaceous Hexactinellida of Great Britain and Northern Ireland. Part II. Monographs of the Paleontographical Society 112: xlvii-xlviii.
    • (1958) Monographs of the Paleontographical Society , vol.112 , pp. 47-48
    • Reid, R.E.H.1
  • 34
    • 0000434356 scopus 로고
    • Notes on a classification of the Hexactinosa
    • Reid REH. 1963. Notes on a classification of the Hexactinosa. Journal of Paleontology 37: 218-231.
    • (1963) Journal of Paleontology , vol.37 , pp. 218-231
    • Reid, R.E.H.1
  • 35
    • 0141882050 scopus 로고    scopus 로고
    • Class Hexactinellida Schmidt, 1870
    • In: Hooper JNA, van Soest RWM, eds. New York: Plenum
    • Reiswig HM. 2002a. Class Hexactinellida Schmidt, 1870. In: Hooper JNA, van Soest RWM, eds. Systema Porifera. A guide to the classification of sponges. New York: Plenum, 1201-1202.
    • (2002) Systema Porifera. A guide to the classification of sponges , pp. 1201-1202
    • Reiswig, H.M.1
  • 36
    • 34047178691 scopus 로고    scopus 로고
    • Family Farreidae Gray, 1872
    • In: Hooper JNA, van Soest RWM, eds. New York: Plenum
    • Reiswig HM. 2002b. Family Farreidae Gray, 1872. In: Hooper JNA, van Soest RWM, eds. Systema Porifera. A guide to the classification of sponges. New York: Plenum, 1332-1340.
    • (2002) Systema Porifera. A guide to the classification of sponges , pp. 1332-1340
    • Reiswig, H.M.1
  • 37
    • 82255197088 scopus 로고    scopus 로고
    • Family Cribrospongiidae Roemer, 1864
    • In: Hooper JNA, van Soest RWM, eds. New York: Plenum
    • Reiswig HM. 2002c. Family Cribrospongiidae Roemer, 1864. In: Hooper JNA, van Soest RWM, eds. Systema Porifera. A guide to the classification of sponges. New York: Plenum, 1290-1292.
    • (2002) Systema Porifera. A guide to the classification of sponges , pp. 1290-1292
    • Reiswig, H.M.1
  • 38
    • 9644271116 scopus 로고    scopus 로고
    • Family Uncinateridae fam. nov
    • In: Hooper JNA, van Soest RWM, eds. New York: Plenum
    • Reiswig HM. 2002d. Family Uncinateridae fam. nov. In: Hooper JNA, van Soest RWM, eds. Systema Porifera. A guide to the classification of sponges. New York: Plenum, 1372-1376.
    • (2002) Systema Porifera. A guide to the classification of sponges , pp. 1372-1376
    • Reiswig, H.M.1
  • 39
    • 9644271116 scopus 로고    scopus 로고
    • Family Tretodictyidae Schulze, 1886
    • In: Hooper JNA, van Soest RWM, eds. New York: Plenum
    • Reiswig HM. 2002e. Family Tretodictyidae Schulze, 1886. In: Hooper JNA, van Soest RWM, eds. Systema Porifera. A guide to the classification of sponges. New York: Plenum, 1341-1354.
    • (2002) Systema Porifera. A guide to the classification of sponges , pp. 1341-1354
    • Reiswig, H.M.1
  • 40
    • 33645603793 scopus 로고    scopus 로고
    • Family Aphrocallistidae Gray, 1867
    • In: Hooper JNA, van Soest RWM, eds. New York: Plenum
    • Reiswig HM. 2002f. Family Aphrocallistidae Gray, 1867. In: Hooper JNA, van Soest RWM, eds. Systema Porifera. A guide to the classification of sponges. New York: Plenum, 1282-1286.
    • (2002) Systema Porifera. A guide to the classification of sponges , pp. 1282-1286
    • Reiswig, H.M.1
  • 41
    • 77957276947 scopus 로고    scopus 로고
    • A new species of Tretodictyum (Porifera: Hexactinellida: Tretodictyidae) from off Cocos Island, tropical eastern Pacific Ocean
    • Reiswig HM. 2010. A new species of Tretodictyum (Porifera: Hexactinellida: Tretodictyidae) from off Cocos Island, tropical eastern Pacific Ocean. Proceedings of the Biological Society of Washington 123: 242-250.
    • (2010) Proceedings of the Biological Society of Washington , vol.123 , pp. 242-250
    • Reiswig, H.M.1
  • 42
    • 40949122717 scopus 로고    scopus 로고
    • Two new tretodictyids (Hexactinellida: Hexactinosida: Tretodictyidae) from the coasts of North America
    • Reiswig HM, Dohrmann M, Pomponi SA, Wörheide G. 2008. Two new tretodictyids (Hexactinellida: Hexactinosida: Tretodictyidae) from the coasts of North America. Zootaxa 1721: 53-64.
    • (2008) Zootaxa , vol.1721 , pp. 53-64
    • Reiswig, H.M.1    Dohrmann, M.2    Pomponi, S.A.3    Wörheide, G.4
  • 43
    • 80053274968 scopus 로고    scopus 로고
    • The Marine Fauna of New Zealand: Hexasterophoran Glass Sponges of New Zealand (Porifera: Hexactinellida: Hexasterophora): Orders Hexactinosida, Aulocalycoida and Lychniscosida
    • Reiswig HM, Kelly M. 2011. The Marine Fauna of New Zealand: Hexasterophoran Glass Sponges of New Zealand (Porifera: Hexactinellida: Hexasterophora): Orders Hexactinosida, Aulocalycoida and Lychniscosida. NIWA Biodiversity Memoirs 124: 1-176.
    • (2011) NIWA Biodiversity Memoirs , vol.124 , pp. 1-176
    • Reiswig, H.M.1    Kelly, M.2
  • 45
    • 0035339194 scopus 로고    scopus 로고
    • Upper Devonian sponges from the holy cross mountains, central Poland
    • Rigby JK, Pisera A, Wrzolek T, Racki G. 2001. Upper Devonian sponges from the holy cross mountains, central Poland. Palaeontology 44: 447-488.
    • (2001) Palaeontology , vol.44 , pp. 447-488
    • Rigby, J.K.1    Pisera, A.2    Wrzolek, T.3    Racki, G.4
  • 46
    • 40549134511 scopus 로고    scopus 로고
    • The mitochondrial genome of the hexactinellid sponge Aphrocallistes vastus: evidence for programmed translational frameshifting
    • Rosengarten RD, Sperling EA, Moreno MA, Leys SP, Dellaporta SL. 2008. The mitochondrial genome of the hexactinellid sponge Aphrocallistes vastus: evidence for programmed translational frameshifting. BMC Genomics 9: 33.
    • (2008) BMC Genomics , vol.9 , pp. 33
    • Rosengarten, R.D.1    Sperling, E.A.2    Moreno, M.A.3    Leys, S.P.4    Dellaporta, S.L.5
  • 47
    • 0035156561 scopus 로고    scopus 로고
    • RNA sequence evolution with secondary structure constraints: comparison of substitution rate models using maximum-likelihood methods
    • Savill NJ, Hoyle DC, Higgs PG. 2001. RNA sequence evolution with secondary structure constraints: comparison of substitution rate models using maximum-likelihood methods. Genetics 157: 399-411.
    • (2001) Genetics , vol.157 , pp. 399-411
    • Savill, N.J.1    Hoyle, D.C.2    Higgs, P.G.3
  • 49
    • 0001531754 scopus 로고
    • Die Kieselspongien der oberen Kreide von Nordwestdeutschland. II Teil Triaxonia (Hexactinellida)
    • Suppl. Bd.
    • Schrammen A. 1912. Die Kieselspongien der oberen Kreide von Nordwestdeutschland. II Teil Triaxonia (Hexactinellida). Paläontographica, Suppl. Bd. 5: 177-385.
    • (1912) Paläontographica , vol.5 , pp. 177-385
    • Schrammen, A.1
  • 52
    • 0036623258 scopus 로고    scopus 로고
    • An approximately unbiased test of phylogenetic tree selection
    • Shimodaira H. 2002. An approximately unbiased test of phylogenetic tree selection. Systematic Biology 51: 492-508.
    • (2002) Systematic Biology , vol.51 , pp. 492-508
    • Shimodaira, H.1
  • 53
    • 33750403801 scopus 로고    scopus 로고
    • RAxML-VI-HPC: maximum likelihood-based phylogenetic analyses with thousands of taxa and mixed models
    • Stamatakis A. 2006. RAxML-VI-HPC: maximum likelihood-based phylogenetic analyses with thousands of taxa and mixed models. Bioinformatics 22: 2688-2690.
    • (2006) Bioinformatics , vol.22 , pp. 2688-2690
    • Stamatakis, A.1
  • 54
    • 50249100595 scopus 로고    scopus 로고
    • A rapid bootstrap algorithm for the RAxML web servers
    • Stamatakis A, Hoover P, Rougemont J. 2008. A rapid bootstrap algorithm for the RAxML web servers. Systematic Biology 57: 758-771.
    • (2008) Systematic Biology , vol.57 , pp. 758-771
    • Stamatakis, A.1    Hoover, P.2    Rougemont, J.3
  • 55
    • 0343835671 scopus 로고
    • Hexactinellid sponges from the mountains of the West Pacific
    • In: Shirshov PP, ed. Moscow: Academy of Sciences of the USSR
    • Tabachnick KR. 1988. Hexactinellid sponges from the mountains of the West Pacific. In: Shirshov PP, ed. Structural and functional researches of the marine benthos. Moscow: Academy of Sciences of the USSR, 49-64.
    • (1988) Structural and functional researches of the marine benthos , pp. 49-64
    • Tabachnick, K.R.1
  • 56
    • 33745133878 scopus 로고    scopus 로고
    • Description of a new species and subspecies of Fieldingia, erection of a new family Fieldingidae, and a new order Fieldingida (Porifera; Hexactinellida; Hexasterophora)
    • Tabachnick KR, Janussen D. 2004. Description of a new species and subspecies of Fieldingia, erection of a new family Fieldingidae, and a new order Fieldingida (Porifera; Hexactinellida; Hexasterophora). Bolletino dei Musei di Istituto Biologia della Università di Genova 68: 623-637.
    • (2004) Bolletino dei Musei di Istituto Biologia della Università di Genova , vol.68 , pp. 623-637
    • Tabachnick, K.R.1    Janussen, D.2
  • 57
    • 74149092109 scopus 로고    scopus 로고
    • Two new Hyalonema species (Hyalonematidae: Amphidiscosida) from eastern and south-eastern Brazil, and further Hexactinellida (Porifera) collected from seamounts off south-eastern Brazil by the RV 'Marion Dufresne' MD55 expedition
    • Tabachnick KR, Menshenina LL, Lopes DA, Hajdu E. 2009. Two new Hyalonema species (Hyalonematidae: Amphidiscosida) from eastern and south-eastern Brazil, and further Hexactinellida (Porifera) collected from seamounts off south-eastern Brazil by the RV 'Marion Dufresne' MD55 expedition. Journal of the Marine Biological Association of the United Kingdom 89: 1243-1250.
    • (2009) Journal of the Marine Biological Association of the United Kingdom , vol.89 , pp. 1243-1250
    • Tabachnick, K.R.1    Menshenina, L.L.2    Lopes, D.A.3    Hajdu, E.4
  • 58
    • 79956340301 scopus 로고    scopus 로고
    • Revision of Aspidoscopulia Reiswig, 2002 (Porifera: Hexactinellida: Farreidae) with description of two new species
    • Tabachnick KR, Menshenina LL, Pisera A, Ehrlich H. 2011. Revision of Aspidoscopulia Reiswig, 2002 (Porifera: Hexactinellida: Farreidae) with description of two new species. Zootaxa 2883: 1-22.
    • (2011) Zootaxa , vol.2883 , pp. 1-22
    • Tabachnick, K.R.1    Menshenina, L.L.2    Pisera, A.3    Ehrlich, H.4
  • 61
  • 62
    • 0028064845 scopus 로고
    • Maximum likelihood phylogenetic estimation from DNA sequences with variable rates over sites: approximate methods
    • Yang Z. 1994. Maximum likelihood phylogenetic estimation from DNA sequences with variable rates over sites: approximate methods. Journal of Molecular Evolution 39: 306-314.
    • (1994) Journal of Molecular Evolution , vol.39 , pp. 306-314
    • Yang, Z.1


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