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Volumn 108, Issue 9, 2011, Pages 2022-2035

Polyhydroxybutyrate production from lactate using a mixed microbial culture

Author keywords

Lactate; Microbial diversity; Mixed microbial culture; Polyhydroxybutyrate; Selective pressure

Indexed keywords

LACTATE; MICROBIAL DIVERSITY; MIXED MICROBIAL CULTURE; POLYHYDROXYBUTYRATE; SELECTIVE PRESSURE;

EID: 79960448009     PISSN: 00063592     EISSN: 10970290     Source Type: Journal    
DOI: 10.1002/bit.23148     Document Type: Article
Times cited : (137)

References (48)
  • 1
    • 34447096992 scopus 로고    scopus 로고
    • Strategies for the development of a side stream process for polyhydroxyalkanoate (PHA) production from sugar cane molasses
    • Albuquerque MGE, Eiroa M, Torres C, Nunes BR, Reis MAM. 2007. Strategies for the development of a side stream process for polyhydroxyalkanoate (PHA) production from sugar cane molasses. J Biotechnol 130(4): 411-421.
    • (2007) J Biotechnol , vol.130 , Issue.4 , pp. 411-421
    • Albuquerque, M.G.E.1    Eiroa, M.2    Torres, C.3    Nunes, B.R.4    Reis, M.A.M.5
  • 2
    • 77953000283 scopus 로고    scopus 로고
    • Polyhydroxyalkanoate (PHA) production by a mixed microbial culture using sugar molasses: Effect of the influent substrate concentration on culture selection
    • Albuquerque MGE, Torres CAV, Reis MAM. 2010. Polyhydroxyalkanoate (PHA) production by a mixed microbial culture using sugar molasses: Effect of the influent substrate concentration on culture selection. Water Res 44(11): 3419-3433.
    • (2010) Water Res , vol.44 , Issue.11 , pp. 3419-3433
    • Albuquerque, M.G.E.1    Torres, C.A.V.2    Reis, M.A.M.3
  • 3
    • 0025294528 scopus 로고
    • Combination of 16s ribosomal-RNA-targeted oligonucleotide probes with flow-cytometry for analyzing mixed microbial-populations
    • Amann RI, Binder BJ, Olson RJ, Chisholm SW, Devereux R, Stahl DA. 1990. Combination of 16s ribosomal-RNA-targeted oligonucleotide probes with flow-cytometry for analyzing mixed microbial-populations. Appl Environ Microbiol 56(6): 1919-1925.
    • (1990) Appl Environ Microbiol , vol.56 , Issue.6 , pp. 1919-1925
    • Amann, R.I.1    Binder, B.J.2    Olson, R.J.3    Chisholm, S.W.4    Devereux, R.5    Stahl, D.A.6
  • 4
    • 0025226099 scopus 로고
    • Occurrence, metabolism, metabolic role, and industrial uses of bacterial polyhydroxyalkanoates
    • Anderson AJ, Dawes EA. 1990. Occurrence, metabolism, metabolic role, and industrial uses of bacterial polyhydroxyalkanoates. Microbiol Rev 54(4): 450-472.
    • (1990) Microbiol Rev , vol.54 , Issue.4 , pp. 450-472
    • Anderson, A.J.1    Dawes, E.A.2
  • 5
    • 0036216330 scopus 로고    scopus 로고
    • Effect of different carbon sources on aerobic storage by activated sludge
    • Beccari M, Dionisi D, Giuliani A, Majone M, Ramadori R. 2002. Effect of different carbon sources on aerobic storage by activated sludge. Wat Sci Technol 45(6): 157-168.
    • (2002) Wat Sci Technol , vol.45 , Issue.6 , pp. 157-168
    • Beccari, M.1    Dionisi, D.2    Giuliani, A.3    Majone, M.4    Ramadori, R.5
  • 6
    • 70350492881 scopus 로고    scopus 로고
    • The utilization of glycogen accumulating organisms for mixed culture production of polyhydroxyalkanoates
    • Bengtsson S. 2009. The utilization of glycogen accumulating organisms for mixed culture production of polyhydroxyalkanoates. Biotechnol Bioeng 104(4): 698-708.
    • (2009) Biotechnol Bioeng , vol.104 , Issue.4 , pp. 698-708
    • Bengtsson, S.1
  • 7
    • 44449108001 scopus 로고    scopus 로고
    • Acidogenic fermentation of industrial wastewaters: Effects of chemostat retention time and pH on volatile fatty acids production
    • Bengtsson S, Hallquist J, Werker A, Welander T. 2008. Acidogenic fermentation of industrial wastewaters: Effects of chemostat retention time and pH on volatile fatty acids production. Biochem Eng J 40(3): 492-499.
    • (2008) Biochem Eng J , vol.40 , Issue.3 , pp. 492-499
    • Bengtsson, S.1    Hallquist, J.2    Werker, A.3    Welander, T.4
  • 8
    • 73749084949 scopus 로고    scopus 로고
    • Production of polyhydroxyalkanoates from fermented sugar cane molasses by a mixed culture enriched in glycogen accumulating organisms
    • Bengtsson S, Pisco AR, Reis MAM, Lemos PC. 2010. Production of polyhydroxyalkanoates from fermented sugar cane molasses by a mixed culture enriched in glycogen accumulating organisms. J Biotechnol 145(3): 253-263.
    • (2010) J Biotechnol , vol.145 , Issue.3 , pp. 253-263
    • Bengtsson, S.1    Pisco, A.R.2    Reis, M.A.M.3    Lemos, P.C.4
  • 9
    • 0036182908 scopus 로고    scopus 로고
    • Poly-beta-hydroxybutyrate metabolism in dynamically fed mixed microbial cultures
    • Beun JJ, Dircks K, Van Loosdrecht MCM, Heijnen JJ. 2002. Poly-beta-hydroxybutyrate metabolism in dynamically fed mixed microbial cultures. Water Res 36(5): 1167-1180.
    • (2002) Water Res , vol.36 , Issue.5 , pp. 1167-1180
    • Beun, J.J.1    Dircks, K.2    Van Loosdrecht, M.C.M.3    Heijnen, J.J.4
  • 11
    • 0030720836 scopus 로고    scopus 로고
    • Process analysis and economic evaluation for poly(3-hydroxybutyrate) production by fermentation
    • Choi JI, Lee SY. 1997. Process analysis and economic evaluation for poly(3-hydroxybutyrate) production by fermentation. Bioprocess Eng 17(6): 335-342.
    • (1997) Bioprocess Eng , vol.17 , Issue.6 , pp. 335-342
    • Choi, J.I.1    Lee, S.Y.2
  • 12
    • 0026554422 scopus 로고
    • Parameters affecting solvent production by Clostridium-Pasteurianum
    • Dabrock B, Bahl H, Gottschalk G. 1992. Parameters affecting solvent production by Clostridium-Pasteurianum. Appl Environ Microbiol 58(4): 1233-1239.
    • (1992) Appl Environ Microbiol , vol.58 , Issue.4 , pp. 1233-1239
    • Dabrock, B.1    Bahl, H.2    Gottschalk, G.3
  • 13
    • 0344172842 scopus 로고    scopus 로고
    • The domain-specific probe EUB338 is insufficient for the detection of all bacteria: Development and evaluation of a more comprehensive probe set
    • Daims H, Bruhl A, Amann R, Schleifer KH, Wagner M. 1999. The domain-specific probe EUB338 is insufficient for the detection of all bacteria: Development and evaluation of a more comprehensive probe set. Syst Appl Microbiol 22(3): 434-444.
    • (1999) Syst Appl Microbiol , vol.22 , Issue.3 , pp. 434-444
    • Daims, H.1    Bruhl, A.2    Amann, R.3    Schleifer, K.H.4    Wagner, M.5
  • 15
    • 0035199069 scopus 로고    scopus 로고
    • Sequencing batch reactor: Influence of periodic operation on performance of activated sludges in biological wastewater treatment
    • Dionisi D, Majone M, Tandoi V, Beccari M. 2001. Sequencing batch reactor: Influence of periodic operation on performance of activated sludges in biological wastewater treatment. Ind Eng Chem Res 40(23): 5110-5119.
    • (2001) Ind Eng Chem Res , vol.40 , Issue.23 , pp. 5110-5119
    • Dionisi, D.1    Majone, M.2    Tandoi, V.3    Beccari, M.4
  • 16
    • 1442277073 scopus 로고    scopus 로고
    • Biodegradable polymers from organic acids by using activated sludge enriched by aerobic periodic feeding
    • Dionisi D, Majone M, Papa V, Beccari M. 2004. Biodegradable polymers from organic acids by using activated sludge enriched by aerobic periodic feeding. Biotechnol Bioeng 85(6): 569-579.
    • (2004) Biotechnol Bioeng , vol.85 , Issue.6 , pp. 569-579
    • Dionisi, D.1    Majone, M.2    Papa, V.3    Beccari, M.4
  • 17
    • 27744540208 scopus 로고    scopus 로고
    • Storage of biodegradable polymers by an enriched microbial community in a sequencing batch reactor operated at high organic load rate
    • Dionisi D, Beccari M, Di Gregorio S, Majone M, Papini MP, Vallini G. 2005. Storage of biodegradable polymers by an enriched microbial community in a sequencing batch reactor operated at high organic load rate. J Chem Technol Biotechnol 80(11): 1306-1318.
    • (2005) J Chem Technol Biotechnol , vol.80 , Issue.11 , pp. 1306-1318
    • Dionisi, D.1    Beccari, M.2    Di Gregorio, S.3    Majone, M.4    Papini, M.P.5    Vallini, G.6
  • 18
    • 30944449439 scopus 로고    scopus 로고
    • Effect of the applied organic load rate on biodegradable polymer production by mixed microbial cultures in a sequencing batch reactor
    • Dionisi D, Majone M, Vallini G, Di Gregorio S, Beccari M. 2006. Effect of the applied organic load rate on biodegradable polymer production by mixed microbial cultures in a sequencing batch reactor. Biotechnol Bioeng 93(1): 76-88.
    • (2006) Biotechnol Bioeng , vol.93 , Issue.1 , pp. 76-88
    • Dionisi, D.1    Majone, M.2    Vallini, G.3    Di Gregorio, S.4    Beccari, M.5
  • 19
    • 35348854346 scopus 로고    scopus 로고
    • Effect of the length of the cycle on biodegradable polymer production and microbial community selection in a sequencing batch reactor
    • Dionisi D, Majone M, Vallini G, Di Gregorio S, Beccari M. 2007. Effect of the length of the cycle on biodegradable polymer production and microbial community selection in a sequencing batch reactor. Biotechnol Progr 23: 1064-1073.
    • (2007) Biotechnol Progr , vol.23 , pp. 1064-1073
    • Dionisi, D.1    Majone, M.2    Vallini, G.3    Di Gregorio, S.4    Beccari, M.5
  • 20
    • 68749083911 scopus 로고    scopus 로고
    • Development of a fermentation process for production of calcium-L-lactate
    • Gao MT, Hirata M, Toorisaka E, Hano T. 2009. Development of a fermentation process for production of calcium-L-lactate. Chem Eng Process 48(1): 464-469.
    • (2009) Chem Eng Process , vol.48 , Issue.1 , pp. 464-469
    • Gao, M.T.1    Hirata, M.2    Toorisaka, E.3    Hano, T.4
  • 21
    • 0034135549 scopus 로고    scopus 로고
    • Factors affecting the fermentative lactic acid production from renewable resources
    • Hofvendahl K, Hahn-Hagerdal B. 2000. Factors affecting the fermentative lactic acid production from renewable resources. Enzyme Microb Technol 26(2-4): 87-107.
    • (2000) Enzyme Microb Technol , vol.26 , Issue.2-4 , pp. 87-107
    • Hofvendahl, K.1    Hahn-Hagerdal, B.2
  • 22
    • 78650582002 scopus 로고    scopus 로고
    • Metabolic modeling of mixed substrate uptake for polyhydroxyalkanoate (PHA) production
    • Jiang Y, Hebly M, Kleerebezem R, Muyzer G, van Loosdrecht MCM. 2011. Metabolic modeling of mixed substrate uptake for polyhydroxyalkanoate (PHA) production. Water Res 45(3): 1309-1321.
    • (2011) Water Res , vol.45 , Issue.3 , pp. 1309-1321
    • Jiang, Y.1    Hebly, M.2    Kleerebezem, R.3    Muyzer, G.4    van Loosdrecht, M.C.M.5
  • 23
    • 79955071653 scopus 로고    scopus 로고
    • Effect of temperature and cycle length on microbial competition in PHB-producing sequencing batch reactor
    • DOI: 10.1038/ismej.2010.174.
    • Jiang Y, Marang L, Kleerebezem R, Muyzer G, van Loosdrecht MCM. 2010a. Effect of temperature and cycle length on microbial competition in PHB-producing sequencing batch reactor. ISME J DOI: 10.1038/ismej.2010.174.
    • (2010) ISME J
    • Jiang, Y.1    Marang, L.2    Kleerebezem, R.3    Muyzer, G.4    van Loosdrecht, M.C.M.5
  • 24
    • 79960446132 scopus 로고    scopus 로고
    • Plasticicumulans acidivorans gen. nov., sp. nov., a polyhydroxyalkanoate-accumulating gammaproteobacterium from a sequencing-batch bioreactor
    • DOI: ijs.0.021410-0.
    • Jiang Y, Sorokin D, Kleerebezem R, Muyzer G, van Loosdrecht MCM. 2010b. Plasticicumulans acidivorans gen. nov., sp. nov., a polyhydroxyalkanoate-accumulating gammaproteobacterium from a sequencing-batch bioreactor. Int J Syst Evol Microbiol DOI: ijs.0.021410-0.
    • (2010) Int J Syst Evol Microbiol
    • Jiang, Y.1    Sorokin, D.2    Kleerebezem, R.3    Muyzer, G.4    van Loosdrecht, M.C.M.5
  • 25
    • 65249097716 scopus 로고    scopus 로고
    • Enrichment of a mixed bacterial culture with a high polyhydroxyalkanoate storage capacity
    • Johnson K, Jiang Y, Kleerebezem R, Muyzer G, van Loosdrecht MCM. 2009a. Enrichment of a mixed bacterial culture with a high polyhydroxyalkanoate storage capacity. Biomacromolecules 10(4): 670-676.
    • (2009) Biomacromolecules , vol.10 , Issue.4 , pp. 670-676
    • Johnson, K.1    Jiang, Y.2    Kleerebezem, R.3    Muyzer, G.4    van Loosdrecht, M.C.M.5
  • 26
    • 69249099736 scopus 로고    scopus 로고
    • Model-based data evaluation of polyhydroxybutyrate producing mixed microbial cultures in aerobic sequencing batch and fed-batch reactors
    • Johnson K, Kleerebezem R, van Loosdrecht MCM. 2009b. Model-based data evaluation of polyhydroxybutyrate producing mixed microbial cultures in aerobic sequencing batch and fed-batch reactors. Biotechnol Bioeng 104(1): 50-67.
    • (2009) Biotechnol Bioeng , vol.104 , Issue.1 , pp. 50-67
    • Johnson, K.1    Kleerebezem, R.2    van Loosdrecht, M.C.M.3
  • 27
    • 76949102359 scopus 로고    scopus 로고
    • Short- and long-term temperature effects on aerobic polyhydroxybutyrate producing mixed cultures
    • Johnson K, van Geest J, Kleerebezem R, van Loosdrecht MCM. 2010. Short- and long-term temperature effects on aerobic polyhydroxybutyrate producing mixed cultures. Water Res 44(6): 1689-1700.
    • (2010) Water Res , vol.44 , Issue.6 , pp. 1689-1700
    • Johnson, K.1    van Geest, J.2    Kleerebezem, R.3    van Loosdrecht, M.C.M.4
  • 28
    • 34249952423 scopus 로고    scopus 로고
    • Mixed culture biotechnology for bioenergy production
    • Kleerebezem R, van Loosdrecht MCM. 2007. Mixed culture biotechnology for bioenergy production. Curr Opin Biotechnol 18(3): 207-212.
    • (2007) Curr Opin Biotechnol , vol.18 , Issue.3 , pp. 207-212
    • Kleerebezem, R.1    van Loosdrecht, M.C.M.2
  • 29
    • 0031719199 scopus 로고    scopus 로고
    • Growth kinetics of suspended microbial cells: From single-substrate-controlled growth to mixed-substrate kinetics
    • Kovarova-Kovar K, Egli T. 1998. Growth kinetics of suspended microbial cells: From single-substrate-controlled growth to mixed-substrate kinetics. Microbiol Mol Biol Rev 62(3): 646-666.
    • (1998) Microbiol Mol Biol Rev , vol.62 , Issue.3 , pp. 646-666
    • Kovarova-Kovar, K.1    Egli, T.2
  • 30
    • 0030298096 scopus 로고    scopus 로고
    • Plastic bacteria? Progress and prospects for polyhydroxyalkanoate production in bacteria
    • Lee SY. 1996. Plastic bacteria? Progress and prospects for polyhydroxyalkanoate production in bacteria. Trends Biotechnol 14(11): 431-438.
    • (1996) Trends Biotechnol , vol.14 , Issue.11 , pp. 431-438
    • Lee, S.Y.1
  • 31
    • 33644548315 scopus 로고    scopus 로고
    • Synthesis of polyhydroxyalkanoates from different short-chain fatty acids by mixed cultures submitted to aerobic dynamic feeding
    • Lemos PC, Serafim LS, Reis MAM. 2006. Synthesis of polyhydroxyalkanoates from different short-chain fatty acids by mixed cultures submitted to aerobic dynamic feeding. J Biotechnol 122(2): 226-238.
    • (2006) J Biotechnol , vol.122 , Issue.2 , pp. 226-238
    • Lemos, P.C.1    Serafim, L.S.2    Reis, M.A.M.3
  • 32
    • 38649129394 scopus 로고    scopus 로고
    • Microbial characterisation of polyhydroxyalkanoates storing populations selected under different operating conditions using a cell-sorting RT-PCR approach
    • Lemos PC, Levantesi C, Serafim LS, Rossetti S, Reis MAM, Tandoi V. 2008. Microbial characterisation of polyhydroxyalkanoates storing populations selected under different operating conditions using a cell-sorting RT-PCR approach. Appl Microbiol Biotechnol 78(2): 351-360.
    • (2008) Appl Microbiol Biotechnol , vol.78 , Issue.2 , pp. 351-360
    • Lemos, P.C.1    Levantesi, C.2    Serafim, L.S.3    Rossetti, S.4    Reis, M.A.M.5    Tandoi, V.6
  • 35
    • 0027408903 scopus 로고
    • Thauera selenatis gen. nov, sp. nov, a member of the beta-subclass of proteobacteria with a novel type of anaerobic respiration
    • Macy JM, Rech S, Auling G, Dorsch M, Stackebrandt E, Sly LI. 1993. Thauera selenatis gen. nov, sp. nov, a member of the beta-subclass of proteobacteria with a novel type of anaerobic respiration. Int J Syst Bacteriol 43(1): 135-142.
    • (1993) Int J Syst Bacteriol , vol.43 , Issue.1 , pp. 135-142
    • Macy, J.M.1    Rech, S.2    Auling, G.3    Dorsch, M.4    Stackebrandt, E.5    Sly, L.I.6
  • 36
    • 0027068041 scopus 로고
    • Phylogenetic oligodeoxynucleotide probes for the major subclasses of proteobacteria-Problems and solutions
    • Manz W, Amann R, Ludwig W, Wagner M, Schleifer KH. 1992. Phylogenetic oligodeoxynucleotide probes for the major subclasses of proteobacteria-Problems and solutions. Syst Appl Microbiol 15(4): 593-600.
    • (1992) Syst Appl Microbiol , vol.15 , Issue.4 , pp. 593-600
    • Manz, W.1    Amann, R.2    Ludwig, W.3    Wagner, M.4    Schleifer, K.H.5
  • 39
    • 0142074115 scopus 로고    scopus 로고
    • Denaturing gradient gel electrophoresis in marine microbial ecology
    • Schäfer H, Muyzer G. 2001. Denaturing gradient gel electrophoresis in marine microbial ecology. Method Microbiol 30(30): 425-468.
    • (2001) Method Microbiol , vol.30 , Issue.30 , pp. 425-468
    • Schäfer, H.1    Muyzer, G.2
  • 40
    • 3042710803 scopus 로고    scopus 로고
    • Optimization of polyhydroxybutyrate production by mixed cultures submitted to aerobic dynamic feeding conditions
    • Serafim LS, Lemos PC, Oliveira R, Reis MAM. 2004. Optimization of polyhydroxybutyrate production by mixed cultures submitted to aerobic dynamic feeding conditions. Biotechnol Bioeng 87(2): 145-160.
    • (2004) Biotechnol Bioeng , vol.87 , Issue.2 , pp. 145-160
    • Serafim, L.S.1    Lemos, P.C.2    Oliveira, R.3    Reis, M.A.M.4
  • 41
    • 0031367143 scopus 로고    scopus 로고
    • Metabolic flux analysis for biosynthesis of poly(beta-hydroxybutyric acid) in Alcaligenes eutrophus from various carbon sources
    • Shi HD, Shiraishi M, Shimizu K. 1997. Metabolic flux analysis for biosynthesis of poly(beta-hydroxybutyric acid) in Alcaligenes eutrophus from various carbon sources. J Ferment Bioeng 84(6): 579-587.
    • (1997) J Ferment Bioeng , vol.84 , Issue.6 , pp. 579-587
    • Shi, H.D.1    Shiraishi, M.2    Shimizu, K.3
  • 42
    • 77954282501 scopus 로고    scopus 로고
    • Investigation of bacterial diversity in membrane bioreactor and conventional activated sludge processes from petroleum refineries using phylogenetic and statistical approaches
    • Silva CC, Jesus EC, Torres APR, Sousa MP, Santiago VMJ, Oliveira VM. 2010. Investigation of bacterial diversity in membrane bioreactor and conventional activated sludge processes from petroleum refineries using phylogenetic and statistical approaches. J Microbiol Biotechnol 20(3): 447-459.
    • (2010) J Microbiol Biotechnol , vol.20 , Issue.3 , pp. 447-459
    • Silva, C.C.1    Jesus, E.C.2    Torres, A.P.R.3    Sousa, M.P.4    Santiago, V.M.J.5    Oliveira, V.M.6
  • 43
    • 0023690652 scopus 로고
    • Cloning and expression in Escherichia coli of the Alcaligenes eutrophus H16 poly-beta-hydroxybutyrate biosynthetic-pathway
    • Slater SC, Voige WH, Dennis DE. 1988. Cloning and expression in Escherichia coli of the Alcaligenes eutrophus H16 poly-beta-hydroxybutyrate biosynthetic-pathway. J Bacteriol 170(10): 4431-4436.
    • (1988) J Bacteriol , vol.170 , Issue.10 , pp. 4431-4436
    • Slater, S.C.1    Voige, W.H.2    Dennis, D.E.3
  • 45
    • 34249932212 scopus 로고    scopus 로고
    • Influence of the pH on (open) mixed culture fermentation of glucose: A chemostat study
    • Temudo MF, Kleerebezem R, van Loosdrecht M. 2007. Influence of the pH on (open) mixed culture fermentation of glucose: A chemostat study. Biotechnol Bioeng 98(1): 69-79.
    • (2007) Biotechnol Bioeng , vol.98 , Issue.1 , pp. 69-79
    • Temudo, M.F.1    Kleerebezem, R.2    van Loosdrecht, M.3
  • 46
    • 53249101309 scopus 로고    scopus 로고
    • Diversity of microbial communities in open mixed culture fermentations: Impact of the pH and carbon source
    • Temudo MF, Muyzer G, Kleerebezem R, van Loosdrecht MCM. 2008. Diversity of microbial communities in open mixed culture fermentations: Impact of the pH and carbon source. Appl Microbiol Biotechnol 80(6): 1121-1130.
    • (2008) Appl Microbiol Biotechnol , vol.80 , Issue.6 , pp. 1121-1130
    • Temudo, M.F.1    Muyzer, G.2    Kleerebezem, R.3    van Loosdrecht, M.C.M.4
  • 47
    • 77950296763 scopus 로고    scopus 로고
    • Effect of pH on the production of bacterial polyhydroxyalkanoates by mixed cultures enriched under periodic feeding
    • Villano M, Beccari M, Dionisi D, Lampis S, Miccheli A, Vallini G, Majone M. 2010. Effect of pH on the production of bacterial polyhydroxyalkanoates by mixed cultures enriched under periodic feeding. Process Biochem 45(5): 714-723.
    • (2010) Process Biochem , vol.45 , Issue.5 , pp. 714-723
    • Villano, M.1    Beccari, M.2    Dionisi, D.3    Lampis, S.4    Miccheli, A.5    Vallini, G.6    Majone, M.7


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