Bioturbation in Burgess Shale-type Lagerstätten - Case study of trace fossil-body fossil association from the Kaili Biota (Cambrian Series 3), Guizhou, China

Palaeogeography, Palaeoclimatology, Palaeoecology

Volumn 292, Issue 1-2, 2010, Pages 245-256

  Lin, Jih Pai ^a,b   Zhao, Yuan Long ^c   Rahman, Imran A ^d   Xiao, Shuhai ^e   Wang, Yue ^c  

^a NANJING INSTITUTE OF GEOLOGY AND PALAEONTOLOGY   (China)

^b YALE UNIVERSITY   (United States)

^c GUIZHOU UNIVERSITY   (China)

^d UNIVERSITY OF MANCHESTER   (United Kingdom)

^e VIRGINIA POLYTECHNIC INSTITUTE AND STATE UNIVERSITY   (United States)

Author keywords

Cambrian; Fecal pellets; Ichnofossils; Konservat Lagerst tten; Sediment reworking

Indexed keywords

BIOTURBATION; DEEP-SEA SEDIMENT; ECHINODERM; ICHNOLOGY; SEAFLOOR; SHALE; TOMOGRAPHY; TRACE FOSSIL; TRILOBITE;

CHINA; GUIZHOU;

ANIMALIA; ARTHROPODA; CREEDIIDAE; CRUZIANA; ECHINODERMATA; GORDIA; PLANOLITES; RUSOPHYCUS; TRICHOPHYCUS; TRILOBITOMORPHA;

EID: 78049297098     PISSN: 00310182     EISSN: None     Source Type: Journal    
DOI: 10.1016/j.palaeo.2010.03.048     Document Type: Article

References (103)

1. Allison, P.A., Brett, C.E., 1995. In situ benthos and paleo-oxygenation in the Middle Cambrian Burgess Shale, British Columbia, Canada. Geology 23, 1079-1082.
2. Allison, P.A., Briggs, D.E.G., 1993a. Burgess Shale biotas: burrowed away? Lethaia 26, 184-185.
3. Allison, P.A., Briggs, D.E.G., 1993b. Exceptional fossil record: distribution of soft-tissue preservation through the Phanerozoic. Geology 21, 527-530.
4. Arakawa, K.Y., 1963. Studies on the molluscan faeces (I). Publications of the Seto Marine Biological Laboratory 11, 185-208.
5. Arakawa, K.Y., 1965. Studies on the molluscan faeces (II). Publications of the Seto Marine Biological Laboratory 13, 1-21.
6. Arakawa, K.Y., 1968. Studies on the molluscan faeces (III). Publications of the Seto Marine Biological Laboratory 16, 127-139.
7. Aronson, R.B., 1992. Decline of the Burgess Shale fauna: ecologic or taphonomic restrictions? Lethaia 25, 225-229.
8. Aronson, R.B., 1993. Burgess Shale-type biotas were not just burrowed away: reply. Lethaia 26, 185.
9. Ausich, W.I., 2001. Echinoderm taphonomy. In: Jangoux, M., Lawrence, J.M. (Eds.), Echinoderm Studies.: A, 6. A. Balkema, Rotterdam, pp. 171-227.
10. Babcock, L.E., Zhang, W.-T., Leslie, S.A., 2001. The Chengjiang Biota: record of the Early Cambrian diversication of life and clues to exceptional preservation of fossils. GSA Today 11 (2), 4-9.
11. Bambach, R.K., Bush, A.M., Erwin, D.H., 2007. Autecology and the lling of ecospace: key metazoan radiations. Palaeontology 50, 1-22.
12. Bartels, C., Briggs, D.E.G., Brassel, G., 1998. The Fossils of the Hunsrück Slate: Marine Life in the Devonian. Cambridge University Press, Cambridge. 324 pp.
13. Bergström, J., Hou, X.-G., Hålenius, U., 2007. Gut contents and feeding in the Cambrian arthropod Naraoia. GFF 129, 71-76.
14. Borkow, P.S., Babcock, L.E., 2003. Turing pyrite concretions outside-in: role of biolms in pyritization of fossils. The Sedimentary Record 1 (3), 4-7.
15. Bottjer, D.J., Hagadorn, J.W., Dornbos, S.Q., 2000. The Cambrian substrate revolution. GSA Today 10 (9), 1-7.
16. Briggs, D.E.G., Erwin, D.H., Collier, F.J., 1994. The Fossils of the Burgess Shale. Smithsonian Institution Press, Washington, D.C. 238 pp.
17. Bromley, R.G., Beuck, L., Taddei Ruggiero, E., 2008. Endolithic sponge versus terebratulid brachiopod, Pleistocene, Italy: accidental symbiosis, bioclaustration and deformity. In: Wisshak, M., Tapanila, L. (Eds.), Current Developments in Bioerosion. Erlangen Earth Conference Series. Springer-Verlag, Berlin Heidelberg, pp. 361-368.
18. Bruthansová, J., Kraft, P., 2003. Pellets independent of or associated with Bohemian Ordovician body fossils. Acta Palaeontol. Pol. 48, 437-445.
19. Buatois, L.A., Mángano, M.G., Maples, C.G., Lanier, W.P., 1998. Ichnology of an Upper Carboniferous uvio-estuarine paleovalley: the Tonganoxie Sandstone, Buildex Quarry, eastern Kansas. J. Paleontol. 72, 152-180.
20. Buttereld, N.J., 1995. Secular distribution of Burgess-Shale-type preservation. Lethaia 28, 1-13.
21. Buttereld, N.J., 2003. Exceptional fossil preservation and the Cambrian explosion. Integrative and Comparative Biology 43, 166-177.
22. Caron, J.-B., Jackson, D.A., 2006. Taphonomy of the Greater Phyllopod Bed community, Burgess Shale. Palaios 21, 451-465.
23. Chamberlain, C.K., 1977. Ordovician and Devonian trace fossils from Nevada. Nevada Bureau of Mines and Geology, Bulletin 90, 1-24.
24. Conway Morris, S., 1985. Cambrian Lagerstätten: their distribution and signicance. Philos. Trans. R. Soc. B-Biol. Sci. 311, 49-65.
25. Conway Morris, S., 1990. Burgess Shale. In: Briggs, D.E.G., Crowther, P.R. (Eds.), Palaeobiology: a Synthesis. Blackwell Scientic, Oxford, pp. 270-274.
26. Conway Morris, S., 1998. The Crucible of Creation: the Burgess Shale and the Rise of Animals. Oxford University Press, Oxford. 242 pp.
27. Donoghue, P.C.J., Dong, X.-P., 2005. Embryos and ancestors. In: Briggs, D.E.G. (Ed.), Evolving Form and Function: Fossils and Development: Proceedings of a Symposium Honoring Adolf Seilacher for his Contributions to Paleontology, in Celebration of his 80th Birthday. Peabody Museum of Natural History, New Haven, pp. 81-99.
28. Donoghue, P.C.J., Kouchinsky, A., Waloszek, D., Bengtson, S., Dong, X.-P., Val'kov, A.K., Cunningham, J.A., Repetski, J.E., 2006. Fossilized embryos are widespread but the record is temporally and taxonomically biased. Evolution & Development 8, 232-238.
29. Droser, M.L., Bottjer, D.J., 1986. A semiquantitative eld classication of ichnofabric = Journal of Sedimentary Research J. Sed. Petrol. 56, 558-559.
30. Droser, M.L., Bottjer, D.J., 1993. Trends and patterns of Phanerozoic ichnofabrics. Annual Review of Earth and Planetary Science 21, 205-225.
31. Fillion, D., Pickerill, R.K., 1984. On Arthraria antiquata Billings, 1872 and its relationship to Diplocraterion Torell, 1870 and Bifungites Desio, 1940. J. Paleontol. 58, 683-696.
32. Fu, S.-P., Werner, F., Brossmann, J., 1994. Computed tomography: application in studying biogenic structures in sediment cores. Palaios 9, 116-119.
33. Gabbott, S.E., Zalasiewicz, J.A., Collins, D., 2008. Sedimentation of the Phyllopod Bed within the Cambrian Burgess Shale Formation of British Columbia. J. Geol. Soc. London 165, 307-318.
34. Gaines, R.R., Droser, M.L., 2005. New approaches to understanding the mechanics of Burgess Shale-type deposits: from the micron scale to the global picture. The Sedimentary Record 3 (2), 4-8.
35. Gaines, R.R., Kennedy, M.J., Droser, M.L., 2005. A new hypothesis for organic preservation of Burgess Shale taxa in the middle Cambrian Wheeler Formation, House Range, Utah. Palaeogeogr. Palaeoclimatol. Palaeoecol. 220, 193-205.
36. Gaines, R.R., Briggs, D.E.G., Zhao, Y.-L., 2008. Cambrian Burgess Shale-type deposits share a common mode of fossilization. Geology 36, 755-758.
37. Guensburg, T.E., Sprinkle, J., 1992. Rise of echinoderms in the Paleozoic evolutionary fauna: signicance of paleoenvironmental controls. Geology 20, 407-410.
38. Häntzschel, W., 1975. Treatise on invertebrate paleontology, Part W, Miscellanea Supplement 1, trace fossils and problematica, Second Edition (Revised and Enlarged). Treatise on invertebrate paleontology. The Geological Society of America, Inc. and The University of Kansas, Boulder. Colorado and Lawrence, Kansas. 269 pp.
39. Hatai, K., Kotaka, T., Noda, H., 1970. Supplementary note on the faecal pellets from the Early Miyagian Kogota Formation, Kogota-Machi, Miyagi Prefecture, Northeast Honshu, Japan. Saito Ho-on Kai Museum of Natural History Research Bulletin 39, 7-11.
40. Hou, X.-G., Aldridge, R.J., Bergström, J., Siveter, D.J., Siveter, D.J., Feng, X.-H., 2004. The Cambrian Fossils of Chengjiang, China: the Flowering of Early Animal Life. Blackwell Publishing, Oxford. 233 pp.
41. Hu, S.-X., 2005. Taphonomy and Palaeoecology of the Early Cambrian Chengjiang Biota from Eastern Yunnan, China. Berliner Paläobiologische Abhandlungen 7, 1-197.
42. Johnston, P.A., Johnston, K.J., Collom, C.J., Powell, W.G., Pollock, R.J., 2009. Palaeontology and depositional environments of ancient brine seeps in the Middle Cambrian Burgess Shale at the Monarch, British Columbia, Canada. Palaeogeogr. Palaeoclimatol. Palaeoecol. 277, 86-105.
43. Lin, J.-P., 2006. Taphonomy of naraoiids (Arthropoda) from the Middle Cambrian Kaili Biota, Guizhou Province, South China. Palaios 21, 15-25.
44. Lin, J.-P., 2007. Preservation of the gastrointestinal system in Olenoides (Trilobita) from the Kaili Biota (Cambrian) of Guizhou, China. Memoirs of the Association of Australasian Palaeontologists 33, 179-189.
45. Lin, J.-P., 2008. From a Fossil Assemblage to a Paleoecological Community: Time, Organism and Environment Based on the Kaili Lagerstätte (Cambrian). South China and Coeval Deposits of Exceptional Preservation. VDM Verlag Dr, Müller, Saarbrücken. 412 pp.
46. Lin, J.-P., 2009. Review of the depositional environment of the Kaili Formation (Cambrian Series 2-3 boundary interval: China). Memoirs of the Association of Australasian Palaeontologists 37, 131-149.
47. Lin, J.-P., Yuan, J.-L., Wang, Y., Zhao, Y.-L., 2005. Introduction to the type section of the Kaili Formation, Danzhai, Guizhou Province, China. In: Peng, S.-C., Babcock, L.E., Zhu, M.-Y. (Eds.), Cambrian System of China and Korea - Guide to Field Excursions. University of Science and Technology of China Press, Hefei, China, pp. 55-61.
48. Lin, J.-P., Gon III, S.M., Gehling, J.G., Babcock, L.E., Zhao, Y.-L., Zhang, X.-L., Hu, S.-X., Yuan, J.-L., Yu, M.-Y., Peng, J., 2006. A Parvanocrina-like arthropod from Cambrian strata of South China. Hist. Biol. 18, 22-45.
49. Lin, J.-P., Ausich, W.I., Zhao, Y.-L., 2008a. Settling strategy of stalked echinoderms from the Kaili Biota (middle Cambrian), Guizhou Province, South China. Palaeogeogr. Palaeoclimatol. Palaeoecol. 258, 213-221.
50. Lin, J.-P., Ausich, W.I., Zhao, Y.-L., Peng, J., 2008b. Taphonomy, palaeoecological implications, and colouration of Cambrian gogiid echinoderms from Guizhou Province, South China. Geol. Mag. 145, 17-36.
51. Luo, H.-L., Yong, L., Hu, S.-X., Fu, X.-P., Hou, S.-G., Liu, X.-Y., Chen, L.-Z., Li, F.-J., Pang, J.-Y., Liu, Q., 2008. Early Cambrian Malong Fauna and Guanshan Fauna from Eastern Yunnan. China, Yunnan Science and Technology Press, Kunming. 134 pp.
52. Maas, A., Braun, A., Müller, K.J., Waloszek, D., 2006. The 'Orsten'-more than a Cambrian Konservat-Lagerstätte yielding exceptional preservation. Palaeoworld 15, 266-282.
53. MacKey, S.D., Bridge, J.S., 1995. Three-dimensional model of alluvial stratigraphy: theory and application. J. Sed. Res. B65, 7-31.
54. Mao, J.-R., Zhao, Y.-L., 1994. Scenella of Kaili Fauna from Taijiang, Guizhou. Acta Palaeontologica Sinica 33, 325-328.
55. Martin, K.D., 2004. A re-evaluation of the relationship between trace fossils and dysoxia. In: McIlroy, D. (Ed.), The Application of Ichnology to Palaeoenvironmental and Stratigraphic Analysis. The Geological Society of London, London, pp. 141-156.
56. Orr, P.J., Benton, M.J., Briggs, D.E.G., 2003. Post-Cambrian closure of the deep-water slope-basin taphonomic window. Geology 31, 769-772.
57. Parsley, R.L., Zhao, Y.-L., 2006. Long stalked eocrinoids in the basal Middle Cambrian Kaili Biota, Taijiang County, Guizhou Province, China. J. Paleontol. 80, 1058-1071.
58. Pemberton, S.G., McEachern, J.A., Saunders, T., 2004. Stratigraphic applications of substrate-specic ichnofacies: delineating discontinuities in the rock record. In: McIlroy, D. (Ed.), The Application of Ichnology to Palaeoenvironmental and Stratigraphic Analysis. The Geological Society of London, London, pp. 29-62.
59. Potter, P.E., Maynard, J.B., Depetris, P.J., 2004. Mud and Mudstones: Introduction and Overview. Springer, Berlin Heidelberg. 297 pp.
60. Powell, W.G., Johnston, P.A., Collom, C.J., 2003. Geochemical evidence for oxygenated bottom waters during deposition of fossiliferous strata of the Burgess Shale Formation. Palaeogeogr. Palaeoclimatol. Palaeoecol. 201, 249-268.
61. Rahman, I.A., Jefferies, R.P.S., Südkamp, W.H., Smith, R.D.A., 2009. Ichnological insights into mitrate palaeobiology. Palaeontology 52, 127-138.
62. Rhoads, D.C., 1974. Organism-sediment relations on the muddy sea oor. Oceanogr. Mar. Biol. Annu. Rev. 12, 263-300.
63. Rhoads, D.C., Young, D.K., 1970. The inuence of deposit-feeding organisms on sediment stability and community trophic structure. J. Mar. Res. 28, 150-178.
64. Richter, R., 1936. Marken und Spuren im Hunsrück-Schiefer. II. Schichtung und Grund-Leben. Senckenbergiana 18, 215-244.
65. Robison, R.A., 1991. Middle Cambrian biotic diversity: examples from four Utah Lagerstätten. In: Simonetta, A.M., Conway Morris, S. (Eds.), The Early Evolution of Metazoa and the Signicance of Problematic Taxa. Cambridge University Press, Cambridge, pp. 77-98.
66. Schieber, J., 2001. A role for organic petrology in integrated studies of mudrocks: examples from Devonian black shales of the eastern US. Int. J. Coal Geol. 47, 171-187.
67. Seilacher, A., 1970. Fossil-Lagerstätten, Nr. 1: Begriff und Bedeutung der Fossil-Lagerstätten. Neues Jahrbuch für Geologie und Paläontologie. Monatshefte 1, 34-39.
68. Seilacher, A., Reif, W.E., Westphal, F., 1985. Sedimentological, ecological and temporal patterns of fossil Lagerstätten. Philos. Trans. R. Soc. B-Biol. Sci. 311, 5-23.
69. Seilacher, A., Buatois, L.A., Mángano, M.G., 2005. Trace fossils in the Ediacaran-Cambrian transition: behavioral diversication, ecological turnover and environmental shift. Palaeogeogr. Palaeoclimatol. Palaeoecol. 227, 323-356.
70. Sprinkle, J., 1973. Morphology and Evolution of Blastozoan Echinoderms. The Museum of Comparative Zoology, Harvard University, Cambridge. 284 pp.
71. Sundberg, F.A., Yuan, J.-L., McCollum, L.B., Zhao, Y.-L., 1999. Correlation of the Lower-Middle Cambrian Boundary of South China and western United States of America Supp Acta Palaeontologica Sinica 38, 102-107.
72. Sutcliffe, O.E., Südkamp, W.H., Jefferies, R.P.S., 2000. Ichnological evidence on the behaviour of mitrates: two trails associated with the Devonian mitrate Rhenocystis. Lethaia 33, 1-12.
73. Sutton, M.D., 2008. Tomographic techniques for the study of exceptionally preserved fossils. Proc. R. Soc. Lond. Ser. B 275, 1587-1593.
74. Sutton, M.D., Briggs, D.E.G., Siveter, D.J., Siveter, D.J., 2001. Methodologies for the visualization and reconstruction of three-dimensional fossils from the Silurian Herefordshire Lagerstätte. Palaeontologia Electronica, Vol. 4. [Online]. Available from World Wide Web:.
75. Sutton, M.D., Briggs, D.E.G., Siveter, D.J., Siveter, D.J., Orr, P.J., 2002. The arthropod Offacolus kingi (Chelicerata) from the Silurian of Herefordshire, England: computer based morphological reconstructions and phylogenetic afnities. Proc. R. Soc. Lond. Ser. B 269, 1195-1203.
76. Tai, T.-S., Deng, S. (Eds.), 2005. Fossils: the Key to the Past, Morphology of Early Life - 500 Million Years Ago. Translated by Peng Jin, Ron Parsley & (Alex) Jih-Pai Lin. Permanent URL: http://hdl.handle.net/1811/24227.
77. Taylor, A.M., Goldring, R., 1993. Description and analysis of bioturbation and ichnofabric. J. Geol. Soc. London 150, 141-148.
78. Taylor, A., Goldring, R., Gowland, S., 2003. Analysis and application of ichnofabrics. Earth-Sci. Rev. 60, 227-259.
79. Uchman, A., 1995. Taxonomy and palaeoecology of ysch trace fossils: the Marnosoarenacea Formation and associated facies (Miocene, Northern Apennines, Italy). Beringeria 15, 3-115.
80. Uchman, A., Pika-Biolzi, M., Hochuli, P.A., 2004. Oligocene trace fossils from temporary uvial plain ponds: an example from the Freshwater Molasse of Switzerland. Eclogae Geol. Helv. 97, 133-148.
81. Vannier, J., Chen, J.-Y., 2002. Digestive system and feeding mode in Cambrian naraoiid arthropods. Lethaia 35, 107-120.
82. Vizcaïno, D., Álvaro, J.J., Monceret, E., 2004. Trilobites and ichnofossils from a new fossil Lagerstätte in the Lower Cambrian Pardailhan Formation, southern Montagne Noire, France. Geobios 37, 277-286.
83. Vorhies, J.S., Gaines, R.R., 2009. Microbial dissolution of clay minerals as a source of iron and silica in marine sediments. Nature Geoscience 2, 221-225.
84. Wang, Y., 2006. A new ichnogenus Kailidiscus produced by attachment from the Middle Cambrian Kaili Formation in Taijiang, Guizhou Province, China. Acta Geol. Sinica 80, 27-37.
85. Wang, Y., Wang, P.-L., 2006. Ichnofossil Treptichnus from the Kaili Formation at Taijiang County, Guizhou Province. Geological Review 52, 1-10.
86. Wang, Y., Zhao, Y.-L., Lin, J.-P., Wang, P.-L., 2004. Relationship between trace fossil Gordia and medusiform fossils Pararotadiscus from the Kaili Biota, Taijiang, Guizhou, and its signicance. Geological Review 50, 113-119.
87. Wang, Y., Yu, Y.-Y., Peng, J., Wang, P.-L., 2006a. Discussion on the sequence stratigraphy and sea-level changes of the Kaili Formation at Balang, Taijiang, Guizhou. Journal of Stratigraphy 30, 34-43.
88. Wang, Y., Zhang, H.-J., Yang, X.-G., Li, S.-L., Wang, Q.-S., 2006b. Trace fossils from the Middle Cambrian Jialao Formation at Nangao Town, Danzhai County, Guizhou Province, China. Geological Bulletin of China 25, 475-481.
89. Wang, Y., Zhou, Z.-C., Zhao, Y.-L., Yu, M.-Y., 2006c. Relationship between ichnocoenosis and sedimentary environment in the Lower-Middle Cambrian Kaili Formation of Taijiang, Guizhou, China. Acta Palaeontologica Sinica 45, 243-251.
90. Wang, Y., Lin, J.-P., Zhao, Y.-L., Orr, P.J., 2009. Palaeoecology of the trace fossil Gordia and its interaction with nonmineralizing taxa from the early Middle Cambrian Kaili Biota, Guizhou province, South China. Palaeogeogr. Palaeoclimatol. Palaeoecol. 277, 141-148.
91. Yang, S.-P., 1994. Trace fossils from Early - Middle Cambrian Kaili Formation in Taijiang, Guizhou. Acta Palaeontologica Sinica 33, 350-358.
92. Yang, R.-D., Zhao, Y.-L., 1999. Discovery on trace fossils from the Early - Middle Cambrian Kaili Formation of Taijiang, Guizhou. Acta Palaeontologica Sinica 38, 58-65 Supp.
93. Zhang, X.-G., Pratt, B.R., 2008. Microborings in Early Cambrian phosphatic and phosphatized fossils. Palaeogeogr. Palaeoclimatol. Palaeoecol. 267, 185-195.
94. Zhang, Z.-H., Sheng, J.-W., Gong, X.-Y., Zhao, Y.-L., Mao, J.-R., Yan, C.-H., 1996. A preliminary discussion on preservation condition of Kaili Fauna, Middle Cambrian, Taijiang, Guizhou. Acta Palaeontologica Sinica 35, 607-622.
95. Zhang, X.-G., Hou, X.-G., Bergström, J., 2006. Early Cambrian priapulid worms buried with their lined burrows. Geol. Mag. 143, 743-748.
96. Zhang, X.-G., Bergström, J., Bromley, R.G., Hou, X.-G., 2007. Diminutive trace fossils in the Chengjiang Lagerstätte. Terra Nova 19, 407-412.
97. Zhao, Y.-L., Huang, Y.-Z., Gong, Z.-Y., 1994. Echinoderm fossils of Kaili Fauna from Taijing, Guizhou. Acta Palaeontologica Sinica 33, 303-324.
98. Zhao, Y.-L., Zhu, M.-Y., Babcock, L.E., Yuan, J.-L., Parsley, R.L., Peng, J., Yang, X.-L., Wang, Y., 2005. Kaili Biota: a taphonomic window on diversication of metazoans from the basal Middle Cambrian: Guizhou, China. Acta Geol. Sinica 79, 751-765.
99. Zhao, Y.-L., Yuan, J.-L., Peng, S.-C., Babcock, L.E., Peng, J., Lin, J.-P., Guo, Q.-J., Wang, Y.-X., 2007. New data on the Wuliu-Zengjiayan section (Balang, South China), GSSP candidate for the base of Cambrian Series 3. Memoirs of the Association of Australasian Palaeontologists 33, 57-65.
100. Zhu, M.-Y., Erdtmann, B.-D., Zhao, Y.-L., 1999. Taphonomy and paleoecology of the early Middle Cambrian Kaili Lagerstätte in Guizhou, China. Acta Palaeontologica Sinica 38, 28-57 Supp.
101. Zhu, M.-Y., Zhao, Y.-L., Chen, J.-Y., 2002. Revision of the Cambrian discoidal animals Stellostomites eumorphus and Pararotadiscus guizhouensis from South China. Geobios 35, 165-185.
102. Zhu, M.-Y., Vannier, J., Van Iten, H., Zhao, Y.-L., 2004. Direct evidence for predation on trilobites in the Cambrian. Proc. R. Soc. Lond. Ser. B 271, S277-S280. Supp.
103. Zhu, M.-Y., Babcock, L.E., Peng, S.-C., 2006. Advances in Cambrian stratigraphy and paleontology: integrating correlation techniques, paleobiology, taphonomy and paleoenvironmental reconstruction. Palaeoworld 15, 217-222.