An integrative approach to distinguishing the late permian dicynodont species Oudenodon bainii and Tropidostoma microtrema (Therapsida: Anomodontia)

Palaeontology

Volumn 50, Issue 5, 2007, Pages 1175-1209

  Botha, J ^a   Angielczyk, K D ^b  

^a NATIONAL MUSEUM   (South Africa)

^b UNIVERSITY OF BRISTOL   (United Kingdom)

Author keywords

Biostratigraphy; Bonehistology; KarooBasin; Morphometrics; Oudenodon bainii; Permian; Tropidostoma microtrema

Indexed keywords

BIOSTRATIGRAPHY; HISTOLOGY; INTEGRATED APPROACH; MORPHOLOGY; MORPHOMETRY; PERMIAN; REPTILE; SEXUAL DIMORPHISM; TOOTH;

AFRICA; KAROO; SOUTH AFRICA; SOUTHERN AFRICA; SUB-SAHARAN AFRICA;

ANOMODONTIA; CISTECEPHALUS; OUDENODON; THERAPSIDA;

EID: 34548766028     PISSN: 00310239     EISSN: 14754983     Source Type: Journal    
DOI: 10.1111/j.1475-4983.2007.00697.x     Document Type: Article

References (126)

1. ANDERSON, J. M. and CRUICKSHANK, A. R. I. 1978. The biostratigraphy of the Permian and the Triassic. Part 5. A review of the classification and distribution of Permo-Triassic tetrapods. Palaeontologia Africana, 21, 15-44.
2. ANGIELCZYK, K. D. 2001. Preliminary phylogenetic analysis and stratigraphic congruence of the dicynodont anomodonts (Synapsida: Therapsida). Palaeontologia Africana, 37, 53-79.
3. ANGIELCZYK, K. D. 2002. Redescription, phylogenetic position, and stratigraphic significance of the dicynodont genus Odontocyclops (Synapsida: Anomodontia). Journal of Paleontology, 76, 1047-1059.
4. ANGIELCZYK, K. D. and KURKIN, A. A. 2003a. Phylogenetic analysis of Russian Iermian dicynodonts (Therapsida: Anomodontia): implications for Permian biostratigraphy and Pangaean biogeography. Zoological Journal of the Linnean Society, 139, 157-212.
5. ANGIELCZYK, K. D. and KURKIN, A. A. 2003b. Has the utility of Dicynodon for Late Permian terrestrial biostratigraphy been overstated? Geology, 31, 363-366.
6. BARRY, T. H. 1958. On the significance of tuskless specimens of Dicynodon grimbeeki Broom. Palaeontologia Africana, 5, 57-66.
7. BERGER, J. and GOMPPER, M. E. 1999. Sex ratios in extant ungulates: products of contemporary predation or past life histories? Journal of Mammology, 80, 1084-1113.
8. BOND, G. 1973. The palaeontology of Rhodesia. Bulletin of the Geological Survey of Rhodesia, 70, 1-121.
9. BOOKSTEIN, F. L. 1989. Principal warps: Thin-plate splines and the decompositions of deformations. IEEE Transactions on Image Analysis and Machine Intelligence, 2, 567-585.
10. BOOKSTEIN, F. L. 1991. Morphometric tools for landmark data. Cambridge University Press, Cambridge, 455 pp.
11. BOOKSTEIN, F. L. 1996. Biometrics, biomathematics, and the morphometric synthesis. Bulletin of Mathematical Biology, 58, 313-365.
12. BOOKSTEIN, F. L. 1998. A hundred years of morphometrics. Acta Zoologica Academiae Scientiarum Hungarica, 44, 7-59.
13. BOONSTRA, L. D. 1938. A report on some Karroo vertebrates from the Luangwa Valley, Northern Rhodesia. Quarterly Journal of the Geological Society of London, 94, 371-384.
14. BOTHA, J. 2003. Biological aspects of the Permian dicynodont Oudenodon (Therapsida: Dicynodontia) deduced from bone histology and cross-sectional geometry. Palaeontologia Africana, 39, 37-44.
15. BOTHA, J. and CHINSAMY, A. 2000. Growth patterns from the bone histology of the cynodonts Diademodon and Cynognathus. Journal of Vertebrate Paleontology, 20, 705-711.
16. BOTHA, J. and CHINSAMY, A. 2004. Growth and lifestyle adaptations of the Triassic non-mammalian cynodont Trirachodon. Acta Palaeontologica Polonica, 49, 619-627.
17. BOTHA, J. and CHINSAMY, A. 2005. Growth patterns of Thrinaxodon, a non-mammalian cynodont from the Lower Triassic of South Africa. Palaeontology, 48, 385-394.
18. BROILI, F. and SCHRÖDER, J. 1935. Beobachtungen an Wirbeltieren der Karrooformation XII. Über einige primitive Anomodontier-Schädel aus den unteren Beaufort-Schichten. Sitzungsberichte der Bayerischen Akademie der Wissenschaften, 1935, 223-278.
19. BROILI, F. 1936. Beobachtungen an Wirbeltieren der Karrooformation XVII. Ein neuer Anomodontier aus der Cistecephalus Zone. Sitzungsberichte der Bayerischen Akademie der Wissenschaften, 1936, 45-60.
20. BROOM, R. 1901. On the structure and affinities of Udenodon. Proceedings of the Zoological Society of London, 1901, 162-190.
21. BROOM, R. 1912. On some points in the structure of the dicynodont skull. Annals of the South African Museum, 7, 337-351.
22. BROOM, R. 1915. On the anomodont genera, Pristerodon and Tropidostoma. Proceedings of the Zoological Society of London, 1915, 355-361.
23. BROOM, R. 1932. The mammal-like reptiles of South Africa and the origin of mammals. H. F. & G. Witherby, London, 376 pp.
24. BROOM, R. 1935. On some new genera and species of Karroo fossil reptiles. Annals of the Transvaal Museum, 18, 55-72.
25. BROOM, R. 1950. Three new species of anomodonts from the Rubidge Collection. Annals of the Transvaal Museum, 21, 246-250.
26. BROOM, R. and HAUGHTON, S. H. 1917. Some new species of Anomodontia (Reptilia). Annals of the South African Museum, 12, 119-125.
27. BULL, J. J. and CHARNOV, E. L. 1988. How fundamental are Fisherian sex ratios? Oxford Surveys in Evolutionary Biology, 5, 96-135.
28. CASTANET, J. and BAEZ, M. 1991. Adaptation and evolution in Gallotia lizards from the Canary Islands: Age, growth, maturity and longevity. Amphibia-Reptilia, 12, 81-102.
29. CHINSAMY, A. 1990. Physiological implications of the bone histology of Syntarsus rhodesiensis (Saurischia: Theropoda). Palaeontologia Africana, 27, 77-82.
30. CHINSAMY, A. 1991. The osteohistology of femoral growth within a clade: a comparison of the crocodile Crocodylus, the dinosaurs Massospondylus and Syntarsus, and the birds Struthio and Sagittarius. Unpublished PhD thesis, University of Witwatersrand, 200 pp.
31. CHINSAMY, A. 1993. Bone histology and growth trajectory of the prosauropod dinosaur Massospondylus carinatus Owen. Modern Geology, 18, 319-329.
32. CHINSAMY, A. 1995. Ontogenetic changes in the bone histology of the Late Jurassic ornithopod Dryosaurus lettowvorbecki. Journal of Vertebrate Paleontology, 15, 96-104.
33. CHINSAMY, A. and RUBIDGE, B. S. 1993. Dicynodont (Therapsida) bone histology: Phylogenetic and physiological implications. Palaeontologia Africana, 30, 97-102.
34. CHINSAMY, A., RICH, T. and VICKERS-RICH, P. 1998. Polar dinosaur bone histology. Journal of Vertebrate Paleontology, 18, 385-390.
35. CLUVER, M. A. and HOTTON, N. III 1981. The genera Dicynodon and Diictodon and their bearing on the classification of the Dicynodontia (Reptilia, Therapsida). Annals of the South African Museum, 83, 99-146.
36. CLUVER, M. A. and KING, G. M. 1983. A reassessment of the relationships of the Permian Dicynodontia (Reptilia, Therapsida) and a new classification of dicynodonts. Annals of the South African Museum, 91, 195-273.
37. COX, C. B. 1959. On the anatomy of a new dicynodont genus with evidence of the position of the tympanum. Proceedings of the Zoological Society of London, 132, 321-367.
38. COX, C. B. 1964. On the palate, dentition, and classification of the fossil reptile Endothiodon and related genera. American Museum Novitates, 2171, 1-25.
39. CRUICKSHANK, A. R. I. 1986. Biostratigraphy and classification of a new Triassic dicynodont from East Africa. Modern Geology, 10, 121-131.
40. De FAUW, S. L. 1986. The appendicular skeleton of African dicynodonts. Unpublished PhD thesis, Wayne State University, 284 pp.
41. DEMPSTER, A. P., LAIRD, N. M. and RUBIN, D. B. 1977. Maximum likelihood estimation from incomplete data via the EM algorithm. Journal of the Royal Statistical Society, Series B (Statistical Methodology), 39, 1-38.
42. DRYSDALL, A. R. and KITCHING, J. W. 1963. A re-examination of the Karroo succession and fossil localities of part of the Upper Luangwa Valley. Memoir of the Geological Survey of Northern Rhodesia, 1, 1-62.
43. ELDREDGE, N. 1995. Species, speciation, and the context of adaptive change in evolution. 39-63. In ERWIN, D. H. and ANSTEY, R. L. (eds). New approaches to speciation in the fossil record. Columbia University Press, New York, NY, 288 pp.
44. ENLOW, D. H. 1957. A comparative histological study of fossil and recent bone tissues. Part II. Texas Journal of Science, 9, 136-214.
45. ENLOW, D. H. and BROWN, S. O. 1956. A comparative histological study of fossil and recent bone tissues. Part I. Texas Journal of Science, 8, 405-443.
46. FAIRBARIN, D. J. 1997. Allometry for sexual size dimorphism: Pattern and process in the coevolution of body size in males and females. Annual Review of Ecology and Systematics, 28, 659-687.
47. FISHER, D. C. 1992. Stratigraphic parsimony. 124-129. In MADDISON, W. P. and MADDISON, D. R. (eds). Macclade, version 3.0 Manual. Sinauer Associates, Sunderland, MA, 404 pp.
48. FISHER, D. C. 1994. Stratocladistics: Morphological and temporal patterns and their relation to phylogenetic process. 133-171. In GRANDE, L. and RIEPPEL, O. (eds). Interpreting the hierarchy of nature. Academic Press, San Diego, CA, 298 pp.
49. FRANCILLON-VIELLOT, H., BUFFRÉNIL, V., CASTANET, J., GERAUDIE, J., MEUNIER, F. J., SIRE, J. Y., ZYLBEBERG, L. and RICQLÈS, A. DE 1990. Microstructure and mineralization of vertebrate skeletal tissues. 471-530. In CARTER, J. G. (ed.). Skeletal biomineralization: Patterns, process and evolutionary trends 1. Van Nostrand Reinhold, New York, NY, 832 pp.
50. GAY, S. A. and CRUICKSHANK, A. R. I. 1999. Biostratigraphy of the Permian tetrapod faunas from the Ruhuhu Valley, Tanzania. Journal of African Earth Sciences, 29, 195-210.
51. GOLUBEV, V. K. 2005. Permian tetrapod stratigraphy. 95-99. In LUCAS, S. G. and ZEIGLER, K. E. (eds). The non-marine Permian. Bulletin of the New Mexico Museum of Natural History and Science, 30, 362 pp.
52. GOODALL, C. R. 1991. Procrustes methods in the statistical analysis of shape. Journal of the Royal Statistical Society, Series B (Statistical Methodology), 53, 285-339.
53. GOULD, S. J. 2002. The structure of evolutionary theory. Belknap Press, Cambridge, MA, 1433 pp.
54. HANCOX, P. J. and RUBIDGE, B. S. 1996. The first specimen of the Middle Triassic dicynodont Angonisaurus from the Karoo of South Africa: implications for the dating and biostratigraphy of the Cynognathus Assemblage Zone, Upper Beaufort Group. South African Journal of Science, 92, 391-392.
55. HANCOX, P. J. 2001. Breakthroughs in the biodiversity, biogeography, biostratigraphy, and basin analysis of the Beaufort Group. Journal of African Earth Sciences, 33, 563-577.
56. HAUGHTON, S. H. 1917. Investigations in South African fossil reptiles and Amphibia. Part 10. Descriptive catalogue of the Anomodontia, with especial reference to the examples in the South African Museum. Annals of the South African Museum, 12, 127-174.
57. HAUGHTON, S. H. 1926. On Karoo vertebrates from Nyasaland. Transactions of the Geological Society of South Africa, 27, 69-83.
58. HONAKER, J., JOSEPH, A., KING, G., SCHEVE, K. and SINGH, N. 2003. Amelia, a program for missing data, Version 2.1. Harvard University, Cambridge, MA. Freely available at: http://GKing.Harvard.Edu/.
59. HORNER, J. R. 2000. Long bone histology of the hadrosaurid dinosaur Maiasaura peeblesorum: Growth dynamics and physiology of an ontogenetic series of skeletal elements. Journal of Vertebrate Paleontology, 20, 115-129.
60. HORNER, J. R., RICQLÈS, A. de and PADIAN, K. 1999. Variation in dinosaur skeletochronology indicators: Implications for age assessment and physiology. Paleobiology, 25, 295-304.
61. HUTTON, J. M. 1986. Age determination of living Nile crocodiles from the cortical stratification of bone. Copeia, 2, 332-341.
62. IVAKHNENKO, M. F. 1994. A new Late Permian dromasaurian (Anomodontia) from Eastern Europe. Paleontological Journal, 28, 96-103.
63. JACOBS, L. L., WINKLER, D. A., NEWMAN, K. D., GOMAIN, E. M. and DEINO, A. 2005. Therapsids from the Permian Chiweta Beds and the age of the Karoo Supergroup in Malawi. Palaeontologia Electronica, 8, 1-23.
64. KEYSER, A. W. 1973a. A re-evaluation of the genus Tropidostoma Seeley. Palaeontologia Africana, 16, 25-35.
65. KEYSER, A. W. 1973b. A preliminary study of the type area of the Cistecephalus Zone of the Beaufort Series, and a revision of the anomodont family Cistecephalidae. Memoirs of the Geological Survey of South Africa, 62, 1-71.
66. KEYSER, A. W. 1975. A reevaluation of the cranial morphology and systematics of some tuskless Anomodontia. Memoirs of the Geological Survey of South Africa, 67, 1-110.
67. KEYSER, A. W. 1993. A re-evaluation of the smaller endothiodontidae. Memoirs of the Geological Survey of South Africa, 82, 1-53.
68. KEYSER, A. W. and SMITH, R. M. H. 1977-78. Vertebrate biozonation of the Beaufort Group with special reference to the western Karoo Basin. Annals of the Geological Survey of South Africa, 12, 1-35.
69. KING, G. M. 1981. The postcranial skeleton of Robertia broomiana, an early dicynodont (Reptilia, Therapsida) from the South African Karoo. Annals of the South African Museum, 84, 203-231.
70. KING, G. M. 1985. The postcranial skeleton of Kingoria nowacki (von Huene) (Therapsida: Dicynodontia). Zoological Journal of the Linnean Society, 84, 263-289.
71. KING, G. M. 1988. Anomodontia. In WELLNHOFER, P. (ed.). Handbuch der Paläoherpetologie, Part 17C. Fischer Verlag, Stuttgart, 174 pp.
72. KING, G. M. 1990. The dicynodonts: A study in palaeobiology. Chapman and Hall, London, 233 pp.
73. KING, G. M. 1992. The palaeobiogeography of Permian anomodonts. Terra Nova, 4, 633-640.
74. KING, G. M. 1993. How many species of Diictodon were there? Annals of the South African Museum, 102, 303-325.
75. KING, G. M., HONAKER, J., JOSEPH, A. and SCHEVE, K. 2001. Analyzing incomplete political science data: An alternative algorithm for multiple imputation. American Political Science Review, 95, 49-69.
76. KITCHING, J. W. 1963. The fossil localities and mammal-like reptiles of the Upper Luangwa Valley, Northern Rhodesia. South African Journal of Science, 59, 259-264.
77. KURKIN, A. A. 2000. New dicynodonts from the Upper Permian of the Vyatka Basin. Paleontological Journal, 34, S203-S210.
78. LUCAS, S. G. 1998. Global Triassic tetrapod biostratigraphy and biochronology. Palaeogeography, Palaeoclimatology, Palaeoecology, 143, 347-384.
79. LUCAS, S. G. 2002. Tetrapods and the subdivision of Permian time. 471-491. In HILLS, L. V., HENDERSON, C. M. and BAMBER, E. W. (eds). Carboniferous and Permian of the world. Canadian Society of Petroleum Geologists, Memoir, 19, 947 pp.
80. LUCAS, S. G. 2004. A global hiatus in the Middle Permian tetrapod fossil record. Stratigraphy, 1, 47-64.
81. LYDEKKER, R. 1890. Catalogue of the fossil Reptilia and Amphibia in the British Museum (Natural History). Part IV. Containing the orders Anomodontia, Ecaudata, Caudata, and Labyrinthodontia; and supplement. British Museum (Natural History), London, 295 pp.
82. MAISCH, M. W. 1999. The tetrapods from the Late Permian of Tanzania in the collections of the Institut und Museum für Geologie und Paläontologie der Universität Tübingen, with special reference to the pristerodontian dicynodonts Rhachiocephalus and Pelanomodon. Unpublished PhD thesis, Universität Tübingen, 350 pp.
83. MAISCH, M. W. 2002a. A new basal lystrosaurid dicynodont from the Upper Permian of South Africa. Palaeontology, 45, 343-359.
84. MAISCH, M. W. 2002b. Observations on Karoo and Gondwana vertebrates. Part 4: The taxonomic status of the Late Permian rhachiocephalid Platycyclops crassus Broom, 1948 (Therapsida: Dicynodontia) from the South African Karoo. Neues Jahrbuch für Geologie und Paläontologie Monatshefte, 2002, 362-372.
85. MAZIN, J. M. and KING, G. M. 1991. The first dicynodont from the Late Permian of Malagasy. Palaeontology, 34, 837-842.
86. OWEN, R. 1860. On some reptilian fossils from South Africa. Quarterly Journal of the Geological Society of London, 16, 49-63.
87. RALLS, K. 1976. Mammals in which females are larger than males. Quarterly Review of Biology, 51, 245-276.
88. RAY, S. 2001. Small Permian dicynodonts from India. Paleontological Research, 5, 177-191.
89. RAY, S. 2005. Lystrosaurus (Therapsida, Dicynodontia) from India: taxonomy, relative growth, and cranial dimorphism. Journal of Systematic Palaeontology, 3, 203-221.
90. RAY, S., BOTHA, J. and CHINSAMY, A. 2004. Bone histology and growth patterns of some non-mammalian therapsids. Journal of Vertebrate Paleontology, 24, 634-648.
91. RAY, S. CHINSAMY, A. and BANDYOPADHYAY, S. 2005. Lystrosaurus murrayi (Therapsida, Dicynodontia): Bone histology, growth and lifestyle adaptations. Palaeontology, 48, 1169-1186.
92. REID, R. E. H. 1996. Bone histology of the Cleveland-Lloyd dinosaurs and of dinosaurs in general, Part I: Introduction: Introduction to bone tissues. Geology Studies, 41, 25-71.
93. RICQLES, A. de 1972. Recherches paléohistologiques sur les os longs des tétrapodes. III. -titanosuchiens, dinocéphales et dicynodontes. Annales de Paléontologie (Vertébrés), 58, 17-60.
94. RICQLES, A. de 1974. Evolution of endothermy: Histological evidence. Evolutionary Theory, 1, 51-80.
95. RICQLES, A. de 1976. On bone histology of fossil and living reptiles, with comments on its functional and evolutionary significance. 123-150. In BELLAIRS, A. D'A. and COX, C. B. (eds). Morphology and biology of reptiles. Academic Press, London, 290 pp.
96. RICQLES, A. de 1980. Tissue structures of dinosaur bone. Functional significance and possible relation to dinosaur physiology. 103-139. In THOMAS, D. K. and OLSON, E. C. (eds). A cold look at the warm-blooded dinosaurs. Westview Press, Boulder, CO, 514 pp.
97. RICQLES, A. de, MEUNIER, F. J., CASTANET, J. and FRANCILLON-VIEILLOT, H. 1991. Comparative microstructure of bone. 1-77. In HALL, B. K. (ed.). Bone. 3. Bone matrix and bone specific products. CRC Press, Boca Raton, FL, 500 pp.
98. ROHLF, F. J. 1990. Rotational fit (Procrustes) method. 227-236. In ROHLF, F. J. and BOOKSTEIN, F. L. (eds). Proceedings of the Michigan Morphometrics Workshop. University of Michigan, Museum of Zoology, Special Publication, 2, 380 pp.
99. ROHLF, F. J. 1998. On applications of geometric morphometrics to studies of ontogeny and phylogeny. Systematic Biology, 47, 147-158.
100. ROHLF, F. J. 2003a. TpsDig 1.39. Department of Ecology and Evolution, State University of New York, Stony Brook, NY.
101. ROHLF, F. J. 2003b. TpsSmall 1.20. Department of Ecology and Evolution, State University of New York, Stony Brook, NY.
102. ROHLF, F. J. 2004. TpsRegr 1.28. Department of Ecology and Evolution, State University of New York, Stony Brook, NY.
103. ROHLF, F. J. and MARCUS, L. F. 1993. A revolution in morphometrics. Trends in Ecology and Evolution, 8, 129-132.
104. RUBIDGE, B. S. 1984. The cranial morphology and palaeoenvironment of Eodicynodon Barry (Therapsida: Dicynodontia). Navorsinge van die Nasionale Museum Bloemfontein, 4, 326-402.
105. RUBIDGE, B. S. (ed.) 1995. Biostratigraphy of the Beaufort Group (Karoo Supergroup). Council for Geoscience, Pretoria, 46 pp.
106. RUBIDGE, B. S. 2005. Re-uniting lost continents - fossil reptiles from the ancient Karoo and their wanderlust. South African Journal of Geology, 108, 135-172.
107. SEELEY, H. G. 1889. Researches on the structure, organization, and classification of the fossil Reptilia. -VI. On the anomodont Reptilia and their allies. Philosophical Transactions of the Royal Society, Series B, 180, 215-296.
108. SHEETS, H. D. 2001. Standard 6 beta. Freely available at: http://www2.canisius.edu/~sheets/morphsoft.html.
109. SHEETS, H. D. 2002. CoordGen 6d. Freely available at: http://www2.canisius.edu/~sheets/morphsoft.html.
110. SHEETS, H. D. 2003a. Regress 6k. Freely available at: http://www2.canisius.edu/~sheets/morphsoft.html.
111. SHEETS, H. D. 2003b. PCAGen 6n. Freely available at: http://www2.canisius.edu/~sheets/morphsoft.html.
112. SHEETS, H. D. 2003c. CVAGen 6j. Freely available at: http://www2.canisius.edu/~sheets/morphsoft.html.
113. SHEETS, H. D. 2003d. TwoGroup 6h. Freely available at: http://www2.canisius.edu/~sheets/morphsoft.html.
114. SHINE, R. 1988. The evolution of large body size in females: A critique of Darwin's 'Fecundity Advantage' model. American Naturalist, 131, 124-131.
115. SULLIVAN, C. and REISZ, R. R. 2005. Cranial anatomy and taxonomy of the Late Permian dicynodont Diictodon. Annals of the Carnegie Museum, 74, 45-75.
116. SULLIVAN, C., REISZ, R. R. and SMITH, R. M. H. 2003. The Permian mammal-like herbivore Diictodon, the oldest known example of sexually dimorphic armament. Proceedings of the Royal Society of London, Series B, 270, 173-178.
117. SURKOV, M. V., KALANDADZE, N. N. and BENTON, M. J. 2005. Lystrosaurus georgi, a dicynodont from the Lower Triassic of Russia. Journal of Vertebrate Paleontology, 25, 402-413.
118. THACKERAY, J. F., DURAND, J. F. and MEYER, L. 1998. Morphometric analysis of South African dicynodonts attributed to Lystrosaurus murrayi (Huxley, 1859) and L. declivis (Owen, 1860): Probabilities of conspecificity. Annals of the Transvaal Museum, 36, 413-420.
119. TOERIEN, M. J. 1953. The evolution of the palate in South African Anomodontia and its classificatory significance. Palaeontologia Africana, 1, 49-118.
120. TOLLMAN, S. M., GRINE, F. E. and HAHN, B. D. 1980. Ontogeny and sexual dimorphism in Aulacephalodon (Reptilia, Anomodontia). Annals of the South African Museum, 81, 159-186.
121. WAGNER, P. J. and ERWIN, D. H. 1995. Phylogenetic patterns as tests of speciation hypotheses. 87-122. In ERWIN, D. H. and ANSTEY, R. L. (eds). New approaches to speciation in the fossil record. Columbia University Press, New York, NY, 288 pp.
122. WATSON, D. M. S. 1914. Dicynodon halli, sp. n., an anomodont reptile from South Africa. Annals and Magazine of Natural History, 8 (14), 95-97.
123. WATSON, D. M. S. 1917. The evolution of the tetrapod shoulder girdle and forelimb. Journal of Anatomy, 52, 1-63.
124. WATSON, D. M. S. 1948. Dicynodon and its allies. Proceedings of the Zoological Society of London, 118, 823-877.
125. WATSON, D. M. S. 1960. The anomodont skeleton. Transactions of the Zoological Society of London, 29, 131-208.
126. ZELDITCH, M. L., SWIDERSKI, D. L., SHEETS, H. D. and FINK, W. L. 2004. Geometric morphometrics for biologists: A primer. Elsevier, Amsterdam, 443 pp.