The interrelationships of Devonian lungfishes (Sarcopterygii: Dipnoi) as inferred from neurocranial evidence and new data from the genus Soederberghia Lehman, 1959

Zoological Journal of the Linnean Society

Volumn 151, Issue 1, 2007, Pages 115-171

  Friedman, Matt ^a,b  

^a UNIVERSITY OF CHICAGO   (United States)

^b UNIVERSITY MUSEUM OF ZOOLOGY   (United Kingdom)

Author keywords

Bayesian inference; Braincases; Chirodipterus; Famennian; Griphognathus; Lungfish phylogeny; Rhynchodipteridae

Indexed keywords

DIPNOI; DIPTERIDAE; HOLODONTIDAE; RHYNCHODIPTERIDAE; SARCOPTERYGII;

EID: 34548531436     PISSN: 00244082     EISSN: 10963642     Source Type: Journal    
DOI: 10.1111/j.1096-3642.2007.00320.x     Document Type: Article

References (231)

1. Agar WE. 1906. The spiracular gill cleft in Lepidosiren and Protopterus. Anatomischer Anzeiger 28 : 298 304.
2. Agassiz L. 1833-1844. Recherches sur les Poissons Fossiles. V Volumes. Neuchâtel : Imprimerie de Petitpierre et Prince.
3. Ahlberg P. 1991. A re-examination of sarcopterygian interrelationships, with special reference to the Porolepiformes. Zoological Journal of the Linnean Society 103 : 241 287.
4. Ahlberg PE, Clack JA, Lukševiès E. 1996. Rapid braincase evolution between Panderichthys and the earliest tetrapods. Nature 381 : 61 64.
5. Ahlberg PE, Johanson Z. 1998. Osteolepiforms and the ancestry of tetrapods. Nature 395 : 792 794.
6. Ahlberg PE, Johanson Z, Daeschler EB. 2001. The Late Devonian lungfish Soederberghia (Sarcopterygii, Dipnoi) from Australia and North America, and its biogeographical implications. Journal of Vertebrate Paleontology 21 : 1 12.
7. Ahlberg PE, Smith MM, Johanson Z. 2006. Developmental plasticity and disparity in early dipnoan (lungfish) dentitions. Evolution and Development 8 : 331 349.
8. Andrews SMA. 1973. Interrelationships of crossopterygians. In : Greenwood PH, Miles RS, Patterson C, eds. Interrelationships of fishes. London : Academic Press, 137 177.
9. Andrews M, Long J, Ahlberg P, Barwick R, Campbell K. 2006. The structure of the sarcopterygian Onychodus jandemarrai n. sp. from Gogo, Western Australia: with a functional interpretation of the skeleton. Transactions of the Rosyal Society of Edinburgh: Earth Sciences 96 : 197 307.
10. Andrews SMA, Westoll TS. 1970. The postcranial skeleton of rhipidistian fishes excluding Eusthenopteron. Transactions of the Royal Society of Edinburgh 68 : 391 498.
11. Arratia G, Schultze H-P. 1991. Palatoquadrate and its ossifications: development and homology within osteichthyans. Journal of Morphology 208 : 1 81.
12. Bartsch P. 1994. Development of the cranium of Neoceratodus forsteri, with a discussion of the suspensorium and opercular apparatus in Dipnoi. Zoomorphology 114 : 1 31.
13. Barwick RE, Campbell KSW. 1996. A Late Devonian dipnoan, Pillararhynchus, from Gogo, Western Australia, and its relationships. Palaeontographica Abteilung A 239 : 1 42.
14. Barwick RE, Campbell KSW, Mark-Kurik E. 1995. Tarachomylax: a new Early Devonian dipnoan from Severnaya Zemlya, and its place in the evolution of the Dipnoi. Geobios 30 : 45 73.
15. Basden AM, Young GC. 2001. A primitive actinopterygian neurocranium from the Early Devonian of southeastern Australia. Journal of Vertebrate Paleontology 21 : 754 766.
16. Basden AM, Young GC, Coates MI, Ritchie A. 2000. The most primitive osteichthyan braincase? Nature 403 : 185 188.
17. de Beer GR. 1937. The development of the vertebrate skull. Oxford : Oxford University Press.
18. Bemis WE. 1987. Feeding systems of living Dipnoi: anatomy and function. Journal of Morphology Supplement 1 : 249 275.
19. Bemis WE, Forey PL. 2001. Occipital structure and the posterior limit of the skull in actinopterygians. In : Ahlberg PE, ed. Major events in early vertebrate evolution: palaeontology, phylogeny, genetics and development. London : Taylor & Francis, 350 369.
20. Bernacsek GM. 1977. A lungfish cranium from the Middle Devonian of the Yukon Territory, Canada. Palaeontographica Abteilung A 157 : 175 200.
21. Bertmar G. 1965. On the development of the jugular and cerebral veins in fishes. Proceedings of the Zoological Society of London 144 : 87 130.
22. Bertmar G. 1968. Evolution and phylogeny in lungfishes. Acta Zoologica 49 : 189 201.
23. Bischoff TLW. 1840. Lepiosiren paraxoxa. Anatomische Untersuch und Beschreiben. Leipzig: Leopold Voss.
24. Bjerring HC. 1967. Does a homology exist between the basicranial muscle and the polar cartilage? Colloque international. CNRS, Problèmes Actuels de Paléontolgie-Évolution des Vertébrés 163 : 223 267.
25. Bjerring HC. 1971. The nerve supply to the second metamere basicranial muscle in osteolepiform vertebrates, with some remarks on the basic composition of the endocranium. Acta Zoologica 52 : 189 225.
26. Bjerring HC. 1978. The 'intracranial joint' versus the 'ventral otic fissure'. Acta Zoologica 59 : 203 214.
27. Blom H, Clack JA, Ahlberg PE, Friedman M. 2007. Devonian vertebrates from East Greenland: a review of faunal composition and distribution. Geodiversitas 29 : 119 144.
28. Bremer K. 1988. The limits of amnio-acid sequence data in angiosperm phylogenetic reconstruction. Evolution 42 : 795 803.
29. Brinkmann H, Venkatesh B, Brenner S, Meyer A. 2004. Nuclear protein-coding genes support lungfish and not the coelacanth as the closest living relatives of land vertebrates. Proceedings of the National Academy of Sciences of the United States of America 101 : 4900 4905.
30. Campbell KSW, Barwick RE. 1982. The neurocranium of the primitive dipnoan Dipnorhynchus sussmilchi (Etheridge). Journal of Vertebrate Paleontology 2 : 286 327.
31. Campbell KSW, Barwick RE. 1984. Speonesydrion, an Early Devonian dipnoan with primitive toothplates. Palaeoichthyologica 2 : 1 48.
32. Campbell KSW, Barwick RE. 1985. An advanced dipnorhynchid lungfish from the Early Devonian of New South Wales, Australia. Records of the Australian Museum 37 : 301 316.
33. Campbell KSW, Barwick RE. 1987. Paleozoic lungfishes - a review. Journal of Morphology Supplement 1 : 93 131.
34. Campbell KSW, Barwick RE. 1988. Uranolophus: a reappraisal of a primitive dipnoan. Memoir of the Association of Australasian Palaeontologists 7 : 87 144.
35. Campbell KSW, Barwick RE. 1990. Paleozoic dipnoan phylogeny: functional complexes and evolution without parsimony. Paleobiology 16 : 143 169.
36. Campbell KSW, Barwick RE. 1998. A new tooth-plated dipnoan from the Upper Devonian Gogo Formation and its relationships. Memoirs of the Queensland Museum 42 : 403 437.
37. Campbell KSW, Barwick RE. 2000. The braincase, mandible and dental structures of the Early Devonian lungfish Dipnorhynchus kurikae from Wee Jasper, New South Wales. Records of the Australian Museum 52 : 103 128.
38. Campbell KSW, Barwick RE. 2001. Diabolepis and its relationship to the Dipnoi. Journal of Vertebrate Paleontology 21 : 227 241.
39. Campbell KSW, Barwick RE. 2002. The axial postcranial structure of Griphognathus whitei from the Upper Devonian Gogo Formation of Western Australia: comparisons with other Devonian dipnoans. Records of the Western Australian Museum 21 : 167 201.
40. Campbell KSW, Barwick RE, Lindley ID. 2000. New data on the structure of the early Devonian lungfish. Dipnorhynchus. Alcheringa 24 : 277 298.
41. Campbell KSW, Smith MM. 1987. The Devonian dipnoan Holodipterus: dental form variation and remodelling growth mechanisms. Records of the Australian Museum 39 : 131 167.
42. Chang M-M. 1982. The braincase of Youngolepis, a Lower Devonian crossopterygian from Yunnan, south-western China. PhD thesis, Department of Geology, University of Stockholm, and Section of Paleozoology, Swedish Museum of Natural History.
43. Chang M-M, Yu X. 1984. Structure and phylogenetic significance of Diabolichthys speratus gen. et sp. nov., a new dipnoan-like form from the Lower Devonian of eastern Yunnan, China. Proceedings of the Linnean Society, New South Wales 107 : 171 184.
44. Chang M-M. 1991. Rhipidistians, dipnoans and tetrapods. In : Schultze HP, Trueb L, eds. Origins of the higher groups of tetrapods: controversy and consenus. Ithaca, NY : Cornell University Press, 3 28.
45. Chang M-M. 1995. Diabolepis and its bearing upon the relationships between porolepiforms and dipnoans. Bulletin Museum National d'Histoire Naturelle, Paris, 4e Series 17 : 235 268.
46. Chang M-M. 2004. Synapomorphies and scenarios - more characters of Youngolepis betraying its affinity to the Dipnoi. In : Arratia G, Wilson MVH, Cloutier R, eds. Recent advances in the origin and early radiation of vertebrates. München : Verlag Dr Friedrich Pfeil, 665 686.
47. Chang M-M, Wang J-Q. 1995. A new Emsian dipnorhynchid (Dipnoi) from Guangnan, southeastern Yunna, China. Geobios 19 : 233 239.
48. Clack JA. 1994. Acanthostega gunnari, a Devonian tetrapod from Greenland; the snout, palate and ventral parts of the braincase, with a discussion of their significance. Meddelelser om Grønland 31 : 1 24.
49. Clack JA. 1998. The neurocranium of Acanthostega gunnari Jarvik and the evolution of the otic region in tetrapods. Zoological Journal of the Linnean Society 122 : 61 97.
50. Clack JA, Ahlberg PE, Finney SM, Dominguez Alonso P, Robinson J, Ketcham RA. 2003. A uniquely specialized ear in a very early tetrapod. Nature 425 : 65 69.
51. Clément G. 2001. Evidence for the lack of choanae in the porolepiformes. Journal of Vertebrate Paleontology 21 : 795 802.
52. Clément G, Boisvert CA. 2006. Lohest's true and false 'Devonian amphibians': evidence for the rhynchodipterid lungfish Soederberghia in the Famennian of Belgium. Journal of Vertebrate Paleontolology 26 : 276 283.
53. Clément G, Janvier P. 2004. Powichthys spitsbergensis sp. nov., a new member of the Dipnomorpha (Sarcopterygii, lobe-finned fishes) from the Lower Devonian of Siptsbergen, with remarks on basal dipnomorph anatomy. Fossils and Strata 50 : 92 112.
54. Cloutier R. 1996a. Dipnoi (Akinetia: Sarcopterygii). In : Schultze H-P, Cloutier R, eds. Devonian fishes and plants of Miguasha, Quebec, Canada. München : Verlag Dr Friedrich Pfeil, 198 226.
55. Cloutier R. 1996b. The primitive actinistian Miguashaia bureaui Schultze (Sarcopterygii). In : Schultze H-P, Cloutier R, eds. Devonian fishes and plants of Miguasha, Quebec, Canada. München : Verlag Dr Friedrich Pfeil, 227 247.
56. Cloutier R. 1997. Morphologie et variation du toit crânien du dipneuste Scaumenacia curta (Whiteaves) (Sarcopterygii), du Dévonien supérieur du Québec. Geodiversitas 19 : 61 105.
57. Cloutier R, Ahlberg PE. 1996. Morphology, characters, and the interrelationships of basal sarcopterygians. In : Stiassny MLJ, Parenti LR, Johnson GD, eds. Interrelationships of fishes. San Diego : Academic Press, 445 479.
58. Cloutier R, Arratia G. 2004. Early diversification of actinopterygians. In : Arratia G, Wilson MVH, Cloutier R, eds. Recent advances in the origin and early radiation of vertebrates. München : Verlag Dr Friedrich Pfeil, 217 270.
59. Coates MI. 1999. Endocranial preservation of a Carboniferous actinopterygian from Lancashire, UK, and the interrelationships of primitive actinopterygians. Philosophical Transactions of the Royal Society of London, Series B 354 : 435 462.
60. Coates MI, Sequeira SEK. 1998. The braincase of a primitive shark. Transactions of the Royal Society of Edinburgh: Earth Sciences 89 : 63 85.
61. Coates MI, Sequeira SEK. 2001a. Early sharks and primitive gnathostome interrelationships. In : Ahlberg PE, ed. Major events in early vertebrate evolution: palaeontology, phylogeny, genetics and development. London : Taylor & Francis, 241 262.
62. Coates MI, Sequeira SEK. 2001b. A new stethacanthid condrichthyan from the Lower Carboniferous of Bearsden, Scotland. Journal of Vertebrate Paletontology 21 : 438 459.
63. Denison R. 1968. Early Devonian lungfishes from Wyoming, Utah, and Idaho. Fieldiana: Geology 17 : 353 413.
64. Denison R. 1978. Placodermi. In : Schultze H-P, ed. Handbook of paleoichthyology, Vol. 2. Stuttgart : Gustav Fischer Verlag, 1 128.
65. Denison R. 1979. Acanthodii. In : Schultze H-P, ed. Handbook of paleoichthyology, Vol. 2. Stuttgart : Gustav Fischer Verlag, 1 62.
66. Felsenstein J. 1985. Confidence limits on phylogenies: an approach using the bootstrap. Evolution 39 : 783 791.
67. Felsenstein J. 2004. Inferring phylogenies. Sunderland, MA : Sinauer Associates.
68. Forey PL. 1987. Relationships of lungfishes. Journal of Morphology Supplement 1 : 75 91.
69. Forey PL. 1998. History of the coelacanth fishes. London : Chapman & Hall.
70. Forey PL, Gardiner BG. 1986. Observations on Ctenurella (Ptyctodontida) and the classification of placoderm fishes. Zoological Journal of the Linnean Society 86 : 43 74.
71. Forey PL, Gardiner BG, Patterson C. 1991. The lungfish, the coelacanth, and the cow revisted. In : Schultze HP, Trueb L, eds. Origins of the higher groups of tetrapods: controversy and consensus. Ithaca, NY : Cornell University Press, 145 172.
72. Forster-Cooper C. 1937. The Middle Devonian fish fauna of Achanarras. Transactions of the Royal Society of Edinburgh 59 : 1 35.
73. Fox H. 1965. Early development of the head and pharynx of Neoceratodus with a consideration of its phylogeny. Journal of Zoology 146 : 470 554.
74. Fox RC, Campbell KSW, Barwick RE, Long JA. 1995. A new osteolepiform fish from the Lower Carboniferous Raymond Formation, Drummond Formation, Queensland. Memoirs of the Queensland Museum 38 : 97 221.
75. Friedman M. 2003. New material of the Devonian lungfish Soederberghia groenlandica (Sarcopterygii: Dipnoi) from East Greenland, with comments on the status of the Rhynchodipteridae. Journal of Vertebrate Paleontology 23 (Suppl. 52A.
76. Friedman M. 2005. Braincases, bodies and biogeography: new characters and their bearing on the interrelationships of early lungfishes. Journal of Vertebrate Paleontology 25 (Suppl. 60A.
77. Friedman M. in press. Styloichthys as the oldest coelacanth: implications for early osteichthyan interrelationships. Journal of Systematic Palaeontology.
78. Friedman M, Blom H. 2006. A new actinopterygian fish from the Famennian of East Greenland and the relationships of Devonian ray-finned fishes. Journal of Paleontology 80 : 1186 1204.
79. Friend PF, Alexander-Marrack PD, Allen KC, Nicholson J, Yeats AK. 1983. Devonian sediments of East Greenland VI: review of results. Meddelelser om Grønland 206 : 1 96.
80. Gardiner BG. 1973. Interrelationships of teleostomes. In : Greenwood PH, Miles RS, Patterson C, eds. Interrelationships of fishes. London : Academic Press, 105 135.
81. Gardiner BG. 1984. The relationships of the palaeoniscid fishes, a review based on the new specimens of Mimia and Moythomasia from the Upper Devonian of Western Australia. Bulletin of the British Museum of Natural History (Geology) 37 : 173 428.
82. Gardiner BG, Schaeffer B. 1989. Interrelationships of lower actinopterygian fishes. Zoological Journal of the Linnean Society 97 : 135 187.
83. Gardiner BG, Schaeffer B, Masserie JA. 2005. A review of the lower actinopterygian phylogeny. Zoological Journal of the Linnean Society 144 : 511 526.
84. Goldman N, Anderson JP, Rodrigo AG. 2000. Likelihood-based tests of topologies in phylogenetics. Systematic Biology 49 : 652 670.
85. Goodrich ES. 1909. Cyclostomes and fishes. In : Lankester ER, ed. A treatise on zoology, part IX. London : Adam and Charles Black, 1 518.
86. Goodrich ES. 1930. Studies on the structure and development of vertebrates. London : Macmillan.
87. Goujet D. 1984. Les Poissons Placodermes du Spitsberg. Arthrodires Dolichothoraci de la Fromation de Wood Bay (Dévonien Inférieur). Paris : Editions du Centre National de la Recherche Scientifique.
88. Grande L, Bemis WE. 1991. Osteology and phylogenetic relationships of fossil and Recent paddlefishes (Polyodontidae) with comments on the interrelationships of Acipenseriformes. Society of Vertebrate Paleontology Memoir 1 : 1 121 supplement to Journal of Vertebrate Paleontology 11.
89. Grande L, Bemis WE. 1998. A comprehensive phylogenetic study of amiid fishes (Amiidae) based on comparative skeletal anatomy. An empirical search for interconnected patterns of natural history. Society of Vertebrate Paleontology Memoir 4 : 1 690 supplement to Journal of Vertebrate Paleontology 18.
90. Gray JE. 1856. Observations on a living African Lepidosiren in the Crystal Palace. Proceedings of the Zoological Society of London 24 : 342 348.
91. Gross W. 1933. Die Wirbeltiere des rhenischen Devons. Abhandlungen der Preussischen Geologischen Landesanstalt 154 : 1 83.
92. Gross W. 1956. Über Crossopterygier und Dipnoer aus dem baltischen Oberdevon im Zusammenhang einer vergleichenden Untersuchung des Porenkanalsystems paläozoischer Agnathen und Fische. Kungliga Svenska Vetenskapsakademiens Handlingar 5 : 1 140.
93. Hilton EJ. 2001. Tongue bite apparatus of osteoglossomorph fishes: variation of a character complex. Copeia 2001 : 372 381.
94. Holmes EB. 1985. Are lungfishes the sister-group of tetrapods? Biological Journal of the Linnean Society 25 : 379 394.
95. Holmgren N, Stensiö E. 1936. Kranium und Visceralskelett der Akranier, Cyclostomen und Fische. In : Bolk L, Göppert E, Kallius E Lubosch W, eds. Handbuch der Vergleichenden Anatomie der Wirbeltiere. Vierter Band. Berlin : Urban & Schwarzenberg, 233 500.
96. Huelsenbeck JP, Ronquist F. 2001. MRBAYES: Bayesian inference of phylogeny. Bioinformatics 17 : 754 755.
97. Huxley TH. 1880. On the applications of the laws of evolution to the arrangement of the Vertebrata and more particularly of the Mammalia. Proceedings of the Zoological Society of London 1880 : 649 662.
98. Jaekel O. 1927. Der Kopf der Wirbeltiere. Zeitschrift für die Gesamte Anatomie. Ergebnisse der Anatomie und Entwicklungsgeschichte 27 : 815 974.
99. Janvier P. 1985. Les Céphalaspides du Spitsberg. Paris : Éditions du Centre National del la Recherche Scientifique.
100. Janvier P. 1993. Patterns of diversity in the skull of jawless fishes. In : Hanken J, Hall BK, eds. The skull, Vol. 2. Patterns of structural and systematic diversity. Chicago : University of Chicago Press, 131 188.
101. Janvier P. 1996. Early vertebrates. Oxford : Clarendon Press.
102. Janvier P. 2004. Wandering nostrils. Nature 432 : 23.
103. Janvier P, Lethiers F, Monod O, Balkaş O. 1984. Discovery of a vertebrate fauna at the Devonian-Carboniferous boundary in SE Turkey (Hakkari Province). Journal of Petroleum Geology 7 : 147 168.
104. Janvier P, Martin M. 1978. Vertébrés Dévoniens de l'Iran central. I: Dipneustes. Geobios 11 : 819 832.
105. Janvier P, Suárez-Riglos M. 1986. The Silurian and Devonian vertebrates of Bolivia. Bulletin de l'Institut Français d'Etudes Andines 15 : 73 114.
106. Jarvik E. 1942. On the structure of the snout of crossopterygians and lower gnathostomes in general. Zoologiska Bidrag Fran Uppsala 21 : 235 675.
107. Jarvik E. 1954. On the visceral skeleton in Eusthenopteron with a discussion of the parasphenoid and palatoquadrate in fishes. Kungliga Svenska Vetenskapsakademiens Handlingar, 4 Serien 5 : 1 104.
108. Jarvik E. 1961. Devonian vertebrates. In : Raasch GO, ed. Geology of the Arctic 1. Toronto : University of Toronto Press, 197 204.
109. Jarvik E. 1967. Remarks on the structure of the snout in Megalichthys and certain other rhipidistid crossopterygians. Arkiv för Zoologi 19 : 41 98.
110. Jarvik E. 1968. The systematic position of the Dipnoi. In : Ørvig T, ed. Current problems of lower vertebrate phylogeny. Stockholm : Almqvist and Wiksell, 223 245.
111. Jarvik E. 1972. Middle and Upper Devonian Porolepiformes from East Greenland with special reference to Glyptolepis groenlandica n.sp. Meddelelser om Grønland 182 : 1 307.
112. Jarvik E. 1977. The systematic position of acanthodian fishes. In : Andrews SM, Miles RS, Walker AD, eds. Problems in vertebrate evolution. London : Academic Press, 199 225.
113. Jarvik E. 1980. Basic structure and evolution of vertebrates, Vol. 1. London : Academic Press.
114. Jarvik E. 1996. The Devonian tetrapod Ichthyostega. Fossils and Strata 40 : 1 213.
115. Jessen HL. 1966. Die Crossopterygier des Oberen Plattenkalkes (Devon) der Bergisch-Gladbach - Paffrather Mulde (Rheinisches Schiefergebirge) unter Berücksichtigung von amerikanischem und europäischem Onychodus-Material. Arkiv för Zoologi 18 : 305 389.
116. Jessen HL. 1975. A new choanate fish, Powichthys thorsteinssoni n. g., n. sp., from the early Lower Devonian of the Canadian Arctic Archipelago. Colloque international. CNRS, Problèmes Actuels de Paléontolgie- Évolution des Vertébrés 218 : 213 222.
117. Jessen HL. 1980. Lower Devonian Porolepiformes from the Canadian Arctic with special reference to Powichthys thorsteinssoni Jessen. Palaeontographica Abteilung A 167 : 180 214.
118. Kass RE, Raftery AE. 1995. Bayes factors. Journal of the American Statistical Association 90 : 773 795.
119. Kemp A. 1999a. Ontogeny of the skull of the Australian lungfish Neoceratodus forsteri (Osteichthyes: Dipnoi). Journal of Zoology, London 248 : 97 137.
120. Kemp A. 1999b. Anomalies in skull bones of the Australian lungfish, Neoceratodus forsteri, compared with aberrations in fossil dipnoan skulls. Journal of Vertebrate Paleontology 19 : 407 429.
121. Kemp A. 2000. Chirodipterus potteri, a new Devonian lungfish from New South Wales, Australia: and the ontogeny of chirodipterid toothplates. Journal of Vertebrate Paleontology 20 : 665 674.
122. Kesteven HL. 1950. The origin of the tetrapods. Proceedings of the Royal Society of Victoria 59 : 93 138.
123. Krupina NI. 1992. Some data on the ontogeny of Devonian dipnoans. In : Mark-Kurik E, ed. Fossil fishes as living animals. Tallinn : Academy of Sciences of Estonia, 215 222.
124. Krupina NI. 1995. New species of Rhinodipterus (Dipnoi) from the Upper Devonian of North Western Russia. Geobios Mémoire Spécial 19 : 269 274.
125. Krupina NI, Reisz RR. 2000. Phylogeny of Devonian dipnoans, with special emphasis on forms from the Russian platform. Paleontological Journal 34 : S147 S153.
126. Krupina NI, Reisz RR, Scott D. 2001. The skull and tooth system of Orlovichthys limnatis, a Late Devonian dipnoan from Russia. Canadian Journal of Earth Sciences 38 : 1301 1311.
127. Kulczycki J. 1960. Porolepis (Crossopterygii) from the Lower Devonian of the Holy Cross Mountains. Acta Palaeontologica Polonica 5 : 5 105.
128. Lebedev OA. 1995. Morphology of a new osteolepidid fish from Russia. Bulletin Museum National d'Histoire Naturelle, Paris, 4e Series 17 : 287 341.
129. Lebedev OA. 2004. A new tetrapod Jakubsonia livensis from the early Famennian (Devonian) of Russia and palaeoecological remarks on the Late Devonian tetrapod habitats. Acta Universitatis Latviensia 679 : 79 98.
130. Lehman J-P. 1959. Les dipnuestes du Dévonien supérieur du Groenland. Meddelelser om Grønland 160 : 1 58.
131. Lehmann W, Westoll TS. 1952. A primitive dipnoan fish from the Lower Devonian of Germany. Proceedings of the Royal Society B 140 : 403 421.
132. Long JA. 1985. A new osteolepidid fish from the Upper Devonian Gogo Formation, Western Australia. Records of the Western Australian Museum 12 : 361 377.
133. Long JA. 1987. A redescription of the lungfish Eoctenodus Hills 1929, with reassessment of other Australian records of the genus Dipterus Sedgwick & Murchison 1828. Records of the Western Autralian Museum 13 : 297 314.
134. Long JA. 1988. New palaeoniscoid fishes from the Late Devonian and Early Carboniferous of Victoria. Memoir of the Australasian Association of Palaeontologists 12 : 1 64.
135. Long JA. 1992a. Gogodipterus paddyensis (Miles), gen. nov., a new chirodipterid lungfish from the Late Devonian Gogo Formation, Western Australia. Beagle, Records of the Northern Territory Museum of Arts and Sciences 9 : 11 20.
136. Long JA. 1992b. Cranial anatomy of two new Late Devonian lungfishes (Pisces: Dipnoi) from Mount Howitt, Victoria. Records of the Australian Museum 44 : 299 318.
137. Long JA. 1993. Cranial ribs in Devonian lungfishes and the origins of dipnoan air-breathing. Memoir of Australasian Palaeontologists 15 : 199 209.
138. Long JA. 2001. The relationships of Psarolepis and the onychodontiform fishes. Journal of Vertebrate Paleontology 21 : 815 820.
139. Long JA. 2005. Interrelationships of the holodontid lungfishes (Osteichthyes, Dipnomorpha) based on new material from the Upper Devonian Gogo Formation of Western Australia. Journal of Vertebrate Paleontology 25 : 84A.
140. Long JA, Barwick RE, Campbell KSW. 1997. Osteology and functional morphology of the osteolepiform fish Gogonasus andrewsae Long, 1985, from the Upper Devonian Gogo Formation, Western Australia. Records of the Western Australian Museum Supplement 53 : 1 89.
141. Maddison WP, Maddison DR. 2000. Macclade: analysis of phylogeny and character evolution, Version 4.0. Sunderland, MA : Sinauer Associates.
142. Maisey JG. 1986. Heads and tails: a chordate phylogeny. Cladistics 2 : 201 256.
143. Maisey JG. 2001. A primitive chondrichthyan braincase from the Middle Devonian of Bolivia. In : Ahlberg PE, ed. Major events in early vertebrate evolution. London : Taylor & Francis, 263 288.
144. Maisey JG. 2005. Braincase of the Upper Devonian shark Cladodoides wildungensis (Chondrichthyes, Elasmobranchii), with observations on the braincase in early chondrichthyans. Bulletin of the American Museum of Natural History 288 : 1 103.
145. Maisey JG, Anderson ME. 2001. A primitive chondrichthyan braincase from the Early Devonian of South Africa. Journal of Vertebrate Paleontology 21 : 702 713.
146. Marshall CR. 1987. Lungfish: phylogeny and parsimony. Journal of Morphology Supplement 1 : 151 162.
147. Marshall JEA, Astin TR, Clack JA. 1999. East Greenland tetrapods are Devonian in age. Geology 27 : 637 640.
148. Miles RS. 1973. Relationships of acanthodians. In : Greenwood PH, Miles RS, Patterson C, eds. Interrelationships of fishes. London : Academic Press, 63 103.
149. Miles RS. 1975. The relationships of the Dipnoi. Colloque international. CNRS, Problèmes Actuels de Paléontolgie-Évolution des Vertébrés 218 : 133 148.
150. Miles RS. 1977. Dipnoan (lungfish) skulls and the relationships of the group: a study based on new species from the Devonian of Australia. Zoological Journal of the Linnean Society 61 : 1 328.
151. Millot J, Anthony J. 1958. Anatomie de Latimeria chalumnae. Tome II. Squelette, Muscles, et Formations du Soutien. Paris : Éditions du Centre National de la Recherche Scientifique.
152. Moy-Thomas JA. 1939. Palaeozoic fishes. London : Methuen Ltd.
153. Moy-Thomas JA, Miles RS. 1971. Palaeozoic fishes. London : Chapman & Hall.
154. Müller J. 1845. Über den Bau und die Grenzen der Ganoiden, and über das natürliche System der Fische. Abhandlungen der Königlichen Akademie der Wissenschaften zu Berlin 1844 : 117 216.
155. Natterer J. 1837. Lepidosiren paradoxa, eine neue Gattung aus der Familie der fischähnlichen Reptilien. Annalen des Wiener Museums der Naturgeschichte 2 : 165 170.
156. Nicholson J, Friend PF. 1976. Devonian sediments of East Greenland V. The central sequence, Kap Graah Group and Mount Celsius Supergroup. Meddelelser om Grønland 206 : 1 117.
157. Nielsen E. 1942. Studies on Triassic fishes I. Meddelelser om Grønland 138 : 1 394.
158. Nielsen E. 1949. Studies on Triassic fishes II. Meddelelser om Grønland 146 : 1 309.
159. Novitskaya LI, Krupina NI. 1985. On the ethmoid of Paleozoic dipnoans. Paleontological Journal 19 : 90 98.
160. Nylander JAA, Ronquist F, Huelsenbeck JP, Nieves-Aldrey JL. 2004. Bayesian phylogenetic analysis of combined data. Systematic Biology 53 : 47 67.
161. Olsen H, Larsen PH. 1993. Lithostratigraphy of the continental Devonian sediments in North-East Greenland. Grønlands Geologiske Undersøgelse 165 : 1 108.
162. Otto M, Bardenheuer P. 1996. Lungfish with dipterid tooth-plates in the Lower Devonian of Central Europe. Modern Geology 20 : 341 350.
163. Panchen AL, Smithsonian TR. 1987. Character diagnosis, fossils and the origin of tetrapods. Biological Reviews 62 : 341 438.
164. Patterson C. 1975. The braincase pholidophorid and leptolepid fishes, with a review of the actinopterygian braincase. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences 269 : 275 579.
165. Pearson DM, Westoll TS. 1979. The Devonian actinopterygian Cheirolepis Agassiz. Transactions of the Royal Society of Edinburgh: Earth Sciences 70 : 337 399.
166. Poplin C. 1981. Les homologies du pont prootique chez les osteichthyens. Cybium, 3e Série 5 : 3 17.
167. Pridmore PA, Campbell KSW, Barwick RE. 1994. Morphology and phylogenetic position of the holodipteran dipnoans of the Upper Devonian Gogo Formation of northwestern Australia. Philosophical Transactions of the Royal Society of London, Series B 344 : 105 164.
168. Rayner DH. 1951. On the cranial structure of an early palaeoniscid, Kentuckia, gen. nov. Transactions of the Royal Society of Edinburgh 62 : 53 83.
169. Reif WE. 1982. Evolution of the dermal skeleton and dentition in vertebrates. Evolutionary Biology 15 : 287 368.
170. Reisz RR, Smith MM. 2001. Lungfish dental pattern conserved for 360 Myr. Nature 411 : 548.
171. Rocco L, Costagliola D, Morescalchi MA, Stingo V. 2004. A molecular approach to systematics of Polypteriformes among Osteichthyes. Italian Journal of Zoology 71 : 347 351.
172. Romer AS. 1933. Vertebrate paleontology. Chicago : University of Chicago Press.
173. Romer AS. 1937. The braincase of the Carboniferous crossopterygian Megalichthys nitidus. Bulletin of the Museum of Comparative Zoology 82 : 1 73.
174. Romer AS. 1955. Herpetichthyes, Amphibioidei, Choanichthyes or Sarcopterygii? Nature 176 : 126.
175. Ronquist F, Huelsenbeck JP. 2003. MRBAYES 3: Bayesian phylogenetic inference under mixed models. Bioinformatics 19 : 1572 1574.
176. Rosen DA, Forey PL, Gardiner BG, Patterson C. 1981. Lungfishes, tetrapods, paleontology, and plesiomorphy. Bulletin of the American Museum of Natural History 167 : 163 264.
177. Ruta M, Coates MI, Quicke DLJ. 2003. Early tetrapod relationships revisited. Biological Reviews 78 : 251 345.
178. Säve-Söderbergh G. 1934. Further contributions to the Devonian stratigraphy of East Greenland. II. Meddelelser om Grønland 96 : 1 74.
179. Säve-Söderbergh G. 1936. On the morphology of the Triassic stegocephalians from Spitsbergen, and the interpretation of the endocranium in the Labyrithodontia. Kungliga Svenska Vetenskapsakademiens Handlingar, 3 Serien 16 : 1 181.
180. Säve-Söderbergh G. 1952. On the skull of Chirodipterus wildungensis Gross, an Upper Devonian dipnoan from Wildungen. Kungliga Svenska Ventenskapsakademiens Handlingar 3 : 5 28.
181. Schaeffer B. 1968. The origin and basic radiation of the Osteichthyes. In : Ørvig T, ed. Current problems of lower vertebrate phylogeny. Stockholm : Almqvist and Wiksell, 207 222.
182. Schaeffer B. 1981. The xenacanth shark neurocranium, with comments on elasmobranch monophyly. Bulletin of the American Museum of Natural History 69 : 3 66.
183. Schultze H-P. 1969. Griphognathus Gross, ein langschnauziger Dipnoer aus dem Oberdevon von Bergisch-Gladbach (Rheinisches Schiefergebirge) und von Lettland. Geologica et Palaeontologica 3 : 21 79.
184. Schultze H-P. 1973. Crossopterygier mit heterozerker schwanzflosse aus dem Oberdevon Kanadas, nebst einer beschreibung von Onychodontida-resten aus dem Mitteldevon Spaniens und aus dem Karbon der USA. Palaeontographica Abteilung A 143 : 188 208.
185. Schultze H-P. 1982. A dipterid dipnoan from the Middle Devonian of Michigan, U.S.A. Journal of Vertebrate Paleontology 2 : 155 162.
186. Schultze H-P. 1987. Dipnoans as sarcopterygians. Journal of Morphology Supplement 1 : 39 74.
187. Schultze H-P. 1991. A comparison of controversial hypotheses on the origin of tetrapods. In : Schultze H-P, Trueb L, eds. Origins of the higher groups of tetrapods: controversy and consensus. Ithaca, NY : Cornell University Press, 29 67.
188. Schultze H-P. 1992a. A new long-headed dipnoan (Osteichthyes) from the Middle Devonian of Iowa, USA. Journal of Vertebrate Paleontology 12 : 42 58.
189. Schultze H-P. 1992b. Early Devonian actinopterygians (Osteichthyes, Pisces) from Siberia. In : Mark-Kurik E, ed. Fossil fishes as living animals. Tallinn : Institute of Geology, 233 242.
190. Schultze H-P. 1992c. Fossilum Catalogus: Dipnoi. Amsterdam : Kugler Publications.
191. Schultze H-P. 1993a. Osteichthyes: Sarcopterygii. In : Benton MJ, ed. The fossil record 2. London : Chapman & Hall, 657 663.
192. Schultze H-P. 1993b. Patterns of diversity in the skull of jawed fishes. In : Hanken J, Hall BK, eds. The skull, vol. 2. patterns of structural and systematic diversity. Chicago : University of Chicago Press, 189 254.
193. Schultze H-P. 2001. Melanognathus: a primitive dipnoan from the Lower Devonian of the Canadian Arctic and the interrelationships of Devonian dipnoans. Journal of Vertebrate Paleontology 21 : 781 794.
194. Schultze H-P, Campbell KSW. 1987. Characterization of the Dipnoi, a monophyletic group. Journal of Morphology Supplement 1 : 25 37.
195. Schultze H-P, Chorn J. 1997. The Permo-Carboniferous genus Sagenodus and the beginning of modern lungfish. Contributions to Zoology 67 : 9 70.
196. Schultze H-P, Chorn J. 1998. Sarcopterygian and other fishes from the marine Upper Devonian of Colorado, U.S.A. Mitteilungen aus dem Museum für Naturkunde in Berlin, Geowissenschaftliche Reihe 1 : 53 72.
197. Schultze H-P, Cumbaa SL. 2001. Dialipina and the characters of basal actinopterygians. In : Ahlberg PE, ed. Major events in early vertebrate evolution. Palaeontology, phylogeny, genetics and development. Taylor & Francis, London, 316 332.
198. Schultze H-P, Marshall CR. 1993. Contrasting the use of functional complexes and isolated characters in lungfish evolution. Memoirs of the Association of Australasian Palaeontologists 15 : 211 224.
199. Smith MM. 1988. The dentition of Palaeozoic lungfishes: a consideration of the significance of teeth, denticles and tooth plates for dipnoan phylogeny. In : Russell DE, Santoro J-P, Sigogneau-Russell D, eds. Teeth revisited. Proceedings of the VIIth International Symposium on Dental Morphology. Mémoires du Muséum National d'Histoire Naturelle(Série C) 53 : 179 194.
200. Song C-Q, Chang M-M. 1991. Discovery of Chirodipterus (Diponi) from lower Upper Devonian of Hunan, South China. In : Chang M-M, Liu Y-H, Zhang G-R, eds. Early vertebrates and related problems of evolutionary biology. Beijing : Science Press, 465 476.
201. Sorenson MD. 1999. Treerot, Version 2. Boston : Boston University.
202. Spjeldnaes N. 1982. Palaeoecology of Ichthyostega and the origin of terrestrial vertebrates. In : Gallitelli EM, ed. Palaeontology, essential of historical geology (Paleontologia come scienzia geostorica). Modena : S.T.E.M. Mucci, 323 343.
203. Stensiö EA. 1937. On the Devonian coelacanthids of Germany with special reference to the dermal skeleton. Kungliga Svenska Vetenskapsakademiens Handlingar 16 : 1 56.
204. Strong EE, Lipscomb D. 2000. Character coding and inapplicable data. Cladistics 15 : 363 371.
205. Swofford DL. 2002. PAUP*. Phylogenetic analysis using parsimony (*and other methods), Version 4. Sunderland, MA : Sinauer Associates.
206. Takezaki N, Figueroa F, Zaleska-Rutczynska Z, Naoyuki T, Klein J. 2004. The phylogenetic relatopnship of tetrapod, coelacanth, and lungfish revealed by the sequences of forty-four nuclear genes. Molecular Biology and Evolution 21 : 1512 1524.
207. Thomson KS, Campbell KSW. 1971. The structure and relationships of the primitive Devonian lungfish - Dipnorhynchus sussmilchi (Etheridge). Peabody Museum of Natural History Bulletin 38 : 1 109.
208. Thorley JL, Page RDM. 2000. RadCon: phylogenetic tree comparison and consensus. Bioinformatics 16 : 486 487.
209. Traquair RH. 1881. On the cranial osteology of Rhizodopsis. Transactions of the Royal Society of Edinburgh 30 : 167 179.
210. Vorobyeva EI. 1977. Morfologiya i osobennosti evolyutsii kisteperykh ryb. Trudy Paleontologicheskogo Instituta 163 : 1 239. [in Russian].
211. Wang S, Drapala V, Barwick RE, Campbell KSW. 1993. The dipnoan species, Sorbitorhynchus deleaskitus, from the Lower Devonian of Guangxi, China. Philosophical Transactions of the Royal Society of London, Series B 340 : 1 24.
212. Westoll TS. 1949. On the evolution of the Dipnoi. In : Jepsen GL, Mayr E, Simpson GG, eds. Genetics, paleontology and evolution. Princeton, NJ : Princeton University Press, 121 184.
213. White EI. 1965. The head of Dipterus valenciennesi Sedgwick & Murchison. Bulletin of the British Museum (Natural History): Geology 11 : 1 45.
214. Wilkinson M. 1994. Common cladistic information and its consensus representation: reduced Adams and reduced cladistic consensus trees and profiles. Systematic Biology 43 : 343 368.
215. Wilkinson M. 1995. More on reduced consensus methods. Systematic Biology 44 : 435 439.
216. Woodward AS. 1891. Catalogue of the Fossil Fishes in the British Museum (Natural History), part II. London : Trustees of the British Museum (Natural History).
217. Woodward AS. 1898. Outlines of vertebrate palaeontology. Cambridge : Cambridge University Press.
218. Young GC. 1979. A new Early Devonian petalichthyid fish from the Taemas/Wee Jasper region of New South Wales. Alcheringa 2 : 103 116.
219. Young GC. 1980. A new Early Devonian placoderms from New South Wales, Australia, with a discussion of placoderm phylogeny. Palaeontographica Abteilung A 167 : 10 76.
220. Young GC, Long JA, Ritchie A. 1992. Crossopterygian fishes from the Devonian of Antarctica: systematics, relationships and biogeographic significance. Records of the Australian Museum Supplement 14 : 1 77.
221. Yu X. 1990. Cladistic analysis of sarcopterygian relationships, with a description of three new genera of Porolepiformes from the Lower Devonian of Yunnan, China. PhD thesis, Yale University.
222. Yu X. 1998. A new porolepiform-like fish, Psarolepis romeri, gen. et sp. nov. (Sarcopterygii, Osteichthyes) from the Lower Devonian of Yunnan, China. Journal of Vertebrate Paleontology 18 : 261 274.
223. Zangerl R. 1981. Chodrichthyes I: Paleozoic Elasmobranchii. In : Schultze H-P, ed. Handbook of paleoichthyology, Vol. 3A. Stuttgart : Gustav Fischer Verlag, 1 115.
224. Zardoya R, Meyer A. 1996. Evolutionary relationships of the coelacanth, lungfishes, and tetrapods based on 28S ribosomal RNA gene. Proceedings of the National Academy of Sciences of the United States of America 93 : 5449 5454.
225. Zhu M, Ahlberg PE. 2004. The origin of the internal nostril of tetrapods. Nature 432 : 94 97.
226. Zhu M, Schultze H-P. 1997. The oldest sarcopterygian fish. Lethaia 30 : 293 304.
227. Zhu M, Schultze H-P. 2001. Interrelationships of basal osteichthyans. In : Ahlberg PE, ed. Major events in early vertebrate evolution: palaeontology, phylogeny, genetics and development. London : Taylor & Francis, 289 314.
228. Zhu M, Yu X. 2002. A primitive fish close to the common ancestor of tetrapods and lungfish. Nature 418 : 767 770.
229. Zhu M, Yu X, Ahlberg PE. 2001. A primitive sarcopterygian fish with an eyestalk. Nature 410 : 81 84.
230. Zhu M, Yu X, Janvier P. 1999. A primitive fossil fish sheds light on the origins of bony fishes. Nature 397 : 607 610.
231. Zhu M, Yu X, Wang W, Zhao W, Jia L. 2006. A primitive fish provides key characteristics bearing on deep osteichthyan phylogeny. Nature 441 : 77 80.