A new trirachodontid cynodont from the lower levels of the Burgersdorp Formation (Lower Triassic) of the Beaufort Group, South Africa and the cladistic relationships of Gondwanan gomphodonts

Zoological Journal of the Linnean Society

Volumn 147, Issue 3, 2006, Pages 383-413

  Abdala, F ^a   Neveling, J ^b   Welman, J ^c  

^a UNIVERSITY OF THE WITWATERSRAND   (South Africa)

^b Western Cape Regional Office   (South Africa)

^c Free State University   (South Africa)

Author keywords

Early Middle Triassic; Gondwana; Phylogeny; Trirachodontidae

Indexed keywords

CYNOGNATHUS; METABOLUS; TRIRACHODON;

EID: 33745230782     PISSN: 00244082     EISSN: 10963642     Source Type: Journal    
DOI: 10.1111/j.1096-3642.2006.00224.x     Document Type: Article

References (128)

1. Abdala F Barberena MC Dornelles J. 2002. A new species of the traversodontid cynodont Exaeretodon from the Santa Maria Formation (Middle/Late Triassic) of southern Brazil. Journal of Vertebrate Paleontology 22 : 313 325.
2. Abdala F Giannini NP. 2000. Gomphodont cynodonts of the Chañares Formation: the analysis of an ontogenetic sequence. Journal of Vertebrate Paleontology 20 : 501 506.
3. Abdala F Hancox PJ Neveling J. 2005. Cynodonts from the uppermost Burgersdorp Formation, South Africa, and their bearing on the biostratigraphy and correlation of the Triassic Cynognathus Assemblage Zone. Journal of Vertebrate Paleontology 25 : 192 199.
4. Abdala F Ribeiro AM. 2003. A new traversodontid cynodont from the Santa Maria Formation (Ladinian-Carnian) of southern Brazil, with a phylogenetic analysis of Gondwanan traversodontids. Zoological Journal of the Linnean Society 139 : 529 545.
5. Abdala F Ribeiro AM Schultz CL. 2001. A rich cynodont fauna of Santa Cruz do Sul, Santa Maria Formation (Middle-Late Triassic), southern Brazil. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 2001 : 669 687.
6. Abdala F Teixeira AMS. 2004. A traversodontid cynodont of African affinity in the South American Triassic. Palaeontologia Africana 40 : 11 22.
7. Bandyopadhyay S Sengupta DP. 1999. Middle Triassic vertebrates of India. Journal of African Earth Sciences 29 : 233 241.
8. Barberena MC. 1974. Contribuição ao conhecimento dos cinodontes gonfodontes (Cynodontia, Tritylodontoidea) do Brasil. Tese para obtenção do título de Livre Docente inédita. Rio Grande do Sul : Ministério de Educação e Cultura, Universidade Federal do Rio Grande do Sul.
9. Barberena MC. 1981. Novos materiais de Traversodon stahleckeri da Formação Santa Maria (Triássico do Rio Grande do Sul). Pesquisas 14 : 149 162.
10. Battail B. 1983. La phylogénie des cynodonts gomphodontes. Acta Palaeontologica Polonica 28 : 19 30.
11. Battail B. 1991. Les cynodontes (Reptilia, Therapsida): une phylogénie. Bulletin du Muséum National d'Histore Naturelle de Paris 13 : 17 105.
12. Battail B. 2004. New data on Scalenodontoides macrodontes (Cynodontia: Traversodontidae). In : Ashwal LD, ed. Geosciences Africa 2004, Abstracts. Johannesburg : University of the Witwatersrand, 49 50.
13. Battail B Surkov MV. 2000. Mammal-like reptiles from Russia. In : Benton MJ Shishkin MA Unwin DM Kurochkin EN, eds. The age of dinosaurs in Russia and Mongolia. Cambridge : Cambridge University Press, 86 119.
14. Bonaparte JF. 1962. Descripción del cráneo y mandíbula de Exaeretodon frenguellii, Cabrera, y su comparación con Diademodontidae, Tritylodontidae y los cinodontes sudamericanos. Publicaciones del Museo Municipal de Ciencias Naturales y Tradición. Mar Del Plata 1 : 135 202.
15. Bonaparte JF. 1963. La Familia Traversodontidae (Terapsida-Cynodontia). Acta Geologica Lilloana 4 : 163 194.
16. Bonaparte JF. 1966. Una nueva 'fauna' triásica de Argentina (Therapsida: Cynodontia Dicynodontia). Consideraciones filogenéticas y paleobiogeográficas. Ameghiniana 4 : 243 296.
17. Bonaparte JF. 1969a. Dos nuevas 'faunas' de reptiles triásicos de Argentina. Gondwana stratigraphy. Mar del Plata : IUGS, 283 306.
18. Bonaparte JF. 1969b. Cynognathus minor n. sp. (Therapsida-Cynodontia). Nueva evidencia de vinculación faunística afrosudamericana a principios del Triásico. Gondwana stratigraphy. Mar del Plata : IUGS, 273 281.
19. Bonaparte JF Martinelli AG Schultz CL Rubert R. 2003. The sister group of mammals: small cynodonts from the Late Triassic of southern Brazil. Revista Brasileira de Paleontologia 5 : 5 27.
20. Brink AS. 1951. Some small cynodonts. South African Journal of Science 47 : 338 342.
21. Brink AS. 1955a. On the Cynognathidae. Palaeontologia Africana 3 : 47 55.
22. Brink AS. 1955b. A study on the skeleton of Diademodon. Palaeontologia Africana 3 : 3 39.
23. Brink AS. 1963. Two cynodonts from the Ntawere Formation in the Luangwa valley of northern Rhodesia. Palaeontologia Africana 8 : 77 96.
24. Brink AS Kitching JW. 1953. On some new Cynognathus Zone specimens. Palaeontologia Africana 1 : 29 48.
25. Broili F Schröder J. 1934. Zur Osteologie des Kopfes von Cynognathus. Sitzungsberichte der Bayerischen Akademie der Wissenchaften, Mathematisch-Naturwissenschaftliche, Abteilung 1934 : 95 128.
26. Broili F Schröder J. 1935a. Beobachtungen an Wirbeltieren der Karrooformation. X. Über die Bezahnung von Trirachodon Seeley. Sitzungsberichte der Bayerischen Akademie der Wissenchaften, Mathematisch-Naturwissenschaftliche, Abteilung 1935 : 189 198.
27. Broili F Schröder J. 1935b. Beobachtungen an Wirbeltieren der Karrooformation. XI. Über den Schädel von Cynidiognathus Haughton. Sitzungsberichte der Bayerischen Akademie der Wissenchaften, Mathematisch-Naturwissenschaftliche, Abteilung 1935 : 199 222.
28. Broili F Schröder J. 1935c. Beobachtungen an Wirbeltieren der Karrooformation. IX Über den Schädel von Gomphognathus Seeley. Sitzungsberichte der bayerischen Akademie der Wissenchaften, mathematisch-naturwissenschaftliche, Abteilung 1935 : 115 182.
29. Broom R. 1903. On the axis, atlas, and proatlas in the higher theriodonts. Proceedings of the Zoological Society of London b 1903 : 177 180.
30. Broom R. 1904. On the theriodonts in the Albany Museum. Records of the Albany Museum 1 : 82 87.
31. Broom R. 1905a. Preliminary notice of some new fossil reptiles collected by Mr. Alfred Brown at Aliwal North, S. Africa. Records of the Albany Museum 1 : 269 275.
32. Broom R. 1905b. On some points in the anatomy of the theriodont reptile Diademodon. Proceedings of the Zoological Society of London b 1905 : 96 102.
33. Broom R. 1907. On the geological horizons of the vertebrate genera of the Karroo Formation. Records of the Albany Museum 2 : 156 163.
34. Broom R. 1911. On the structure of the skull in cynodont reptiles. Proceedings of the Zoological Society of London b 1911 : 893 925.
35. Broom R. 1915a. On some new carnivorous therapsids in the collection of the British Museum. Proceedings of the Zoological Society of London b 1915 : 163 173.
36. Broom R. 1915b. Catalogue of types and figures specimens of fossil vertebrates in the American Museum of Natural History. II. Permian, Triassic and Jurassic reptiles of South Africa. Bulletin of the American Museum of Natural History 25 : 105 164.
37. Broom R. 1919. On the genus Gomphognathus and its allies. Records of the Albany Museum 3 : 223 232.
38. Broom R. 1932. The mammal-like reptiles of South Africa and the origin of mammals. London : H. F. & G. Witherby.
39. Chatterjee S. 1982. A new cynodont reptile from the Triassic of India. Journal of Paleontology 56 : 203 214.
40. Coddington J Scharff N. 1994. Problems with zero-length branches. Cladistics 10 : 415 423.
41. Crompton AW. 1955. On some Triassic cynodonts from Tanganyika. Proceedings of the Zoological Society of London 125 : 617 669.
42. Crompton AW. 1963. On the lower jaw of Diarthrognathus and the origin of the mammalian lower jaw. Proceedings of the Zoological Society of London 140 : 697 749.
43. Crompton AW. 1972a. Postcanine occlusion in Cynodonts and Tritylodontids. Bulletin of the British Museum of Natural History, Geology 21 : 27 71.
44. Crompton AW. 1972b. The evolution of the jaw articulation in cynodonts. In : Joysey KA Kemp TS, eds. Studies in vertebrate evolution. Edinburgh : Oliver & Boyd, 231 251.
45. Crompton AW Ellenberger F. 1957. On a new cynodont from the Molteno Beds and the origin of the tritylodontids. Annals of the South African Museum 44 : 1 13.
46. Damiani R. 2004. Temnospondyls from the Beaufort Group (Karoo Basin) of South Africa and their biostratigraphy. Gondwana Research 7 : 165 173.
47. Damiani R Hancox PJ. 2003. New mastodonsaurid temnospondyls from the Cynognathus Assemblage Zone (Upper Beaufort Group; Karoo Basin) of South Africa. Journal of Vertebrate Palaeontology 23 : 54 66.
48. Flynn JJ Parrish JM Rakotosamimanana B Ranivoharimanana L Simpson WF Wyss AR. 2000. New traversodontids (Synapsida: Eucynodontia) from the Triassic of Madagascar. Journal of Vertebrate Paleontology 20 : 422 427.
49. Flynn JJ Parrish JM Rakotosamimanana B Simpson WF Whatley RL Wyss AR. 1999. A Triassic fauna from Madagascar, including early dinosaurs. Science 286 : 763 765.
50. Fourie S. 1974. The cranial morphology of Thrinaxodon liorhinus Seeley. Annals of the South African Museum 65 : 337 400.
51. Goloboff PA. 1993. Estimating character weights during tree search. Cladistics 9 : 83 91.
52. Goloboff PA Farris JS Nixon KC. 2003. T.N.T. tree analysis using new technology. http://www.zmuc.dk/public/phylogeny.
53. Goñi RG. 1986. Reemplazo de dientes postcaninos en Andescynodon mendozensis Bonaparte (Cynodontia, Traversodontidae). Actas del IV Congreso Argentino de Paleontología y Bioestratigrafia 2 : 7 14.
54. Goñi RG Goin FJ. 1988. Morfología dentaria y biomecánica maticatoria de los cinodontes (Reptilia-Therapsida) del Triásico argentino. I. Andescynodon mendozensis Bonaparte (Cynodontia, Traversodontidae). Ameghiniana 25 : 139 148.
55. Gow CE. 1986. The side wall of the braincase in cynodont therapsids, and a note on the homology of the mammalian promontorium. South African Journal of Zoology 21 : 136 148.
56. Gow CE Hancox PJ. 1993. First complete skull of the Late Triassic Scalenodontoides (Reptilia, Cynodontia) from southern Africa. New Mexico Museum of Natural History & Science Bulletin 3 : 161 168.
57. Gradstein FM Ogg JG. 2004. Geologic time scale 2004 - why, how, and where next! Lethaia 37 : 175 181.
58. Grine FE. 1977. Postcanine tooth function and jaw movements in the gomphodont cynodont Diademodon (Reptilia; Therapsida). Palaeontologia Africana 20 : 123 135.
59. Groenewald GH Welman J MacEachern JA. 2001. Vertebrate burrow complexes from the Early Triassic Cynognathus Zone (Driekoppen Formation, Beaufort Group) of the Karoo Basin, South Africa. Palaios 16 : 148 160.
60. Hammer WR. 1995. New therapsids from the Upper Fremouw Formation (Triassic) of Antarctica. Journal of Vertebrate Paleontology 15 : 105 112.
61. Hancox PJ. 2000. The Continental Triassic of South Africa. Zentralblatt für Geologie und Paläontologie, Teil I, Heft 11-12 1998 : 1285 1324.
62. Hancox PJ Rubidge BS. 1994. A new dicynodont therapsid from South Africa: implications for the biostratigraphy of the Upper Beaufort (Cynognathus Assemblage Zone). South African Journal of Science 90 : 98 99.
63. Hancox PJ Rubidge BS. 1996. The first specimen of the Mid-Triassic dicynodont Angonisaurus from the Karoo of South Africa: implications for the dating and biostratigraphy of the Upper Beaufort Group. South African Journal of Science 92 : 391 392.
64. Hancox PJ Rubidge BS. 2001. Breakthroughs in the biodiversity, biogeography, biostratigraphy and basin analysis of the Beaufort Group. Journal of African Earth Sciences 33 : 563 577.
65. Hancox PJ Shishkin MA Rubidge BS Kitching JW. 1995. A threefold subdivision of the Cynognathus Assemblage Zone (Beaufort Group, South Africa) and its palaeogeographic implications. South African Journal of Science 91 : 143 144.
66. Hopson JA. 1971. Postcanine replacement in the gomphodont cynodont Diademodon. Zoological Journal of the Linnean Society 50 : 1 21.
67. Hopson JA. 1984. Late Triassic traversodont cynodonts from Nova Scotia and southern Africa. Palaeontologia Africana 25 : 181 201.
68. Hopson JA. 1985. Morphology and relationships of Gomphodontosuchus brasiliensis von Huene (Synapsida, Cynodontia, Tritylodontoidea) from the Triassic of Brazil. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 1985 : 285 299.
69. Hopson JA. 1991. Systematics of the nonmammalian Synapsida and implications for patterns of evolution in synapsids. In : Schultze H-D Trueb L, eds. Origin of the higher groups of tetrapods: controversy and consensus. Ithaca : Comstock Publishing Associates, Cornell University Press, 635 693.
70. Hopson JA Barghusen HR. 1986. An analysis of therapsid relationships. In : Hotton N III. MacLean PD Roth JJ Roth EC, eds. The ecology and biology of mammal-like reptiles. Washington : Smithsonian Institution Press, 83 106.
71. Hopson JA Kitching JW. 1972. A revised classification of cynodonts (Reptilia, Therapsida). Palaeontologia Africana 14 : 71 85.
72. Hopson JA Kitching JW. 2001. A probainognathian cynodont from South Africa and the phylogeny of nonmammalian cynodonts. Bulletin of the Museum of Comparative Zoology 156 : 5 35.
73. von Huene F. 1928. Ein Cynodontier aus des Trias Brasiliens. Centralblatt für Mineralogie, Geologie und Paläontologie 1928B : 251 270.
74. von Huene F. 1936. Die fossilen Reptilien der südamerikanischen Gondwanalandes. Ergebnisse der Sauriergrabungen in Südbrasilien 1928-29. Lieferung 2. Tübingen : Verlag Franz F. Heine, 93 159.
75. Kemp TS. 1979. The primitive cynodont Procynosuchus: functional anatomy of the skull and relationships. Philosophical Transactions of the Royal Society of London B 285 : 73 122.
76. Kemp TS. 1980. Aspects of the structure and functional anatomy of the Middle Triassic cynodont Luangwa. Journal of Zoology 191 : 193 239.
77. Kemp TS. 1982. Mammal-like reptiles and the origin of mammals. London : Academic Press.
78. Kemp TS. 1983. The relationships of mammals. Zoological Journal of the Linnean Society 77 : 353 384.
79. Keyser AW. 1973. A new vertebrate fauna from South West Africa. Palaeontologia Africana 16 : 1 15.
80. Keyser AW Smith RMH. 1977-78. Vertebrate biozonation of the Beaufort Group with special reference to the western Karoo Basin. Annals of the Geological Survey 12 : 1 35.
81. Kitching JW. 1977. The distribution of the Karoo vertebrate fauna. Memoir 1. Johannesburg : Bernard Price Institute for Palaeontological Research, University of the Witwatersrand.
82. Kitching JW. 1995. Biostratigraphy of the Cynognathus Assemblage Zone. In : Rubidge B, ed. Biostratigraphy of the Beaufort Group (Karoo Supergroup). Biostratigraphic Series 1. Pretoria : South African Committee for Stratigraphy, 40 45.
83. von Koenigswald W Sander PM. 1997. Glossary of terms used for enamel microstructures. In : von Koenigswald W Sander PM, eds. Tooth enamel microstructure. Rotterdam : Balkema, 267 280.
84. Kokogian DA Spalletti L Morel E Artabe A Martínez RN Alcober OA Milana JP Zavattieri AM Papú OH. 1999. Los depósitos continentales Triásicos. Geología Argentina. Anales del Instituto de Geología y Recursos Minerales 29 : 377 398.
85. Lucas SG Estep JW Heckert AB Hunt AP. 1999. Cynodont teeth from the Upper Triassic of New Mexico, USA. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 1999 : 331 344.
86. Lucas SG Hancox PJ. 2001. Tetrapod-based correlation of the nonmarine Upper Triassic of Southern Africa. Albertiana 25 : 5 9.
87. Lucas SG Luo Z. 1993. Adelobasileus from the Upper Triassic of west Texas: the oldest mammal. Journal of Vertebrate Paleontology 13 : 309 334.
88. Luo Z. 1994. Sister-group relationships of mammals and transformations of diagnostic mammalian characters. In : Fraser NC Sues H-D, eds. The shadow of the dinosaurs - early Mesozoic tetrapods. Cambridge : Cambridge University Press, 98 128.
89. Luo Z Crompton AW. 1994. Transformation of the quadrate (incus) through the transition from non-mammalian cynodonts to mammals. Journal of Vertebrate Paleontology 14 : 341 374.
90. Martínez RN May CL Forster CA. 1996. A new carnivorous cynodont from the Ischigualasto Formation (Late Triassic, Argentina), with comments on eucynodont phylogeny. Journal of Vertebrate Paleontology 16 : 271 284.
91. Morel EM Artabe AE Spalletti LA. 2003. Triassic floras of Argentina: biostratigraphy, floristic events and comparison with other areas of Gondwana and Laurasia. Alcheringa 27 : 231 243.
92. Neveling J. 2002. Biostratigraphic and sedimentological investigation of the contact between the Lystrosaurus and Cynognathus Assemblage Zones (Beaufort Group: Karoo Supergroup). PhD Thesis, University of the Witwatersrand, Johannesburg.
93. Neveling J. 2004. Stratigraphic and sedimentological investigation of the contact between the Lystrosaurus and Cynognathus Assemblage Zones (Beaufort Group: Karoo Supergroup). Bulletin of the Council for Geosciences, South Africa, 137 : 1 165.
94. Neveling J Rubidge BS Hancox PJ. 1999. A lower Cynognathus Assemblage Zone fossil from the Katberg Formation (Beaufort Group, South Africa). South African Journal of Science 95 : 555 556.
95. Parrington FR. 1946. On the cranial anatomy of cynodonts. Proceedings of the Zoological Society of London 116 : 181 197.
96. Rogers RR Arcucci AB Abdala F Sereno PC Forster CA May CL. 2001. Paleoenvironment and taphonomy of the Chañares Formation tetrapod assemblage (Middle Triassic), north-western Argentina: spectacular preservation in volcanogenic concretions. Palaios 16 : 461 481.
97. 39Ar dating of dinosaurs origins. Science 260 : 794 797.
98. Romer AS. 1967. The Chañares (Argentina) Triassic reptile fauna. III. Two new gomphodonts, Massetognathus pascuali and Massetognathus teruggii. Breviora 264 : 1 25.
99. Romer AS. 1972. The Chañares (Argentina) Triassic reptile fauna. XVII. The Chañares gomphodonts. Breviora 396 : 1 9.
100. Rougier GW Wible JR Hopson JA. 1992. Reconstruction of the cranial vessels in the Early Cretaceous mammal Vincelestes neuquenianus: implications for the evolution of the mammalian cranial vascular system. Journal of Vertebrate Paleontology 12 : 188 216.
101. Rowe T. 1988. Definition, diagnosis and origin of mammalia. Journal of Vertebrate Paleontology 8 : 241 264.
102. Rowe T. 1993. Phylogenetic systematics and the early history of mammals. In : Szalay FS Novacek MJ McKenna MC, eds. Mammal phylogeny. Mesozoic differentiation, multituberculates, monotremes, early therians, and marsupials. New York : Springer, 129 145.
103. Rubidge BS Johnson MR Kitching JW Smith RMH Keyser AW Groenewald GH. 1995. An introduction to the biozonation of the Beaufort Group. In : Rubidge B, ed. Biostratigraphy of the Beaufort Group (Karoo Supergroup). Biostratigraphic Series 1. Pretoria : South African Committee for Stratigraphy, 1 2.
104. Rubidge BS Sidor CA. 2001. Evolutionary patterns among Permo-Triassic therapsids. Annual Review of Ecology and Systematics 32 : 449 480.
105. Sander PM. 1999. The microstructure of reptilian tooth enamel: terminology, function and phylogeny. Münchner Geowissenschaftliche Abhandlungen 38 : 1 102.
106. Seeley HG. 1892. Researches on the structure, organization, and classification of the Fossil Reptilia. VII. Further observations on Pareiasaurus. Philosophical Transactions of the Royal Society of London 183 : 311 370.
107. Seeley HG. 1894. Researches on the structure, organization, and classification of the Fossil Reptilia. Part IX, Section 3. On Diademodon. Philosophical Transactions of the Royal Society of London 185 : 1029 1041.
108. Seeley HG. 1895a. Researches on the structure, organization, and classification of the Fossil Reptilia. Part IX, Section 4. On the Gomphodontia. Philosophical Transactions of the Royal Society of London 186 : 1 57.
109. Seeley HG. 1895b. Researches on the structure, organization, and classification of the Fossil Reptilia. Part IX, Section 5. On the skeleton in new Cynodontia from the Karroo rocks. Philosophical Transactions of the Royal Society of London 186 : 59 148.
110. Seeley HG. 1908. Additional evidence as to the dentition and structure of the skull in the South African fossil reptile genus Diademodon. Proceedings of the Zoological Society of London b 1908 : 611 617.
111. Shishkin MA Rubidge BS Hancox PJ. 1995. Vertebrate biozonation of the Upper Beaufort Series of South Africa - A new look on correlation of the Triassic biotic events in Euramerica and southern Gondwana. In : Sun A Wang Y, eds. Sixth symposium on Mesozoic terrestrial ecosystems and biota, short papers. Beijing : China Ocean Press, 39 41.
112. Sidor CA Smith RMH. 2004. A new galesaurid (Therapsida: Cynodontia) from the Lower Triassic of South Africa. Palaeontology 47 : 535 556.
113. Smith MM. 1989. Distribution and variation in enamel structure in the oral teeth of sarcopterygians: its significance for the evolution of a protoprismatic enamel. Historical Biology 3 : 97 126.
114. Smith RMH Swart R. 2002. Changing fluvial environments and vertebrate taphonomy in response to climatic drying in a Mid-Triassic rift valley fill: the Omingonde Formation (Karoo Supergroup) of central Namibia. Palaios 17 : 249 267.
115. Sues H-D. 1985. First record of the tritylodontid Oligokyphus (Synapsida) from the Lower Jurassic of western North America. Journal of Vertebrate Paleontology 5 : 328 335.
116. Sues H-D. 1986. The skull and dentition of two tritylodontid synapsids from the Lower Jurassic of western North America. Bulletin of the Museum of Comparative Zoology 151 : 217 268.
117. Sues H-D Hopson JA Shubin NH. 1992. Affinities of ?Scalenodontoides plemmyridon Hopson, 1984 (Synapsida: Cynodontia) from the Upper Triassic of Nova Scotia. Journal of Vertebrate Paleontology 12 : 168 171.
118. Sun A. 1984. Skull morphology of the tritylodont genus Bienotheroides of Sichuan. Scientia Sinica 27 : 970 984.
119. Sun A. 1988. Additional study on Sinognathus gracilis (Cynodontia; Reptilia). Vertebrata Palasiatica 26 : 173 180.
120. Tatarinov LP. 1973. Cynodonts of Gondwanan habit in the Middle Triassic of the USSR. Paleontological Journal 1973 : 200 205.
121. Tatarinov LP. 1988. On the morphology and systematic position of the gomphodont cynodont Antecosuchus ochevi. Paleontological Journal 1988 : 82 90.
122. Tatarinov LP. 2002. Gomphodont cynodonts (Reptilia, Theriodontia) from the Middle Triassic of the Orenburg Region. Paleontological Journal 36 : 176 179.
123. Watson DMS. 1911. The skull of Diademodon, with notes on those of some other cynodonts. Annals and Magazine of Natural History 8 : 293 330.
124. Watson DMS. 1914. The zones of the Beaufort beds of the Karroo System in South Africa. Geological Magazine, New Series 6 : 203 208.
125. Welman J Groenewald GH Kitching JW. 1991. Confirmation of the occurrence of Cynognathus Zone (Kannemeyeria-Diademodon Assemblage-zone) deposits (uppermost Beaufort Group) in the northeastern Orange Free State, South Africa. South African Journal of Geology 94 : 245 248.
126. Wible JR. 1991. Origin of Mammalia: the craniodental evidence reexamined. Journal of Vertebrate Paleontology 11 : 1 28.
127. Young C-C. 1959. Note on the first cynodont from the Sinokannemeyeria faunas in Shansi, China. Vertebrata Palasiatica 3 : 124 131.
128. Zavattieri AM Arcucci AB. 2003. Edad y posición estratigráfica de los tetrápodos de la localidad Cerro Bayo de Potrerillos (Triásico), Mendoza, Argentina. Ameghiniana 40 : 75R.