A revision of the goby genus Gnatholepis Bleeker (Teleostei, Gobiidae, Gobionellinae), with description of a new species

Zootaxa

Volumn , Issue 3529, 2012, Pages 1-69

  Larson, Helen K ^a   Buckle, Duncan J ^b  

^a Museum and Art Gallery Northern Territory   (Australia)

^b CHARLES DARWIN UNIVERSITY   (Australia)

Author keywords

Atlantic; Gnatholepis; Gobiidae; Gobionellinae; Indo Pacific; New species

Indexed keywords

EID: 84868135069     PISSN: 11755326     EISSN: 11755334     Source Type: Journal    
DOI: 10.11646/zootaxa.3529.1.1     Document Type: Article

References (146)

1. Allen, G.R. (1997) Marine Fishes of Tropical Australia and South-East Asia. Western Australian Museum, Perth, 292 pp.
2. Allen, G.R. & Adrim, M. (2003) Coral reef fishes of Indonesia. Zoological Studies, 42(1), 1-72. (Pubitemid 36646135)
3. Allen, G.R. & Smith-Vaniz, W.F. (1994) Fishes of the Cocos (Keeling) Islands. Atoll Research Bulletin, 412, 1-21.
4. Allen, G R. & Swainston, R. (1988) The Marine Fishes of North-Western Australia. A Field Guide for Anglers And Divers. Western Australian Museum, Perth, 201 pp.
5. Anderson, M.J., Gorley, R.N. & Clarke, K.R. (2008) PERMANOVA+ for PRIMER: Guide to software and statistical methods. Primer-E, Plymouth, UK.
6. Anderson, M.J. & Robinson, J. (2003) Generalized discriminant analysis based on distances. Australian & New Zealand Journal of Statistics, 45(3), 301-318. (Pubitemid 37023230)
7. Anderson, M.J. & Willis, T.J. (2003) Canonical analysis of principal coordinates: a useful method of constrained ordination for ecology. Ecology, 84(2), 511-525. (Pubitemid 36893087)
8. Anderson, R.C. (2005) Reef Fishes of the Maldives. Manta Marine, Malé, 130 pp.
9. Aoyagi, H. (1957) General Notes on the Freshwater Fishes of the Japanese Archipelago. Taishukan [in Japanese].
10. Araújo, R. & Freitas, M. (2002) First record of the goldspot goby Gnatholepis thompsoni Jordan, 1904 (Pisces: Gobiidae) in Madeira Island (NE Atlantic Ocean). Bocagiana (Funchal), 209, 1-4.
11. Birdsong, R.S., Murdy, E.O. & Pezold, F.L. (1988) A study of the vertebral column and median fin osteology in gobioid fishes with comments on gobioid relationships. Bulletin of Marine Science, 42(2), 174-214.
12. Bleeker, P. (1851) Over eenige nieuwe soorten van Blennioïden en Gobioïden van den Indischen Archipel. Natuurkundig Tijdschrift voor Nederlandsch-Indie, 1, 236-258.
13. Bleeker, P. (1853) Diagnostische beschrijvingen van nieuwe of weinig bekende vischsoorten van Sumatra. Tiental V-X. Natuurkundig Tijdschrift voor Nederlansch-Indië, 4, 243-302.
14. Bleeker, P. (1854) Bijdrage tot de kennis der ichthyologische fauna van de Kokos-eilanden. Natuurkundig Tijdschrift voor Nederlansch-Indië, 7, 37-48.
15. Bleeker, P. (1874) Esquisse d'un système naturel des Gobioïdes. Archives Néerlandaises des Sciences exactes et naturelles, 9, 289-331.
16. Bleeker, P. (1983) Atlas Ichthyologique des Indes Orientales Néêrlandaises, par M.- P. Bleeker. (Plates originally planned for planned tomes XI-XIV published here for the first time). Smithsonian Institution Press: Washington, 186 pp.
17. Blyth, E. (1860) Report on some fishes received chiefly from the Sitang River and its tributary streams, Tenasserim Provinces. Journal of the Asiatic Society of Bengal, 29(2), 138-174.
18. Böhlke, J.E. (1953) A catalogue of the type specimens of Recent fishes in the Natural History Museum of Stanford University. Stanford Ichthyological Bulletin, 5, 1-168.
19. Böhlke, E.B. (1984) Catalog of type specimens in the ichthyological collection of the Academy of Natural Sciences of Philadelphia. Academy of Natural Sciences of Philadelphia Special Publication, 14, 1-246.
20. Brito, A. & Miller, P.J. (2001) Gobiid fishes from the Cape Verde Islands, including two new species of Gobius. Journal of Natural History, 35(2), 253-277. (Pubitemid 32106432)
21. Coene, H. & Ollevier, F. (2000) Resource partitioning in a tropical goby assemblage. Abstract. Meeting on Mangrove Macrobenthos: Biology, Ecology and Exploitation, Mombasa, Kenya, 7-11 September 2000.
22. Cole, K. S. (1990) Patterns of gonad structure in hermaphroditic gobies (Teleostei: Gobiidae). Environmental Biology of Fishes, 28, 125-142.
23. Collette, B.B., Williams, J.T., Thacker, C.E. & Smith, M.L. (2003) Shore fishes of Navassa Island, West Indies: a case study on the need for rotenone sampling in reef fish biodiversity studies. Aqua, Journal of Ichthyology and Aquatic Biology, 6(3), 89-131.
24. Day, F. (1870) Remarks on some of the fishes in the Calcutta Museum. Part I. Proceedings of the Zoological Society of London, 1869 (Part 3), 511-527.
25. Debelius, H. (1997) Mediterranean and Atlantic Fish Guide. IKAN-Unterwasser Archiv, Frankfurt, 305 pp.
26. Debelius, H. (1998) Red Sea Reef Guide. IKAN-Unterwasser Archiv, Frankfurt, 321 pp.
27. De Vis, C.W. (1884) New Australian fishes in the Queensland Museum. No.4. Proceedings of the Linnean Society of New South Wales, 9(3), 685-698.
28. DiSalvo, L.H., Randall, J.E. & Cea, A. (1988) Ecological reconnaissance of the Easter Island sublittoral marine environment. National Geographic Research, 4(4), 451-473.
29. Dor, M. (1984) CLOFRES. Checklist of the Fishes of the Red Sea. Israel Academy of Sciences and Humanities, Jerusalem, 437
30. Edwards, A.J. & Glass, C.W. (1987) The fishes of Saint Helena Island, South Atlantic Ocean. I. The shore fishes. Journal of Natural History, 21(3), 617-686.
31. Francis, M.P. (1993) Checklist of the coastal fishes of Lord Howe, Norfolk and Kermadec Islands, southwest Pacific Ocean. Pacific Science, 47(2), 136-170. (Pubitemid 24412451)
32. Fricke, R. (1999) Fishes of the Mascarene Islands (Réunion, Mauritius, Rodriguez). An Annotated Checklist, with Descriptions of New Species. Theses Zoologicae Volume 31. Koeltz Scientific Books, Koenigstein, 759 pp.
33. Fricke, R., Mulochau, T., Durville, P., Chabanet, P., Tessier, E. & Letourneur, Y. (2009) Annotated checklist of the fish species (Pisces) of La Réunion, including a Red List of threatened and declining species. Stuttgarter Beiträge zur Naturkunde A, Neue Serie 2, 1-168.
34. Galzin, R., Lecchini, D., Williams, J.T., Planes, S. & Menou, J.-L. (2002) Diversité de l'ichtyofaune corallienne à Rapa (Polynésie française). Cybium, 30(3), 221-234. (Pubitemid 46167076)
35. Gasparini, J. L. & Floeter, S. R. (2001) The shore fishes of Trindade Island, western South Atlantic. Journal of Natural History, 35(11), 1639-1656. (Pubitemid 33037620)
36. Gill, A.C. & Reader, S. (1992) Fishes. Pp. 90-93, 193-118.
37. In: Reef biology: a survey of Middleton & Elizabeth Reefs, South Pacific. Kowari, 3, 1-230.
38. Goren, M. (1979) The Gobiinae of the Red Sea (Pisces: Gobiidae). Senckenbergiana Biologica, 60(1-2), 13-64.
39. Goren, M. (1981) Three new species and three new records of gobies from New Caledonia. Cybium, 5(3), 93-101.
40. Goren, M. & Dor, M. (1994) An Updated Checklist of the Fishes of the Red Sea. CLOFRES II. Israel Academy of Sciences and Humanities, and Interuniversity Institute for Marine Sciences, Elat, 120 pp.
41. Greenfield, D.W. & Randall, J.E. (2004) The marine gobies of the Hawaiian Islands. Proceedings of the California Academy of Sciences, 55(27), 498-549.
42. Günther, A. (1861) Catalogue of the acanthopterygian fishes in the collection of the British Museum, Vol. III, British Museum, London, 153 pp.
43. Harrison, I.J. (1989) Specialization of the gobioid palatopterygoquadrate complex and its relevance to gobioid systematics. Journal of Natural History, 23, 325-353.
44. Hayashi, M. & Shiratori, T. (2003) Gobies of Japanese waters. TBS Encyclopaedia Britannica/Hankyu, Tokyo, 223 pp.
45. Heemstra, E., Heemstra, P., Smale, M., Hooper, T. & Pelicier, D. (2004) Preliminary checklist of coastal fishes from the Mauritian island of Rodrigues. Journal of Natural History, 38, 3315-3344. (Pubitemid 39342486)
46. Herre, A.W.C.T. (1927) Gobies of the Philippines and the China Sea. Monographs of the Philippine Bureau of Science, Manila, Philippine Islands, 23, 1-352.
47. Herre, A.W.C.T. (1931) A check list of the fishes recorded from the New Hebrides. Journal of the Pan-Pacific Research Institution, 6(4), 11-14.
48. Herre, A.W.C.T. (1935) New fishes obtained by the Crane Pacific expedition. Field Museum of Natural History Publications in Zoology Series, 18(12), 383-438.
49. Herre, A.W.C.T. (1936) Fishes of the Crane Pacific Expedition. Field Museum of Natural History Publications in Zoology Series, 21, 1-472.
50. Herre, A.W.C.T. (1953) A new species of Gnatholepis with a key to the tropical Pacific species. Philippine Journal of Science, 82(2), 193-197.
51. Hoese, D.F. (1986) Family No. 240: Gobiidae. In: Smith, M.M. & Heemstra, P.C. (Eds), Smith's Sea Fishes. Springer-Verlag, New York, pp. 774-807.
52. Hoese, D.F. & Allen, G.R. (1977) Signigobius biocellatus, a new genus and species of sand-dwelling coral reef gobiid fish from the western tropical Pacific. Japanese Journal of Ichthyology, 23(4), 199-207.
53. Hoese, D.F. & Larson, H.K. (2006) Gobiidae. Gobies. In: Hoese, D.F., Bray, D.J., Paxton, J.R. & Allen, G.R. (Eds), Zoological Catalogue of Australia. Volume 35. Fishes. Part 3(of 3). ABRS and CSIRO Publishing, Canberra, pp 1612-1697.
54. Hoese, D.F. & Winterbottom, R. (1979) A new species of Lioteres (Pisces, Gobiidae) from Kwazulu, with a revised checklist of South African gobies and comments on the generic relationships and endemism of western Indian Ocean gobioids. Royal Ontario Museum, Life Sciences Occasional Paper, 31, 1-13.
55. Hubbs, C.L. & Lagler, K.F. (1970) Fishes of the Great Lakes Region. University of Michigan Press, Ann Arbor, 213 pp.
56. Hutchins, B. (2001) Checklist of the fishes of Western Australia. Records of the Western Australian Museum, Supplement, 63, 9-50.
57. Ibarra, M. & Stewart, D.J. (1987) Catalogue of type specimens of Recent fishes in Field Museum of Natural History. Fieldiana Zoology (New Series), 35, 1-112.
58. Jatzow, R. & Lenz, H. (1898) Fische von Ost-Afrika, Madagaskar und Aldabra. Abhandlungen Senckenbergischen Naturforschenden Gesellschaft Frankfurt, 21, 497-531.
59. Jordan, D.S. (1904) Notes on the fishes collected in the Tortugas Archipelago. Bulletin of the United States Fisheries Commission, 22 [1902], 539-544.
60. Jordan, D.S. & Evermann, B.W. (1903) Descriptions of new genera and species of fishes from the Hawaiian Islands. Bulletin of the United States Fisheries Commission, 22 [1902], 161-208.
61. Jordan, D.S. & Evermann, W.B. (1905) The aquatic resources of the Hawaiian Islands. Part II. The shore fishes. Bulletin of the United States Fisheries Commission, 23 [1903], 1-574.
62. Jordan, D.S. & Evermann, W.B. (1926) A check list of the fishes of Hawaii. Journal of the Pan-Pacific Research Institution, 1(1), 1-15.
63. Kimura, S., Satapoomin, U. & Matsuura, K. (Eds) (2009) Fishes of Andaman Sea. West Coast of Southern Thailand. National Museum of Nature and Science, Tokyo, 346 pp.
64. Klunzinger, C.B. (1871) Synopsis der Fische des Rothen Meeres, II. Verhandlunger der Kaiserlich-Königlichen Zoologisch-botanischen Gesellschaft in Wien, 21, 441-668.
65. Koumans, F.P. (1931) A preliminary revision of the genera of the gobioid fishes with united ventral fins. Proefschrift Drukkerij Imperator N.V. Lisse, 1-174.
66. Koumans, F.P. (1940) Results of a reexamination of types and specimens of gobioid fishes, with notes on the fishfauna of the surroundings of Batavia. Zoologische Mededeelingen, 22, 121-210.
67. Kuiter, R.H. (1993) Coastal fishes of South-Eastern Australia. Crawford House Press, Bathurst, 437 pp.
68. Kuiter, R.H. & Tonozuka, T. (2001) Pictorial Guide to Indonesian Reef Fishes Part 3 Jawfishes - sunfishes. Zoonetics, Melbourne, pp 623-859.
69. Kulbicki, M. & Williams, J.T. (1997) Checklist of the shorefishes of Ouvéa Atoll, New Caledonia. Atoll Research Bulletin, 444, 1-26.
70. Laboute, P. & Grandperrin, R. (2000) Poissons de Nouvelle- Calédonie. Editions Catherine Ledru, Nouméa, 520 pp.
71. Larson, H.K. (1999) A review of the mangrove goby genus Hemigobius (Gobioidei, Gobiidae, Gobionellinae). The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 15, 23-42.
72. Larson, H.K. (2001) A revision of the gobiid fish genus Mugilogobius (Teleostei: Gobioidei), with discussion of its systematic placement. Records of the Western Australian Museum, Supplement No. 62, 1-233.
73. Larson, H.K. & Buckle, D. (2005) A new species of the circumtropical goby genus Gnatholepis Bleeker (Teleostei, Gobiidae, Gobionellinae) from northern Australia. The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 21, 67-72.
74. Larson, H.K. & Murdy, E.O. (2001) Gobiidae. Gobies. In: Carpenter, K.E. & Niem, V.H. (Eds), FAO Species Identification Guide for Fishery Purposes. The Living Marine Resources of the Western Central Pacific. Volume 6. Bony fishes part 4 (Labridae to Latimeriidae). FAO, Rome, pp. 3578-3603.
75. Larson, H.K. & Wright, J. (2003) A new genus for the species Gobius baliurus Valenciennes and Gnatholepis calliurus Jordan and Seale (Teleostei, Gobiidae, Gobiinae). The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 19, 127-135.
76. Leviton, A.E., Gibbs, R.H., Heal, E. & Dawson, C.E. (1985) Standards in Herpetology and Ichthyology: Part 1. Standard symbolic codes for institutional resource collections in Herpetology and Ichthyology. Copeia, 3, 802-832.
77. Lobel, P.S. (2003) Marine Life of Johnston Atoll, Central Pacific Ocean. Natural World Press, Orgeon, 128 pp.
78. Lobel, P.S. & Lobel, L.K. (2004) Annotated checklist of the fishes of Wake Atoll. Pacific Science, 58(1), 65-90. (Pubitemid 38492763)
79. Lubbock, R. (1980) The shore fishes of Ascension Island. Journal of Fish Biology, 17, 283-303.
80. Manilo, L.G. & Bogorodsky, S.V. (2003) Taxonomic composition, diversity and distribution of coastal fishes of the Arabian Sea. Journal of Ichthyology, 43(Suppl. 1), S75-149.
81. Masuda, H., Amaoka, K., Araga, C., Ueno, T. & Yoshino, T. (Eds) (1984) The Fishes of the Japanese Archipelago. Tokai University Press, Tokyo, 437 pp.
82. Maugé, A.L. (1986) Gobiidae. In: Daget, J., Gosse, J.-P. & van den Audenaerde, D.F.E. (Eds), Check-list of the Fresh-Water Fishes of Africa. CLOFFA 2. ISNB, Brussells; MRAC, Tervuren; & ORSTOM, Paris, pp. 358-388.
83. Motomura, H., Kuriiwa, K., Katayama, E., Senou, H., Ogihara, G., Meguro, M., Matsunuma, M., Takat, Y., Yoshida, T., Yamashita, M., Kimuira, S., Endo, H., Murase, A., Iwatsuki, Y., Sakurai, Y., Harazaki, S., Hidaka, K., Izumi, H. & Matsuura, K. (2010) Annotated checklist of marine and estuarine fishes of Yaku-shima Island, Kagoshima, southern Japan. In: Motomura, H. & Matsuura, K. (Eds), Fishes of Yaku-shima Island. A World Heritage Island in the Osumi Group, Kagoshima Prefecture, Southern Japan. National Museum of Science and Nature, Tokyo, pp. 65-247.
84. Michiels, N.K., Anthes, N., Hart, N.S., Herler, J., Meixner, A.J., Schleifenbaum, F., Schulte, G., Siebeck, U.E., Sprenger, D. & Wucherer, M.F. (2008) Red fluorescence in reef fish: A novel signalling mechanism? BMC Ecology 8, 1-14 (doi:10.1186/1472-6785-8-16)
85. Miller, P.J. (1990) Gobiidae. In: Quero, J.C., Hureau, J.C., Karrer. C., Post, A. & Saldanha, L. (Eds), Clofeta II. Checklist of the Fishes of the Eastern Tropical Atlantic. UNESCO, Portugal, pp 925-961.
86. Mooi, R.D. & Gill, A.C. (2010) Phylogenies without synapomorphies - a crisis in fish systematics: time to show some character. Zootaxa, 2450, 26-40.
87. Murdy, E.O. (1985) A review of the gobiid fish genera Exyrias and Macrodontogobius, with description of a new species of Exyrias. Indo-Pacific Fishes, 10, 1-14.
88. Murdy, E.O. & Hoese, D.F. (1984) The monotypic gobiid fish genus Macrodontogobius, including synonymization of Gnatholepis hendersoni. Copeia, 1, 227-229.
89. Murdy, E.O. & Hoese, D.F. (2003) Gobiidae. In: Carpenter, K. E. (Ed.), The Living Marine Resources of the Western Central Atlantic. Volume 3: Bony Fishes part 2 (Opistognathidae to Molidae). FAO species identification guide for fishery purposes and American Society of Ichthyologist and Herpetologists Special Publication No. 5, FAO, Rome, pp. 1781-1796.
90. Myers, R.F. (1991) Micronesian Reef Fishes. A Practical Guide to the Identification of the Coral Reef Fishes of the Tropical and Western Pacific. Second edition. Coral Graphics, Barrigada, 298 pp.
91. Myers, R.F. (1999) Micronesian reef fishes. A Comprehensive Guide to the Coral Reef Fishes of Micronesia. Third revised edition. Coral Graphics, Barrigada, 330 pp.
92. Myers, R.F. & Donaldson, T.J. (2003) The fishes of the Mariana Islands. Micronesica, 35-36, 594-648.
93. Nakabo, T. (2002) (Ed.) Fishes of Japan with Pictorial Keys to the Species, English Edition II. Tokai University Press, Tokyo, 1748 pp.
94. Nelson, J.S., Crossman, E.J., Espinosa-Pérez, H., Findley, L.T., Gilbert, C.R., Lea, R.N. & Williams, J.D. (2004) Common and Scientific Names of Fishes From the United States, Canada, and Mexico. Sixth Edition. American Fisheries Society, Special Publication, pp. 291-386.
95. Palmer, G. (1970) New records, and one new species, of teleost fishes from the Gilbert Islands. Bulletin of the British Museum (Natural History) Zoology, 19(6), 211-234.
96. Parenti, L.R. & Thomas, K.R. (1998) Pharyngeal jaw morphology and homology in sicydiine gobies (Teleostei: Gobiidae) and allies. Journal of Morphology, 237, 257-274. (Pubitemid 28422194)
97. Peters, W. (1855a) Übersicht der in Mossambique beobachteten Seefische. Monatsberichte der Königllich Preussischen Akademie Wissenschaften Berlin, 1855, 428-466.
98. Peters,W. (1855b) Uebersicht der in Mossambique beobachteten Fische. Archiv fur Naturgeschichte, 21(Part 2-3), 234-282.
99. Pezold, F. (1993) Evidence for a monophyletic Gobiinae. Copeia, 3, 634-643.
100. Randall, J.E. (1968) Caribbean Reef Fishes. T.F.H. Publications, Neptune City, 318 pp.
101. Randall, J.E. (1985) Fishes. In: Delesalle, B., Galzin, R. & Salvat, B. (Eds) Fifth International Coral Reef Congress, Tahiti, 27 May-1 June, 1985, Volume 1, French Polynesian Coral Reefs, pp. 379-520.
102. Randall, J.E. (1995) Coastal Fishes of Oman. Crawford House, Bathurst, 439 pp.
103. Randall, J.E. (1999) Report on fish collections from the Pitcairn Islands. Atoll Research Bulletin, 461, 1-36 (+14 unnumbered pages of figures).
104. Randall, J.E. (2009) A review of the gobiid fishes of Easter Island, with description of a new species. aqua, International Journal of Ichthyology, 15(4), 177-190.
105. Randall, J.E. & Earle, J.L. (2000) Annotated checklist of the shore fishes of the Marquesas Islands. Bishop Museum Occasional Papers, 66, 1-39.
106. Randall, J.E. & Goren, M. (1993) A review of the gobioid fishes of the Maldives. Ichthyological Bulletin of the J.L.B. Smith Institute of Ichthyology, 58, 1-37.
107. Randall, J.E. & Greenfield, D.W. (2001) A preliminary review of the Indo-Pacific gobiid fishes of the genus Gnatholepis. Ichthyological Bulletin, 69, 1-17.
108. Randall, J.E. & Greenfield, D.W. (2007) Redescription of Gnatholepis cauerensis (Bleeker, 1853), with discussion of the validity of the species. Zoologische Mededeelingen Leiden, 81(16), 303-308.
109. Randall, J.E. & Lim, K.K.P. (2000) A checklist of the fishes of the South China Sea. Raffles Bulletin of Zoology, Supplement, 8, 569-667.
110. Randall, J.E. & Randall, H.A. (1987) Annotated checklist of the fishes of Enewatak Atoll and other Marshall Islands. In: Devaney, D.M., Reese, E.S., Burch, B.L. & Helfrich, P. (Eds), The Natural History of Enewetak Atoll, Volume II, Biogeography and Systematics. U.S. Department of Energy, Office of Scientific and Technical Information, Oak Ridge, Tennessee, pp. 289-324.
111. Randall, J.E., Lobel, P.S. & Chave, E.H. (1985) Annotated checklist of the fishes of Johnston Island. Pacific Science, 39(1), 24-80.
112. Randall, J.E., Allen, G.R. & Steene, R.C. (1990a) Fishes of the Great Barrier Reef and Coral Sea. Crawford House Press, Bathurst, 506 pp.
113. Randall, J.E., Smith, C.L. & Feinberg, M.N. (1990b) Report on fish collections from Rapa, French Polynesia. American Museum Novitates, 2996, 1-42.
114. Randall, J.E., Earle, J.L., Pyle, R.L., Parrish, J.D. & Hayes, T. (1993) Annotated checklist of the fishes of Midway Atoll, northwestern Hawaiian Islands. Pacific Science, 47(4), 356-400. (Pubitemid 24417315)
115. Randall, J.E., Cea, A.E. & Meléndez, R.C. (2005) Checklist of shore and epipelagic fishes of Easter Island, with twelve new records. Boletin del Museo Nacional de Historia Natural, Chile, 54, 41-55.
116. Randall, J.E., Williams, J.T., Smith, D.G., Kulbicki, M., Mou Tham, G., Labrosse, P. & others. (2004) Checklist of the shore and epipelagic fishes of Tonga. Atoll Reseach Bulletin, 502, 1-35.
117. Robins, C.R. & Ray, G.C. (1986) A Field Guide to Atlantic Coast Fishes. North America. Houghton Mifflin Company, Boston, 354 pp.
118. Rocha, L.A. & Rosa, I.L. (2001) Baseline assessment of reef fish assemblages of Parcel Manuel Luiz Marine State Park, Maranhão, north-east Brazil. Journal of Fish Biology, 58, 985-998. (Pubitemid 32546033)
119. Rocha, L.A., Robertson, D.R., Rocha, C.R., Van Tassell, J.L., Craig, M.T. & Bowen, B.W. (2005) Recent invasion of the tropical Atlantic by an Indo-Pacific coral reef fish. Molecular Ecology, 14, 3921-3928. (Pubitemid 44943727)
120. Ross, S.W. & Rhode, F.C. (2004) The gobioid fishes of North Carolina (Pisces: Gobioidei). Bulletin of Marine Science, 74(2), 287-323. (Pubitemid 38826239)
121. Roxas, H.A. & Ablan, G.L. (1940) New Philippine gobioid fishes. The Philippine Journal of Science, 73(3), 301-311.
122. Sanzo, L. (1911) Distribuzione delle papille cutanee (organi ciatiformi) e suo valore sistematico nei Gobi. Mitteilungen aus der Zoologischen Station zu Neapel, 20, 249-328.
123. Sauvage, H.E. (1891) Histoire physique, naturelle et politique de Madagascar. Volume XVI. Histoire naturelle des poissons. Histoire Naturelle des Poissons de Madagascar, 1-543.
124. Schwartz, F.J. (1999) Gobioid fishes of North Carolina. Journal of the Elisha Mitchell Scientific Society, 115(4), 281-293.
125. Seale, A. (1901) Director's report for 1900. Part II - Fishes. Occasional Papers of the Bernice Pauahi Bishop Museum of Polynesian Ethnology and Natural History, 1(3), 61-128.
126. Seale, A. (1910) New species of Philippine fishes. Philippine Journal of Science, 4(6), 491-543.
127. Senou, H., Suzuki, T., Shibukawa, K. & Yano, K. (2004) A Photographic Guide to the Gobioid Fishes of Japan. Heibonsha, Tokyo, 533 pp.
128. Shulman, M.J. & Bermingham, E. (1995) Early life histories, ocean currents, and the population genetics of Caribbean reef fishes. Evolution, 49(5), 897-910. (Pubitemid 3002811)
129. Smith, C.L., Tyler, J.C., Davis, W.P., Jones, R.S., Smith, D.G. & Baldwin, C.C. (2003) Fishes of the Pelican Cays, Belize. Atoll Reseach Bulletin, 497, 1-88.
130. Smith, J.J.F. (1989) The response of Asterropteryx semipunctatus and Gnatholepis anjerensis (Pisces, Gobiidae) to chemical stimuli from injured conspecifics, an alarm response in gobies. Ethology, 81, 279-290.
131. Smith-Vaniz, W.F., Collette, B.B. & Luckhurst, B.E. (1999) Fishes of Bermuda: History, Zoogeography, Annotated Checklist, and Identification Keys. American Society of Ichthyologists and Herpetologists, Special Publication No. 4, i-x + 1-424, 1-12.
132. Snyder, J.O. (1909) Descriptions of new genera and species of fishes from Japan and the Riu Kiu Islands. Proceedings of the U. S. National Museum, 36(1688), 597-610.
133. Sponaugle, S. & Cowen, R.K. (1994) Larval durations and recruitment patterns of two Caribbean gobies (Gobiidae): contrasting early life histories in demersal spawners. Marine Biology, 120, 133-143.
134. Suzuki, T. & Randall, J.E. (2009) Gnatholepis yoshinoi, a new gobiid fish from Okinawa, Japan. Bulletin of the National Museum of Natural Science, Series A, 35(2), 83-88.
135. Thacker, C. (2003) Molecular phylogeny of the gobioid fishes (Teleostei: Perciformes: Gobioidei). Molecular Phylogenetics and Evolution, 26, 354-368. (Pubitemid 36386495)
136. Thacker, C. (2004a) Phylogeny and species boundaries in the gobiid genus Gnatholepis (Teleostei: Perciformes). Zoological Journal of the Linnean Society, 142b, 573-582. (Pubitemid 40068272)
137. Thacker, C. (2004b) Population structure in two species of the reef goby Gnatholepis (Teleostei: Perciformes) among four South Pacific island groups. Coral Reefs, 23, 357-366. (Pubitemid 39320597)
138. Thacker, C. (2009) Phylogeny of Gobioidei and placement within Acanthomorpha, with a new classification and investigation of diversification and character evolution. Copeia, 1, 93-104.
139. Thacker, C.R. & Cole, K.S. (2002) Phylogeny and evolution of the gobiid genus Coryphopterus. Bulletin of Marine Science, 70(3), 837-850. (Pubitemid 36128280)
140. Thacker, C., Thompson, A.R., Roje, D.M. & Shaw, E.Y. (2008) New expansions in old clades: population genetics and phylogeny of Gnatholepis species (Teleostei: Gobioidei) in the Pacific. Marine Biology, 153, 375-385. (Pubitemid 350234020)
141. Wass, R.C. (1984) An annotated checklist of the fishes of Samoa. NOAA Technical Report, National Marine Fisheries Service, Special Scientific Report - Fisheries, 781, 1-43.
142. Watson, R.E. (1992) A review of the gobiid fish genus Awaous from insular streams of the Pacific Plate. Ichthyological Exploration of Freshwaters, 3(2), 161-176.
143. Whitley, G.P. (1958) Descriptions and records of fishes. Proceedings of the Royal Zoological Society of New South Wales, 1956-1957, 28-51.
144. Williams, J.T., Wantiez, L., Chauvet, C., Galzin, R., Harmelin-Vivien, M., Jobet, E., Juncker, Mou-tham, G., Planes, S. & Sasal, P. (2006) Checklist of the shorefishes of Wallis Islands (Wallis and Futuna French Territories, South-Central Pacific). Cybium, 30(3), 247-260. (Pubitemid 46167078)
145. Winterbottom, R. & Emery, A.R. (1986) Review of the gobioid fishes of the Chagos Archipelago, central Indian Ocean. Royal Ontario Museum, Life Sciences Contributions, (142), 1-82.
146. Zug, G.R., Springer, V.G., Williams, J.T. & Johnson, G.D. (1989) The vertebrates of Rotuma and surrounding waters. Atoll Reseach Bulletin, 316, 1-25.