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Volumn 7, Issue 3, 2012, Pages

Why do species co-occur? a test of alternative hypotheses describing abiotic differences in sympatry versus allopatry using spadefoot toads

Author keywords

[No Author keywords available]

Indexed keywords

ALLOPATRY; ARTICLE; CLIMATE CHANGE; COMPARATIVE STUDY; CONTROLLED STUDY; ECOLOGICAL NICHE; ENVIRONMENTAL FACTOR; ENVIRONMENTAL TEMPERATURE; FEMALE; GEOGRAPHICAL VARIATION (SPECIES); HYBRIDIZATION; MALE; MAXENT; NONHUMAN; ORGANISMAL INTERACTION; PREDICTION AND FORECASTING; PROBABILITY; REPRODUCTION; SPEA BOMBIFRONS; SPEA MULTIPLICATA; SPECIES COEXISTENCE; SPECIES DIFFERENCE; SPECIES DIVERSITY; SPECIES HABITAT; STATISTICAL MODEL; SURVIVAL; SYMPATRY; TOAD; ACCLIMATIZATION; ANIMAL; ANURA; BIODIVERSITY; BIOLOGICAL MODEL; CLASSIFICATION; ENVIRONMENT; GENETIC SELECTION; GENETICS; GEOGRAPHY; GROWTH, DEVELOPMENT AND AGING; NORTH AMERICA; SPECIES DIFFERENTIATION;

EID: 84859113850     PISSN: None     EISSN: 19326203     Source Type: Journal    
DOI: 10.1371/journal.pone.0032748     Document Type: Article
Times cited : (27)

References (58)
  • 1
    • 0000180725 scopus 로고
    • Laws of temperature control the geographic distribution of terrestrial animals and plants
    • Merriam CH, (1894) Laws of temperature control the geographic distribution of terrestrial animals and plants. National Geographic 6: 229-238.
    • (1894) National Geographic , vol.6 , pp. 229-238
    • Merriam, C.H.1
  • 2
    • 0002404879 scopus 로고
    • Observations on the behavior of some species at the edges of their ranges
    • Griggs RF, (1914) Observations on the behavior of some species at the edges of their ranges. Bulletin of the Torrey Botanical Club 41: 25-49.
    • (1914) Bulletin of the Torrey Botanical Club , vol.41 , pp. 25-49
    • Griggs, R.F.1
  • 3
    • 0000952820 scopus 로고
    • Field tests of theories concerning distributional control
    • Grinnell J, (1917) Field tests of theories concerning distributional control. Am Nat 51: 115-128.
    • (1917) Am Nat , vol.51 , pp. 115-128
    • Grinnell, J.1
  • 4
    • 0000267002 scopus 로고
    • Population studies - animal ecology and demography - concluding remarks
    • Hutchinson GE, (1957) Population studies- animal ecology and demography- concluding remarks. Cold Spring Harb Symp Quant Biol 22: 415-427.
    • (1957) Cold Spring Harb Symp Quant Biol , vol.22 , pp. 415-427
    • Hutchinson, G.E.1
  • 7
    • 0001262894 scopus 로고
    • The influence of interspecific competition and other factors on the distribution of the barnacle Chthamalus stellatus
    • Connell JH, (1961) The influence of interspecific competition and other factors on the distribution of the barnacle Chthamalus stellatus. Ecology 42: 710-723.
    • (1961) Ecology , vol.42 , pp. 710-723
    • Connell, J.H.1
  • 8
    • 0021595669 scopus 로고
    • On the relationship between abundance and distribution of species
    • Brown JH, (1984) On the relationship between abundance and distribution of species. Am Nat 124: 255-279.
    • (1984) Am Nat , vol.124 , pp. 255-279
    • Brown, J.H.1
  • 10
    • 53349128037 scopus 로고    scopus 로고
    • Reproductive interference between animal species
    • Gröning J, Hochkirch A, (2008) Reproductive interference between animal species. Q Rev Biol 83: 257-282.
    • (2008) Q Rev Biol , vol.83 , pp. 257-282
    • Gröning, J.1    Hochkirch, A.2
  • 11
    • 70349330222 scopus 로고    scopus 로고
    • Character displacement: Ecological and reproductive responses to a common evolutionary problem
    • Pfennig K, Pfennig D, (2009) Character displacement: Ecological and reproductive responses to a common evolutionary problem. Q Rev Biol 84: 253-276.
    • (2009) Q Rev Biol , vol.84 , pp. 253-276
    • Pfennig, K.1    Pfennig, D.2
  • 12
    • 33646466356 scopus 로고    scopus 로고
    • GIS-based niche models reveal unifying climatic mechanisms that maintain the location of avian hybrid zones in a North American suture zone
    • 10.1111/j.1420-9101.2005.01066.x
    • Swenson NG, (2006) GIS-based niche models reveal unifying climatic mechanisms that maintain the location of avian hybrid zones in a North American suture zone. J Evol Biol 19: 717-725 10.1111/j.1420-9101.2005.01066.x.
    • (2006) J Evol Biol , vol.19 , pp. 717-725
    • Swenson, N.G.1
  • 13
    • 67650128115 scopus 로고    scopus 로고
    • The role of climate in the dynamics of a hybrid zone in Appalachian salamanders
    • 10.1111/j.1365-2486.2009.01867.x
    • Walls SC, (2009) The role of climate in the dynamics of a hybrid zone in Appalachian salamanders. Global Change Biol 15: 1903-1910 10.1111/j.1365-2486.2009.01867.x.
    • (2009) Global Change Biol , vol.15 , pp. 1903-1910
    • Walls, S.C.1
  • 14
    • 0029860602 scopus 로고    scopus 로고
    • Some genetic consequences of ice ages, and their role in divergence and speciation
    • 10.1111/j.1095-8312.1996.tb01434.x
    • Hewitt GM, (1996) Some genetic consequences of ice ages, and their role in divergence and speciation. Biol J Linn Soc 58: 247-276 10.1111/j.1095-8312.1996.tb01434.x.
    • (1996) Biol J Linn Soc , vol.58 , pp. 247-276
    • Hewitt, G.M.1
  • 15
    • 0030302587 scopus 로고    scopus 로고
    • The geographic range: Size, shape, boundaries, and internal structure
    • 10.1146/annurev.ecolsys.27.1.597
    • Brown JH, Stevens GC, Kaufman DM, (1996) The geographic range: Size, shape, boundaries, and internal structure. Annu Rev Ecol Syst 27: 597-623 10.1146/annurev.ecolsys.27.1.597.
    • (1996) Annu Rev Ecol Syst , vol.27 , pp. 597-623
    • Brown, J.H.1    Stevens, G.C.2    Kaufman, D.M.3
  • 16
    • 0001998834 scopus 로고    scopus 로고
    • Predicting the potential distribution of plant species in an alpine environment
    • 10.2307/3237224
    • Guisan A, Theurillat J, Kienast F, (1998) Predicting the potential distribution of plant species in an alpine environment. Journal of Vegetation Science 9: 65-74 10.2307/3237224.
    • (1998) Journal of Vegetation Science , vol.9 , pp. 65-74
    • Guisan, A.1    Theurillat, J.2    Kienast, F.3
  • 17
    • 0028178320 scopus 로고
    • Positive interactions in communities
    • 10.1016/0169-5347(94)90088-4
    • Bertness MD, Callaway R, (1994) Positive interactions in communities. Trends in Ecology & Evolution 9: 191-193 10.1016/0169-5347(94)90088-4.
    • (1994) Trends in Ecology & Evolution , vol.9 , pp. 191-193
    • Bertness, M.D.1    Callaway, R.2
  • 18
  • 19
    • 0037349347 scopus 로고    scopus 로고
    • Inclusion of facilitation into ecological theory
    • DOI: 10.1016/S0169-5347(02)00045-9
    • Bruno JF, Stachowicz JJ, Bertness MD, (2003) Inclusion of facilitation into ecological theory. Trends in Ecology & Evolution 18: 119-125 DOI:10.1016/S0169-5347(02)00045-9.
    • (2003) Trends in Ecology & Evolution , vol.18 , pp. 119-125
    • Bruno, J.F.1    Stachowicz, J.J.2    Bertness, M.D.3
  • 20
    • 0041709466 scopus 로고    scopus 로고
    • Facilitated invasion by hybridization of Sarcocornia species in a salt-marsh succession
    • 10.1046/j.1365-2745.2003.00794.x
    • Figueroa ME, Castillo JM, Redondo S, Luque T, Castellanos EM, et al. (2003) Facilitated invasion by hybridization of Sarcocornia species in a salt-marsh succession. J Ecol 91 (4): 616-626 10.1046/j.1365-2745.2003.00794.x.
    • (2003) J Ecol , vol.91 , Issue.4 , pp. 616-626
    • Figueroa, M.E.1    Castillo, J.M.2    Redondo, S.3    Luque, T.4    Castellanos, E.M.5
  • 21
    • 33646132044 scopus 로고    scopus 로고
    • Ecological opportunity and phenotypic plasticity interact to promote character displacement and species coexistence
    • Pfennig DW, Rice AM, Martin RA, (2006) Ecological opportunity and phenotypic plasticity interact to promote character displacement and species coexistence. Ecology 87: 769-779.
    • (2006) Ecology , vol.87 , pp. 769-779
    • Pfennig, D.W.1    Rice, A.M.2    Martin, R.A.3
  • 22
    • 33745080242 scopus 로고    scopus 로고
    • Neutral theory and the evolution of ecological equivalence
    • 10.1890/0012-9658(2006)87[1387:NTATEO]2.0.CO;2
    • Hubbell SP, (2006) Neutral theory and the evolution of ecological equivalence. Ecology 87: 1387-1398 10.1890/0012-9658(2006)87[1387:NTATEO]2.0.CO;2.
    • (2006) Ecology , vol.87 , pp. 1387-1398
    • Hubbell, S.P.1
  • 23
    • 24344493920 scopus 로고    scopus 로고
    • Predicting species distribution: Offering more than simple habitat models
    • 10.1111/j.1461-0248.2005.00792.x
    • Guisan A, Thuiller W, (2005) Predicting species distribution: Offering more than simple habitat models. Ecol Lett 8: 993-1009 10.1111/j.1461-0248.2005.00792.x.
    • (2005) Ecol Lett , vol.8 , pp. 993-1009
    • Guisan, A.1    Thuiller, W.2
  • 24
    • 22544437325 scopus 로고    scopus 로고
    • Energy, water and large-scale patterns of reptile and amphibian species richness in Europe
    • DOI: 10.1016/j.actao.2005.02.006
    • Rodríguez MÁ, Belmontes JA, Hawkins BA, (2005) Energy, water and large-scale patterns of reptile and amphibian species richness in Europe. Acta Oecol 28: 65-70 DOI:10.1016/j.actao.2005.02.006.
    • (2005) Acta Oecol , vol.28 , pp. 65-70
    • Rodríguez, M.A.1    Belmontes, J.A.2    Hawkins, B.A.3
  • 25
    • 0035989038 scopus 로고    scopus 로고
    • How fluctuating competition and phenotypic plasticity mediate species divergence
    • Pfennig DW, Murphy PJ, (2002) How fluctuating competition and phenotypic plasticity mediate species divergence. Evolution 56: 1217-1228.
    • (2002) Evolution , vol.56 , pp. 1217-1228
    • Pfennig, D.W.1    Murphy, P.J.2
  • 26
    • 33947142818 scopus 로고    scopus 로고
    • Field and experimental evidence for competition's role in phenotypic divergence
    • 10.1111/j.1558-5646.2007.00034.x
    • Pfennig DW, Rice AM, Martin RA, (2007) Field and experimental evidence for competition's role in phenotypic divergence. Evolution 61: 257-271 10.1111/j.1558-5646.2007.00034.x.
    • (2007) Evolution , vol.61 , pp. 257-271
    • Pfennig, D.W.1    Rice, A.M.2    Martin, R.A.3
  • 27
    • 0036753060 scopus 로고    scopus 로고
    • Differential selection to avoid hybridization in two toad species
    • Pfennig KS, Simovich MA, (2002) Differential selection to avoid hybridization in two toad species. Evolution 56: 1840-1848.
    • (2002) Evolution , vol.56 , pp. 1840-1848
    • Pfennig, K.S.1    Simovich, M.A.2
  • 28
    • 36048934935 scopus 로고    scopus 로고
    • Facultative mate choice drives adaptive hybridization
    • 10.1126/science.1146035
    • Pfennig KS, (2007) Facultative mate choice drives adaptive hybridization. Science 318: 965-967 10.1126/science.1146035.
    • (2007) Science , vol.318 , pp. 965-967
    • Pfennig, K.S.1
  • 32
    • 0004106843 scopus 로고
    • Analysis of a hybrid zone between the spadefoot toads Scaphiopus multiplicatus and Scaphiopus bombifrons
    • Ph.D. Dissertation. University of California, Riverside
    • Simovich MA, (1985) Analysis of a hybrid zone between the spadefoot toads Scaphiopus multiplicatus and Scaphiopus bombifrons. Ph.D. Dissertation. University of California, Riverside.
    • (1985)
    • Simovich, M.A.1
  • 33
    • 0033532050 scopus 로고    scopus 로고
    • Diet composition of three Anurans from the playa wetlands of northwest Texas
    • Anderson AM, Haukos DA, Anderson JT, (1999) Diet composition of three Anurans from the playa wetlands of northwest Texas. Copeia 1999: 515-520.
    • (1999) Copeia , vol.1999 , pp. 515-520
    • Anderson, A.M.1    Haukos, D.A.2    Anderson, J.T.3
  • 34
    • 0033693971 scopus 로고    scopus 로고
    • Character displacement in polyphenic tadpoles
    • Pfennig DW, Murphy PJ, (2000) Character displacement in polyphenic tadpoles. Evolution 54: 1738-1749.
    • (2000) Evolution , vol.54 , pp. 1738-1749
    • Pfennig, D.W.1    Murphy, P.J.2
  • 35
    • 0342536016 scopus 로고
    • Introgressive hybridization between the spadefoot toads Scaphiopus bombifrons and S. multiplicatus (Salientia: Pelobatidae)
    • Sattler PW, (1985) Introgressive hybridization between the spadefoot toads Scaphiopus bombifrons and S. multiplicatus (Salientia: Pelobatidae). Copeia 1985: 324-332.
    • (1985) Copeia , vol.1985 , pp. 324-332
    • Sattler, P.W.1
  • 36
    • 0347765893 scopus 로고    scopus 로고
    • A test of alternative hypotheses for the evolution of reproductive isolation between spadefoot toads: Support for the reinforcement hypothesis
    • Pfennig KS, (2003) A test of alternative hypotheses for the evolution of reproductive isolation between spadefoot toads: Support for the reinforcement hypothesis. Evolution 57: 2842-2851.
    • (2003) Evolution , vol.57 , pp. 2842-2851
    • Pfennig, K.S.1
  • 37
    • 33947636115 scopus 로고    scopus 로고
    • Ecological selection and hybrid fitness: Hybrids succeed on parental resources
    • Pfennig KS, Chunco AJ, Lackey ACR, (2007) Ecological selection and hybrid fitness: Hybrids succeed on parental resources. Evol Ecol Res 9: 341-354.
    • (2007) Evol Ecol Res , vol.9 , pp. 341-354
    • Pfennig, K.S.1    Chunco, A.J.2    Lackey, A.C.R.3
  • 38
    • 0027091072 scopus 로고
    • Polyphenism in spadefoot toad tadpoles as a locally adjusted evolutionarily stable strategy
    • Pfennig DW, (1992) Polyphenism in spadefoot toad tadpoles as a locally adjusted evolutionarily stable strategy. Evolution 46: 1408-1420.
    • (1992) Evolution , vol.46 , pp. 1408-1420
    • Pfennig, D.W.1
  • 39
    • 27644576843 scopus 로고    scopus 로고
    • Character displacement as the "best of a bad situation": Fitness trade-offs resulting from selection to minimize resource and mate competition
    • Pfennig KS, Pfennig DW, (2005) Character displacement as the "best of a bad situation": Fitness trade-offs resulting from selection to minimize resource and mate competition. Evolution 59: 2200-2208.
    • (2005) Evolution , vol.59 , pp. 2200-2208
    • Pfennig, K.S.1    Pfennig, D.W.2
  • 40
    • 27944446350 scopus 로고    scopus 로고
    • Maximum entropy modeling of species geographic distributions
    • 10.1016/j.ecolmodel.2005.03.026
    • Phillips SJ, Anderson RP, Schapire RE, (2006) Maximum entropy modeling of species geographic distributions. Ecol Model 190: 231-259 10.1016/j.ecolmodel.2005.03.026.
    • (2006) Ecol Model , vol.190 , pp. 231-259
    • Phillips, S.J.1    Anderson, R.P.2    Schapire, R.E.3
  • 41
    • 33645917058 scopus 로고    scopus 로고
    • Novel methods improve prediction of species' distributions from occurrence data
    • 10.1111/j.2006.0906-7590.04596.x
    • Elith J, Graham CH, Anderson RP, Dudík M, Ferrier S, et al. (2006) Novel methods improve prediction of species' distributions from occurrence data. Ecography 29: 129-151 10.1111/j.2006.0906-7590.04596.x.
    • (2006) Ecography , vol.29 , pp. 129-151
    • Elith, J.1    Graham, C.H.2    Anderson, R.P.3    Dudík, M.4    Ferrier, S.5
  • 43
    • 29544443445 scopus 로고    scopus 로고
    • Very high resolution interpolated climate surfaces for global land areas
    • 10.1002/joc.1276
    • Hijmans RJ, Cameron SE, Parra JL, Jones PG, Jarvis A, (2005) Very high resolution interpolated climate surfaces for global land areas. Int J Climatol 25: 1965-1978 10.1002/joc.1276.
    • (2005) Int J Climatol , vol.25 , pp. 1965-1978
    • Hijmans, R.J.1    Cameron, S.E.2    Parra, J.L.3    Jones, P.G.4    Jarvis, A.5
  • 44
    • 70350396934 scopus 로고    scopus 로고
    • Niche shift versus niche conservatism? Climatic characteristics of the native and invasive ranges of the mediterranean house gecko (Hemidactylus turcicus)
    • 10.1111/j.1466-8238.2009.00477.x
    • Roedder D, Loetters S, (2009) Niche shift versus niche conservatism? Climatic characteristics of the native and invasive ranges of the mediterranean house gecko (Hemidactylus turcicus). Global Ecol Biogeogr 18: 674-687 10.1111/j.1466-8238.2009.00477.x.
    • (2009) Global Ecol Biogeogr , vol.18 , pp. 674-687
    • Roedder, D.1    Loetters, S.2
  • 45
    • 35348859799 scopus 로고    scopus 로고
    • Biotic interactions improve prediction of boreal bird distributions at macro-scales
    • 10.1111/j.1466-8238.2007.00345.x
    • Heikkinen RK, Luoto M, Virkkala R, Pearson RG, Körber J, (2007) Biotic interactions improve prediction of boreal bird distributions at macro-scales. Global Ecology & Biogeography 16: 754-763 10.1111/j.1466-8238.2007.00345.x.
    • (2007) Global Ecology & Biogeography , vol.16 , pp. 754-763
    • Heikkinen, R.K.1    Luoto, M.2    Virkkala, R.3    Pearson, R.G.4    Körber, J.5
  • 46
    • 41449100102 scopus 로고    scopus 로고
    • Modeling of species distributions with Maxent: New extensions and a comprehensive evaluation
    • 10.1111/j.0906-7590.2008.5203.x
    • Phillips SJ, Dudik M, (2008) Modeling of species distributions with Maxent: New extensions and a comprehensive evaluation. Ecography 31: 161-175 10.1111/j.0906-7590.2008.5203.x.
    • (2008) Ecography , vol.31 , pp. 161-175
    • Phillips, S.J.1    Dudik, M.2
  • 47
    • 0030953391 scopus 로고    scopus 로고
    • A review of methods for the assessment of prediction errors in conservation presence/absence models
    • Fielding AH, Bell JF, (1997) A review of methods for the assessment of prediction errors in conservation presence/absence models. Environ Conserv 24: 38-49.
    • (1997) Environ Conserv , vol.24 , pp. 38-49
    • Fielding, A.H.1    Bell, J.F.2
  • 48
    • 28444445926 scopus 로고    scopus 로고
    • R Development Core Team. R Foundation for Statistical Computing, ISBN 3-900051-07-0
    • R Development Core Team. R Foundation for Statistical Computing (2005) R: A language and environment for statistical computing. 2.11.1. ISBN 3-900051-07-0.
    • (2005) R: A language and environment for statistical computing
  • 49
    • 70350755588 scopus 로고    scopus 로고
    • Biogeography and conservation of taxa from remote regions: An application of ecological-niche based models and GIS to North-African canids
    • DOI: 10.1016/j.biocon.2009.08.001
    • Brito JC, Acosta AL, Álvares F, Cuzin F, (2009) Biogeography and conservation of taxa from remote regions: An application of ecological-niche based models and GIS to North-African canids. Biol Conserv 142: 3020-3029 DOI:10.1016/j.biocon.2009.08.001.
    • (2009) Biol Conserv , vol.142 , pp. 3020-3029
    • Brito, J.C.1    Acosta, A.L.2    Álvares, F.3    Cuzin, F.4
  • 50
    • 42149161192 scopus 로고    scopus 로고
    • GIS-based niche models identify environmental correlates sustaining a contact zone between three species of European vipers
    • 10.1111/j.1472-4642.2007.00446.x
    • Martínez-Freiría F, Sillero N, Lizana M, Brito JC, (2008) GIS-based niche models identify environmental correlates sustaining a contact zone between three species of European vipers. Divers Distrib 14: 452-461 10.1111/j.1472-4642.2007.00446.x.
    • (2008) Divers Distrib , vol.14 , pp. 452-461
    • Martínez-Freiría, F.1    Sillero, N.2    Lizana, M.3    Brito, J.C.4
  • 51
    • 42049108849 scopus 로고    scopus 로고
    • Analysis of range expansion in two species undergoing character displacement: Why might invaders generally 'win' during character displacement?
    • 10.1111/j.1420-9101.2008.01518.x
    • Rice AM, Pfennig DW, (2008) Analysis of range expansion in two species undergoing character displacement: Why might invaders generally 'win' during character displacement? J Evol Biol 21: 696-704 10.1111/j.1420-9101.2008.01518.x.
    • (2008) J Evol Biol , vol.21 , pp. 696-704
    • Rice, A.M.1    Pfennig, D.W.2
  • 52
    • 0025587155 scopus 로고
    • The adaptive significance of an environmentally-cued developmental switch in an Anuran tadpole
    • 10.1007/BF00317349
    • Pfennig D, (1990) The adaptive significance of an environmentally-cued developmental switch in an Anuran tadpole. Oecologia 85: 101-107 10.1007/BF00317349.
    • (1990) Oecologia , vol.85 , pp. 101-107
    • Pfennig, D.1
  • 54
    • 34347233473 scopus 로고    scopus 로고
    • Projected distributions of novel and disappearing climates by 2100 AD
    • Williams JW, Jackson ST, Kutzbach JE, (2007) Projected distributions of novel and disappearing climates by 2100 AD. Proc Natl Acad Sci U S A 104: 5738-5742.
    • (2007) Proc Natl Acad Sci U S A , vol.104 , pp. 5738-5742
    • Williams, J.W.1    Jackson, S.T.2    Kutzbach, J.E.3
  • 55
    • 77954577840 scopus 로고    scopus 로고
    • Squeezed at the top: Interspecific aggression may constrain elevational ranges in tropical birds
    • Jankowski JE, Robinson SK, Levey DJ, (2010) Squeezed at the top: Interspecific aggression may constrain elevational ranges in tropical birds. Ecology 91: 1877-1884.
    • (2010) Ecology , vol.91 , pp. 1877-1884
    • Jankowski, J.E.1    Robinson, S.K.2    Levey, D.J.3
  • 56
    • 34249947530 scopus 로고    scopus 로고
    • Model projections of an imminent transition to a more arid climate in southwestern North America
    • 10.1126/science.1139601
    • Seager R, Ting M, Held I, Kushnir Y, Lu J, et al. (2007) Model projections of an imminent transition to a more arid climate in southwestern North America. Science 316: 1181-1184 10.1126/science.1139601.
    • (2007) Science , vol.316 , pp. 1181-1184
    • Seager, R.1    Ting, M.2    Held, I.3    Kushnir, Y.4    Lu, J.5
  • 57
    • 34548411422 scopus 로고    scopus 로고
    • Poleward shifts in winter ranges of North American birds
    • La Sorte FA, Thompson FR, (2007) Poleward shifts in winter ranges of North American birds. Ecology 88: 1803-1812.
    • (2007) Ecology , vol.88 , pp. 1803-1812
    • La Sorte, F.A.1    Thompson, F.R.2


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