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note
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We thank N. Chiariello, C. Field, T. Tobeck, E. Cleland, E. Hadly, J. Kriewall, D. Croll, R. Shaw, the Carnegie Institution of Washington, and the Jasper Ridge Biological Preserve for their valuable contributions and assistance. K. Andonian, D. Doak, J. Dukes, C. Gilbert, P. Holloran, D. Hooper, K. Honey, D. Letourneau, J. Levine, B. Tershy, and two anonymous reviewers improved the manuscript. This project was generously supported by a David H. Smith Conservation Research Fellowship through The Nature Conservancy.
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