A reassessment of bunodont metatherians from the paleogene of itaboraí (Brazil): Systematics and age of the Itaboraian salma

Revista Brasileira de Paleontologia

Volumn 14, Issue 2, 2011, Pages 105-136

  Oliveira, Édison V ^a   Goin, Francisco J ^b  

^a FEDERAL UNIVERSITY OF PERNAMBUCO   (Brazil)

^b FACULTAD DE CIENCIAS NATURALES Y MUSEO   (Argentina)

Author keywords

"Ameridelphia"; Australidelphia; Chronostratigraphy; Didelphimorphia; Eocene; Metatheria

Indexed keywords

DIDELPHIDAE; DIDELPHIMORPHIA; GUGGENHEIMELLA; MASTODONTOIDEA; METATHERIA;

EID: 80052497038     PISSN: 15197530     EISSN: 22361715     Source Type: Journal    
DOI: 10.4072/rbp.2011.2.01     Document Type: Article

References (110)

1. Almeida, F.F.M.de.; Carneiro, C.R. & Mizusaki, A.M.P. 1996. Correlação do magmatismo das bacias da margem continental brasileira com o das áreas emersas adjacentes. Revista Brasileira de Geociências, 26(3):125-138.
2. Ameghino, F. 1894. Synoptic enumeration of the fossil mammalian species from the Eocene formations of Patagonia. Boletín de la Academia de Ciencias de Córdoba, 13:259-452.
3. Aplin, K.O. & Archer, M. 1987. Recent advances in marsupial systematics with a new syncretic classification. In: M. Archer (ed.) Possums and Opossums: Studies in Evolution, Surrey Beatty & Sons and the Royal Zoological Society of New South Wales, p. 15-72.
4. Archer, M. 1984. The Australian marsupial radiation. In: M. Archer & G. Clayton (eds.) Vertebrate zoogeography and evolution in Australasia, Hesperian Press, p. 633-808.
5. Archer, M.; Godthelp, H. & Hand, S.J. 1993. Early Eocene marsupial from Australia. Kaupia, 3:193-200.
6. Beck, R.M.D.; Godthelp, H.; Weisbecker, V.; Archer, A. & Hand, S.J. 2008. Australia's oldest marsupial fossils and their biogeographical implications. PLoSONE, 3(3):e1858. doi:10.1371/journal.pone.0001858
7. Bergqvist, L.P.; Mansur, K.; Rodrigues, M.A.; Rodrigues-Francisco, B.H.; Perez, R.A.R. & Beltrão, M.C.M.C. 2008. Itaboraí Basin, State of Rio de Janeiro - the cradle of mammals in Brazil. In: M. Winge; C. Schobbenhaus; C.R.G. Souza; A.C.S. Fernandes; M. Berbert-Born & E.T. Queiroz (eds.) Sítios Geológicos e Paleontológicos do Brasil, CPRM, p.1-19.
8. Bond, M.; Carlini, A.A.; Goin, F.J.; Legarreta, L.; Ortiz-Jaureguizar, E.; Pascual, R. & Uliana, M.A. 1995. Episodes in South American land mammal evolution and sedimentation: testing their apparent concurrence in a Paleocene succession from central Patagonia. In: CONGRESO ARGENTINO DE PALEONTOLOGÍA Y BIOESTRATIGRAFÍA, 6, 1995. Actas, Trelew, p. 47-58.
9. Case, J.; Goin, F.J. & Woodburne, M. 2005. South American" Marsupials from the Late Cretaceous of North America and the Origin of Marsupial Cohorts. Journal of Mammalian Evolution, 12:461-494. doi: 10.1007/s10914-005-7329-3
10. Cifelli, R.L. 1993. Early Cretaceous mammal from North America and the evolution of marsupial dental characters. Proceedings National Academy of Sciences, 90:9413-9416.
11. Ciffeli, R.L. & Johanson, Z. 1994. New marsupial from the upper Cretaceous of Utah. Journal of Vertebrate Paleontology, 14(2):292-295.
12. Cifelli, R.L. & Muizon, C. de 1997. Dentition and jaw of Kokopellia juddi, a primitive marsupial or near-marsupial from the medial Cretaceous of Utah. Journal of Mammalian Evolution, 4:242-258.
13. Cifelli, R.L. & Muizon, C. de 1998. Marsupial mammal from the upper Cretaceous North Horn Formation, Central Utah. Journal of Paleontology, 72(3):532-537.
14. Clemens, W.A. 1966. Fossil mammals of the type Lance Formation Wyoming. Part II. Marsupialia. University of California Publications in Geological Sciences, 62:1-122.
15. Clemens, W.A. 1979. Marsupialia. In: J.A. Lillegraven; Z. Kielan- Jaworowska & W. Clemens (eds.) Mesozoic mammals: the first two-thirds of mammalian history. University of California Press, p.192-220.
16. Cordani, U.G. 1970. Idade do vulcanismo no Oceano Atlântico Sul. Boletim IGA, 1:9-75.
17. Crochet, J.-Y. 1979. Diversité systématique des Didelphidae (Marsupialia) européens tertiaires. Géobios, 12(3):365-378.
18. Crochet, J.-Y. 1980. Les Marsupiaux du Tertiaire d'Europe. Paris, Éditions de la Fondation Singer-Polignac, 279 p.
19. Crochet, J.-Y. & Sigé, B. 1983. Les mammiferes montiens de Hainin (Paleocene moyen de Belgique) Part III: Marsupiaux. Palaeovertebrata, 13(3):51-64.
20. Davis, B.M. 2007. A revision of "pediomyid" marsupials from the Late Cretaceous of North America. Acta Palaeontologica Polonica, 52(2):217-256.
21. Estravís-Fernandez, C. 1992. Estudo dos mamíferos do Eocénico inferior de Silveirinha (Baixo Mondego). Faculdade de Ciências e Tecnologia, Universidade Nova de Lisboa, Tese de Doutorado, 248 p.
22. Ferrari, A.L. 2001. Evolução tectônica do Graben da Guanabara. Programa de Pós-Graduação em Geociências, Universidade de São Paulo, Tese de Doutorado, 412 p.
23. França, R.L.; Del Rey, A.C.; Tagliari, C.V.; Brandão, J.R. & Fontanelli, P.R. 2007. Bacia de Mucuri. Boletim de Geociências da Petrobrás, 15(2):493-499.
24. Fox, R.C. 1983. Notes on the North American marsupials Herpetotherium and Peradectes. Canadian Journal Earth Sciences, 17:1489-1498.
25. Gelfo, J.N.; Goin, F.J.; Woodburne, M.O. & Muizon, C. de 2009. Biochronological relationships of the earliest South American paleogene mammalian faunas. Palaeontology, 52(1):251-269. doi: 10.1111/j.1475-4983.2008.00835.x
26. Godthelp, H.; Archer, M.; Cifelli, R.; Hand, S.J. & Coral, F.G. 1999. Earliest known Australian Tertiary mammal fauna. Nature, 356:514-516.
27. Godthelp, H.; Wroe, S. & Archer, M. 1999. A new marsupial from the Early Eocene Tingamarra local fauna of Murgon, Southeastern Queensland: a prototypical Australian marsupial? Journal of Mammalian Evolution, 6(3):289-313.
28. Goin, F.J. 1991. Los Didelphoidea (Mammalia, Marsupialia) del Cenozoico tardío de la Región Pampeana. Facultad de Ciencias Naturales y Museo de La Plata, Tese de Doutorado, 327 p.
29. Goin, F.J. 2003. Early marsupial radiations in South America. In: M. Jones; C. Dickman, & M. Archer (eds.) Predators with Pouches: the Biology of Carnivorous Marsupials, Commonwealth Scientific & Industrial Research Organization, p. 30-42.
30. Goin, F.J. 2006. A review of the Caroloameghiniidae, Paleogene South American "primate- like" marsupials (Didelphimorphia, Peradectoidea). Palaeontographica Abteilung A, 278:57-67.
31. Goin, F.J. & Candela, A.M. 1996. A new early Eocene Polydolopimorphian (Mammalia, Marsupialia) from Patagonia. Journal of Vertebrate Paleontology, 16(2):292-296.
32. Goin, F.J. & Candela, A.M. 2004. New Paleogene marsupials from the Amazon Basin of eastern Peru. In: K.E. Campbell (ed.) The Paleogene mammalian fauna of Santa Rosa, Amazonian Peru, Science Series, Natural History Museum of Los Angeles, p. 15-60.
33. Goin, F.J. & Oliveira, E.V. 2007. A new species of Gashternia (Marsupialia) from Itaboraí (Brazil). Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 24:309-313. doi:10.1127/0077-7749/2007/0245-0309
34. Goin, F.J.; Candela, A.M. & Forasiepi, A. 1997. New, middle Paleocene marsupials from Central Patagonia. In: ANNUAL MEETING OF THE SOCIETY OF VERTEBRATE PALEONTOLOGY, 57, 1997. Abstracts, New York, SVP, p. 49A.
35. Goin, F.J.; Oliveira, E.V. & Candela, A.M. 1998a. Carolocoutoia ferigoloi nov. gen. and sp. (Protodidelphidae), a new Paleocene "opossum-like" marsupial from Brazil. Palaeovertebrata, 27:145-154.
36. Goin, F.J.; A.M. Candela; Bond, M.; Pascual, R. & Escribano, V. 1998b. Una nueva ''comadreja'' (Mammalia, Marsupialia,?Polydolopimorphia) del Paleoceno de Patagonia, Argentina. Buenos Aires, Asociación Paleontológica Argentina, p. 71-78 (Publicación Especial 5).
37. Goin, F.J.; Zimicz, N.; Reguero, M.A.; Santillana, S.N.; Marenssi, S.A. & Moly, J.J. 2007. New marsupial (Mammalia) from the Eocene of Antarctica, and the origins and affinities of the Microbiotheria. Revista de la Asociación Geológica Argentina, 62:597-603.
38. Goin, F.J.; Candela, A.M.; Abello, M. A.; & Oliveira, E.V. 2010. Earliest South American paucituberculatans and their significance in the understanding of 'pseudodiprotodont' marsupial radiations. Zoological Journal of the Linnean Society, 155:867-884. doi: 10.1111/j.1096-3642.2008.00471.x
39. Hardenbol, J.; Thierry J.; Farley M.B.; Jacquin T.; de Graciansky P.-C. & Vail, P.R. 1998. Mesozoic and Cenozoic sequence chronostratigraphic framework of European basins. Jurassic sequence chronostratigraphy. In: P.C.de. Graciansky; J. Hardenbol; T. Jacquin, & P.R. Vail (eds.) Mesozoic and Cenozoic sequence stratigraphy of European Basins. SEPM, Special Publication, 60:3-13.
40. Hershkovitz, P. 1982. The staggered marsupial lower third incisor (I3). Géobios, Mémoire Special, 6:191-200.
41. Hershkovitz, P. 1995. The staggered marsupial lower incisor: hallmark of cohort Didelphimorphia, and description of a new genus and species with i3 from the Albian (Lower Cretaceous) of Texas. Bonner Zoologische Beiträge, 45(3/4):153-169.
42. Horovitz, I.; Martin, T.; Bloch, J.; Ladevèze, S.; Kurz, C. & Sánchez-Villagra, M. 2009. Cranial anatomy of the earliest marsupials and the origin of opossums. PLoS ONE, 4(12): e8278. doi:10.1371/journal.pone.0008278
43. Horovitz, I.; Ladèveze, S.; Argot, C.; Macrini, T.E.; Martin, T.; Hooker, J.J.; Kurz, C.; Muizon, C. de & Sánchez-Villagra, M.R. 2008. The anatomy of Herpetotherium cf. fugax Cope, 1873, a metatherian from the Oligocene of North America. Palaeontographica Abteilung A, 284:109-141.
44. Johanson, Z. 1996. Revison of the late Cretaceous North American marsupial genus Alphadon. Palaeontographica Abteilung A, 242:127-184.
45. Korth, W.W. 1994. Middle Tertiary marsupials (Mammalia) from North America. Journal of Paleontology, 68:376-96.
46. Ladevèze, S. 2004. Metatherian petrosals from the Late Paleocene of Itaboraí (Brazil), and their phylogenetic implications. Journal of Vertebrate Paleontology, 24:202-213.
47. Ladevèze, S. 2007. Petrosal bones of metatherian mammals from the Late Paleocene of Itaboraí (Brazil), and a cladistic analysis of petrosal features in metatherians. Zoological Journal of the Linnean Society, 150:85-115. doi: 10.1111/j.1096-3642.2007.00282.x
48. Ladevèze, S. & Muizon, C. de. 2010. Evidence of early evolution of Australidelphia (Metatheria, Mammalia) in South America: phylogenetic relationships of the metatherians from the Late Palaeocene of Itaboraí (Brazil) based on teeth and petrosal bones. Zoological Journal of the Linnean Society, 159:746-784. doi: 10.1111/j.1096-3642.2009.00577.x
49. Legarreta, L. & Uliana, M.A. 1994. Asociaciones de fósiles y hiatus en el Supracretácico- Neógeno de Patagonia: una perspective estratigráfico-secuencial. Ameghiniana, 31(3): 257-281.
50. Lima, M.R. & Souza-Cunha, F.L. 1986. Análise palinológica de um nível linhítico da Bacia de São José de Itaboraí, Terciário do Estado do Rio de Janeiro, Brasil. Anais da Academia Brasileira de Ciências, 58:579-588.
51. Lucas, P.W. 2004. Dental functional morphology. How teeth work. Cambridge, Cambridge University Press, 372 p.
52. Maas, M.C.; Thewissen, J.G.M. & Kappelman, J. 1998. Hypsamasia seni (Mammalia: Embrithopoda) and other mammals from the Eocene Kartal Formation of Turkey. Bulletin of Carnegie Museum of Natural History, 34:286-297.
53. Macedo, A.C.M. 1975. Microfósseis de uma fácies clástico-química da bacia calcária de São José de Itaboraí, RJ. Anais da Academia Brasileira de Ciências, 47(3/4):578.
54. Marshall, L.G. 1981. Review of the Hathlyacyninae, an extinct subfamily of South American "dog-like" marsupials. Fieldiana Geology, 7:1-120.
55. Marshall, L.G. 1982a. Systematics of the extinct South American marsupial Family Polydolopidae. Fieldiana Geology, 12:1-109.
56. Marshall, L.G. 1982b. A new genus of Caroloameghiininae (Marsupialia: Didelphoidea: Didelphidae) from the Paleocene of Brazil. Journal of Mammalogy, 63:709-716.
57. Marshall, L.G. 1984. The lower jaw of Eobrasilia coutoi Simpson, 1947, a unique didelphoid (not borhyaenoid) marsupial from the Paleocene of Brazil. Journal of Paleontology, 58:173-177.
58. Marshall, L.G. 1985. Geochronology and land-mammal biochronology of the Transamerican faunal interchange. In: F.G. Stehli, & S.D. Webb (eds.) The Great American Biotic Interchange, New York, Plenum Press, p. 49-85.
59. Marshall, L.G. 1987. Systematics of Itaboraian (Middle Paleocene) age "opossum-like" marsupials from the limestone quarry at São José de Itaboraí, Brazil. In: M. Archer (ed.) Possums and Opossums: Studies in Evolution. Surrey Beatty & Sons and the Royal Zoological Society of New South Wales, p. 91-160.
60. Marshall, L.G. & Muizon, C. de 1984. A new didelphid marsupial (Itaboraian) of São José de Itaboraí (Brazil). Comptes rendus de l'Académie des Sciences de Paris, 299(18): 1297-300.
61. Marshall, L.G.; Case, J. & Woodburne, M.O. 1990. Phylogenetic relationships of the families of marsupials. In: H.H. Genoways (ed.) Current Mammalogy, Plenum Press, v. 2, p. 433-505.
62. Marshall, L.G.; Sempere, T. & Butler, R.F. 1997. Chronostratigraphy of the Mammal-Bearing Paleocene of South America. Journal of South American Earth Sciences, 10(1):49-70.
63. Marshall, L.G.; Butler, R.F.; Drake, R.E. & Curtis, G.H. 1981. Calibration of the beginning of the Age of Mammals in Patagonia. Science, 212:43-45.
64. Martin, J.E.; Case, J.A.; Jagt, J.W.M.; Schulp, A.S. & Mulder, E.W.A. 2005. A new European marsupial indicates a late Cretaceous high-latitude transatlantic dispersal route. Journal of Mammalian Evolution, 12(3/4):495-511. doi:10.1007/s10914-005-7330-x
65. MacPhee, R.D.E. & Reguero, M. 2010. Reinterpretation of a Middle Eocene Record of Tardigrada (Pilosa, Xenarthra, Mammalia) from La Meseta Formation, Seymour Island, West Antarctica. American Museum Novitates, 3689:1-21. doi:10.1206/703.1
66. McKenna, M.C. & Bell, S.K. 1997. Classification of mammals above the species level. New York, Columbia University Press, 631 p.
67. Medeiros, R.A. & Bergqvist, L.P. 1999. Paleocene of the São José de Itaboraí Basin, Rio de Janeiro, Brazil: lithostratigraphy and biostratigraphy. Acta Geologica Leopoldensia, 22(48):3-22.
68. Milani, E.J.; Rangel, H.D.; Bueno, G.V.; Stica, J.M.; Winter, W.R.; Caixeta, J.M. & Neto, O.C.P. 2007. Brazilian sedimentary basins - stratigraphic charts. Boletim de Geociências da Petrobras, 15(2):183-205.
69. Mizusaki, A.M.P. & Mohriak W.U. 1992. Sequências vulcanosedimentares na região da Plataforma Continental de Cabo Frio, RJ. In: CONGRESSO BRASILEIRO DE GEOLOGIA, 37, 1992. Anais, São Paulo, SBG, 2:468-469.
70. Mizusaki A.M.P.; Alves, D.B. & Conceição J.C.J. 1994. Eventos magmáticos nas bacias do Espírito Santo, Mucuri e Cumuruxatiba. In: CONGRESSO BRASILEIRO DE GEOLOGIA, 38, 1994. Anais, Camboriú, SBG, p. 566-568.
71. Mizusaki, A.M.P.; Thomaz Filho, A. & Césero, P. de 1998. Ages of the magmatism and the opening of the South Atlantic Ocean. Pesquisas em Geociências, 25(2):47-57.
72. Mizusaki, A.M.P.; Thomaz-Filho A.; Milani E.J. & de Césero P. 2002. Mesozoic and Cenozoic igneous activity and its tectonic control in Northeastern Brazil. Journal of South American Earth Sciences, 15:183-198.
73. de Muizon, C. 1991. La fauna de mamíferos de Tiupampa (Paleoceno Inferior, Formación Santa Lucia), Bolivia. Revista Técnica de Yacimientos Petrolíferos Fiscales Bolivianos, 12(3/4):575-624.
74. de Muizon, C. Brito, I.M. 1993. Le bassin calcaire de São José de Itaboraí (Rio de Janeiro, Brésil): ses relations fauniques avec le site de Tiupampa (Cochabamba, Bolivie). Annales de Paléontologie, 79(3):233-269.
75. Oliveira, E.V. 1998. Taxonomia, filogenia e paleobiogeografia de marsupiais "poliprotodontes" do Mesopaleoceno da Bacia de Itaborai ́, Rio de Janeiro, Brasil. Programa de pós-graduação em Geociências, Universidade Federal do Rio Grande do Sul, Tese de Doutorado, 327 p.
76. Oliveira, E.V. 1999. Systematics of Itaboraian (Medial Paleocene) age "opossum like" marsupials from the Itaboraí Basin, Rio de Janeiro, Brazil. Anais da Academia Brasileira de Ciências, 71:811-812.
77. Oliveira, E.V. & Goin, F.J. 1999. Robertbutleria Marshall 1987: un probable sinónimo de Protodidelphis Paula Couto 1952 (Marsupialia, Didelphimorphia, Protodidelphidae). Ameghiniana, 36:17R-18R.
78. Oliveira, E.V. & Goin, F.J. 2006. Marsupiais do início do Terciário do Brasil: origem, irradiação e história biogeográfica. In: N.C. Cáceres & E.L.A. Monteiro-Filho (eds.) Os Marsupiais do Brasil, Universidade Federal de Santa Maria, p. 608-649.
79. de Paula Couto, C. 1952a. Fossil mammals from the begining of the Cenozoic in Brazil. Marsupialia: Polydolopidae and Borhyaenidae. American Museum Novitates, 1559:1-27.
80. de Paula Couto, C. 1952b. Fossil mammals from the beginning of the Cenozoic in Brazil. Marsupialia: Didelphidae. American Museum Novitates, 1567:1-26.
81. de Paula Couto, C. 1952c. A new name for Mirandaia ribeiroi Paula Couto, 1952. Journal of Mammalogy, 33:503.
82. de Paula Couto, C. 1961. Marsupiais fósseis do Paleoceno do Brasil. Anais da Academia Brasileira de Ciências, 33(3/4):321-333.
83. de Paula Couto, C. 1962. Didelfídeos fósiles del Paleoceno de Brasil. Revista del Museo Argentino de Ciencias Naturales "Bernardino Rivadavia", 112:135-66.
84. Paula Couto, C. de 1970. News on the fossil marsupials from the Riochican of Brazil. Anais da Academia Brasileira de Ciências, 42(1):19-34.
85. Paula Couto, C. de 1979. Tratado de Paleomastozoologia. Rio de Janeiro, Academia Brasileira de Ciências, 590 p.
86. Pascual, R. & Ortiz-Jaureguizar, E. 1991. El Ciclo faunístico Cochabambiano (Paleoceno temprano): su incidencia en la historia biogeográfica de los mamíferos sudamericanos. Revista Técnica de Yacimientos Petrolíferos Fiscales Bolivianos, 12:559-574.
87. Price, L.Y & de Paula Couto, C. 1946. Vertebrados fósseis do Eoceno Inferior de Itaboraí. Rio de Janeiro, Departamento Nacional da Produção Mineral, Divisão de Geologia e Mineralogia, 3 p. (Notas Preliminares e Estudos 31).
88. Reig, O.; Kirsch, J.A. & Marshall, L.G. 1987. Systematic relationships of the living and Neocenozoic American "Opossum-like" marsupials (Suborder Didelphimorphia), with comments on the classification of these and of the Cretaceous and Paleogene New World and European metatherians. In: M. Archer (ed.) Possums and Opossums: Studies in Evolution, Royal Zoological Society of New South Wales, p. 1-89.
89. Riccomini, C. 1989. O Rift Continental do Sudeste do Brasil. Programa de Pós-Graduação em Geociências, Universidade de São Paulo, Tese de Doutorado, 256 p.
90. Riccomini, C. & Rodrigues-Francisco, B.H. 1992. Idade potássioargôniodo derrame de ankaramito da Bacia de Itaboraí, Rio de Janeiro, Brasil: implicações tectônicas. In: CONGRESSO BRASILEIRO DE GEOLOGIA, 37, 1992. Boletim de Resumos, São Paulo, SBG, v. 2, p. 469-470.
91. Riccomini, C.; Coimbra, A.M.; Sant'Anna, L.G.; Brandt Neto, M. & Valarelli, J.V. 1996. Argilominerais do paleolago Tremembé e sistemas deposicionais relacionados (Paleógeno, Rift Continental do Sudeste do Brasil). Revista Brasileira de Geociências, 26:167-180.
92. Ride, W.D.L. 1964. A review of Australian fossil marsupials. Journal of the Royal Society of Western Australia, 47(4):97-131.
93. Rougier, G.W.; Wible, J.R. & Novacek, M.J. 1998. Implications of Deltatheridium specimens for early marsupial history. Nature, 396:459-463.
94. Sánchez-Villagra, M.R.; Ladevèze, S.; Horovitz I.; Argot, C.; Hooker, J.J.; Macrini, T.; Martin, T.; Moore-Fay, S.; Muizon, C. de; Schmelzle, T. & Asher, R.J. 2007. Exceptionally preserved North American Paleogene metatherians: adaptation and discovery of a major gap in the opossum fossil record. Proceedings of the Royal Society, Biology Letters, 3:318-322.
95. Sant'Anna, L.G., 1999. Geologia, mineralogia e gênese das esmectitas dos depósitos paleogênicos do Rift Continental do Sudeste do Brasil. Programa de Pós-graduação em Geociências, Universidade de São Paulo, Tese de Doutorado, 291 p.
96. Sigé, B.; Archer, M.; Crochet, J.Y.; Godthep, H.; Hand, S. & Beck, R. 2009. Chulpasia and Thylacotinga, late Paleocene-earliest Eocene trans-Antarctic Gondwanan bunodont marsupials: new data from Australia. Geobios, 42:813-823. doi:10.1016/j.geobios.2009.08.001
97. Simpson, G.G. 1935. Occurrence and relationships of Rio Chico fauna of Patagonia. American Museum Novitates, 818:1-21.
98. Simpson, G.G. 1945. The principles of the classification and a classification of mammals. Bulletin of the American Museum of Natural History, 85:1-350.
99. Simpson, G.G. 1947. A new Eocene marsupial from Brazil. American Museum Novitates, 1357:1-7.
100. Swofford, D.L. 2003. PAUP*: Phylogenetic Analysis Using Parsimony (* and Other Methods). Version 4. Sunderland, Sinauer Associates.
101. Szalay, F.S. 1982. A new appraisal of marsupial phylogeny and classification. In: M. Archer (ed.) Carnivorous Marsupials, Royal Zoological Society of New South Wales, p. 621-640.
102. Szalay, F.S. 1994. Evolutionary history of the Marsupialia and an analysis of osteological characters. New York, Cambridge University Press, 455 p.
103. Szalay, F.S. & Sargis, E.J. 2001. Model-based analysis of postcranial osteology of marsupials from the Palaeocene of Itaboraí (Brazil) and the phylogenetics and biogeography of Metatheria. Geodiversitas, 23(2):139-302.
104. Takahashi, F. 1974. Variação morfológica de incisivos em didelfídeos (Marsupialia, Didelphinae). Anais da Academia Brasileira de Ciências, 46(3/4):413-416.
105. Tejedor, M.F.; Goin, F.J.; Gelfo, J.N.; López, G.; Bond, M.; Carlini, A.A.; Scillato-Yané, G.J.; Woodburne, M.O.; Chornogubsky, L.; Aragón, E.; Reguero, M.A.; Czaplewski, N.J.; Vincon, S.; Martin, G.M. & Ciancio, M.R. 2009. New early Eocene mammalian fauna from western Patagonia, Argentina. American Museum Novitates, 3638:1-43. doi: 10.1206/577.1
106. Thomaz-Filho A.; de Cesero, P.; A.M.P. Mizusaki & Leão, J.G. 2005. Hot spot volcanic tracks and their implications for South American plate motion, Campos Basin (Rio de Janeiro State), Brazil. Journal of South American Earth Sciences, 18(3/4):383-389. doi:10.1016/j.jsames.2004.11.006
107. Tibana, P.; Castro, J.C., & Barrocas, S.L.S. 1984. Bacia de Itaboraí. In: CONGRESSO BRASILEIRO DE GEOLOGIA, 33, 1984. Anais, Rio de Janeiro, SBG, v. 12, p. 5309-5316.
108. Voss, R.S. & Jansa, S.A. 2009. Phylogenetic relationships and classification of didelphid marsupials, an extant radiation of New World Metatherian mammals. Bulletin of the American Museum of Natural History, 322:1-177. doi: 10.1206/322.1
109. Woodburne, M.O. & Case, J.A. 1996. Dispersal, vicariance and the late Cretaceous to early Tertiary land mammal biogeography from South America to Australia. Journal of Mammalian Evolution, 3:121-61.
110. Wroe, S.; Ebach, M.; Ahyong, S.; Muizon, C. de & Muirhead, J. 2000. Cladistic analysis of dasyuromorphian (Marsupialia) phylogeny using cranial and dental characters. Journal of Mammalogy, 81:1008-1024.