A review of Pachyvaranus crassispondylus Arambourg, 1952, a pachyostotic marine squamate from the latest Cretaceous phosphates of Morocco and Syria

Geological Magazine

Volumn 148, Issue 2, 2011, Pages 237-249

  Houssaye, Alexandra ^a   Bardet, Nathalie ^a   Rage, Jean Claude ^a   Suberbiola, Xabier Pereda ^b   Bouya, Baâdi ^c   Amaghzaz, Mbarek ^c   Amalik, Mohamed ^d  

^a MUSÉUM NATIONAL D'HISTOIRE NATURELLE   (France)

^b UNIVERSITY OF THE BASQUE COUNTRY UPV EHU   (Spain)

^c Centre Minier de Khouribga   (Morocco)

^d Centre Minier de Ben Guerir   (Morocco)

Author keywords

Maastrichtian; pachyosteosclerosis; Pachyvaranidae clade nov.; Pachyvaranus; Southern Tethys; Varanoidea

Indexed keywords

MAASTRICHTIAN; PACHYOSTEOSCLEROSIS; PACHYVARANIDAE CLADE NOV; PACHYVARANUS; SOUTHERN TETHYS; VARANOIDEA;

PHOSPHATES;

FOSSIL RECORD; LIZARD; MAASTRICHTIAN; NEW RECORD; NEW SPECIES; PALEOBIOGEOGRAPHY; PALEOECOLOGY; PALEOENVIRONMENT; PALEONTOLOGY; PHOSPHATE;

MOROCCO; SYRIAN ARAB REPUBLIC;

REPTILIA; SQUAMATA; TELEOSTEI; VARANOIDEA;

EID: 79951774205     PISSN: 00167568     EISSN: 14695081     Source Type: Journal    
DOI: 10.1017/S0016756810000580     Document Type: Article

References (34)

1. ARAMBOURG, C. 1952. Les vertébrés fossiles des gise-ments de phosphates (Maroc-Algérie-Tunisie). Notes et Mémoires du Service Géologique du Maroc 92, 1-372.
2. BARDET, N. In press. Maastrichtian marine reptiles of the Mediterranean Tethyan realm: a palaeobiogeographical approach. Bulletin de la Société géologique de France.
3. BARDET, N. & PEREDA SUBERBIOLA, X. 1996. Las faunas de reptiles marinos del Cretácico final de Europa (margen norte del Tetis mediterráneo). Revista Española de Paleontología 11(1), 91-9.
4. BARDET, N., CAPPETTA, H., PEREDA SUBERBIOLA, X., MOUTYS, M., ALMALEH, A. K., AHMAD, A. M., KHRATA, O. & GANNOUM, N. 2000. The marine vertebrate faunas from the Late Cretaceous phosphates of Syria. Geological Magazine 137(3), 269-90. (Pubitemid 30464312)
5. BARDET, N., PEREDA SUBERBIOLA, X., BOUYAHYAOUI, F., IAROCHENE, M., BOUYA, B. & AMAGHZAZ, M. 2004. Mosasaurus beaugei Arambourg, 1952 (Squamata, Mosasauridae) from the Late Cretaceous Phosphates of Morocco. Geobios 37(3), 315-24. (Pubitemid 38728589)
6. BARDET, N., PEREDA SUBERBIOLA, X., IAROCHENE, M., AMALIK, M. & BOUYA, B. 2005a. Durophagous Mosasauridae (Squamata) from the Upper Cretaceous phosphates of Morocco, with the description of a new species of Globidens. Netherlands Journal of Geosciences 84(3), 167-75. (Pubitemid 41569047)
7. BARDET, N., PEREDA SUBERBIOLA, X., IAROCHENE, M., BOUYA, B. & AMAGHZAZ, M. 2005b. A new species of Halisaurus from the Late Cretaceous phosphates of Morocco, and the phylogenetical relationships of the Halisaurinae (Squamata: Mosasauridae). Zoological Journal of the Linnean Society 143, 447-72. (Pubitemid 40406216)
8. BUFFRÉNIL, V. DE & RAGE, J. C. 1993. La " pachyostose" vertébrale de Simoliophis (Reptilia, Squamata): données comparatives et considérations fonctionnelles. Annales de Paléontologie 79, 315-35. (Pubitemid 24425002)
9. BUFFRÉNIL, V. DE, BARDET, N., PEREDA SUBERBIOLA, X. & BOUYA, B. 2008. Specialization of bone structure in Pachyvaranus crassispondylus Arambourg, 1952, an aquatic squamate from the Late Cretaceous of the southern Tethyan margin. Lethaia 41, 59-69. (Pubitemid 351255419)
10. CALDWELL, M. W. 2000. On the aquatic squamate Dolicho-saurus longicollis Owen, 1850 (Cenomanian, Upper Cretaceous), and the evolution of elongate necks in Squamates. Journal of Vertebrate Paleontology 20(4), 720-35. (Pubitemid 33228305)
11. CALDWELL, M. W. & CALVO, J. 2008. Details of a new skull and articulated cervical column of Dinilysia patagonica Woodward, 1901. Journal of Vertebrate Paleontology 28(2), 349-62. (Pubitemid 351983299)
12. CAMP, C. L. 1923. Classification of the lizards. Bulletin of the American Museum of Natural History 48, 289-481.
13. CAPPETTA, H. 1987. Mesozoic and Cenozoic Elasmo-branchii, Chondrichthyes II. In Handbook of Paleoichthyology (ed. H.-P. Schultze). Stuttgart, New York: Gustav Fischer Verlag, 193 pp.
14. CHALIFA, Y. & LEWY, Z. 1991. Early Maastrichtian marine teleosts from the Northern Negev, Israel. Israel Journal of Earth Sciences 40, 91-105.
15. COPE, E. D. 1872. On the families of fishes of the Cretaceous formations in Kansas. Proceedings of the American Philosophical Society 12, 327-57.
16. DHONDT, A. V., MALCHUS, N., BOUMAZA, L. & JAILLARD, E. 1999. Cretaceous oysters from North Africa: origin and distribution. Bulletin de la Société géologique de France 170, 67-76. (Pubitemid 29199331)
17. HOFFSTETTER, R. 1939. Contribution à l'étude des Elapidae actuels et fossiles et de l'ostéologie des ophidiens. Archives du Muséum d'Histoire Naturelle de Lyon 15, 1-78.
18. HOFFSTETTER, R. & GASC, J. P. 1967. Observations sur le squelette cervical et spécialement sur les hypapophyses des sauriens varanoïdes actuels et fossiles. Bulletin du Muséum National d'Histoire Naturelle 39(6), 1028-43.
19. HOFFSTETTER, R. & GASC, J. P. 1969. Vertebrae and ribs of modern reptiles. In Biology of the Reptilia (ed. C. Gans), pp. 201-310. London and New York: Academic Press.
20. HOUSSAYE, H. 2009. "Pachyostosis" in aquatic amniotes: a review. Integrative Zoology 4, 325-40.
21. LEE, M. S. Y. 1997. The phylogeny of varanoid lizards and the affinities of snakes. Philosophical Transactions of the Royal Society 352, 53-91. (Pubitemid 127738814)
22. LEE, M. S. Y. & SCANLON, J. D. 2002. The Cretaceous marine squamate Mesoleptos and the origin of snakes. Bulletin of The Natural History Museum of London (Zoology) 68(2), 131-42.
23. LEE, M. S. Y., CALDWELL, M. W. & SCANLON, J. D. 1999. A second primitive marine snake: Pachyophis woodwardi from the Cretaceous of Bosnia-Herzegovina. Journal of Zoology London 248, 509-20. (Pubitemid 29365575)
24. LUCAS, J. & PRÉVÔT-LUCAS, L. 1996. Tethyan phosphates and bioproductites. In The Ocean Basins and Margins. The Tethys Ocean (eds A. E. M. Nairn, L.-E. Ricou, B. Vrielynck & J. Dercourt), pp. 367-91. New York: Plenum Press.
25. MOLNAR, R. 2004. Dragons in the Dust: The Paleobiology of the Giant Monitor Lizard Megalania. Bloomington: Indiana University Press.
26. OPPEL, M. 1811. Die Ordnungen, Familien und Gattungen der Reptilien als Prodrom einer Naturgeschichte derselben. München: Lindauer.
27. RAGE, J. C. & NÉRAUDEAU, D. 2004. A new pachyostotic squamate reptile from the Cenomanian of France. Palaeontology 47(5), 1195-210. (Pubitemid 39338214)
28. RIEPPEL, O. 1980. The postcranial skeleton of Lanthanotus borneensis (Reptilia, Lacertilia). Amphibia-Reptilia 1, 95-112.
29. RIEPPEL, O. & GRANDE, L. 2007. The anatomy of the fossil varanid lizard Saniwa ensidens Leidy, 1870, based on a newly discovered complete skeleton. Journal of Paleontology 81(4), 643-65. (Pubitemid 47104921)
30. RUSSELL, D. A. 1967. Systematics and morphology of American mosasaurs. Peabody Museum of Natural History, Bulletin 23, 240 pp.
31. SMITH, K. T., BHULLAR, B. A. S. & HOLROYD, P. A. 2008. Earliest African record of the Varanus stem-clade (Squamata: Varanidae) from the Early Oligocene of Egypt. Journal of Vertebrate Paleontology 28(3), 909-13.
32. TAYLOR, M. A. 2000. Functional significance of bone ballast in the evolution of buoyancy control strategies by aquatic tetrapods. Historical Biology 14, 15-31. (Pubitemid 30314924)
33. WATKINS, J. 1999. Mechanical characteristics of musculo-skeletal components. In Structure and Function of the Musculoskeletal System (ed. J. Watkins), pp. 285-308. Champaign, IL: Human Kinetics.
34. ZABORSKI, P. M. & MORRIS, N. J. 1999. The Late Cretaceous ammonite genus Libycoceras in the Iullemmeden Basin (West Africa) and its palaeogeographical significance. Cretaceous Research 20, 63-79. (Pubitemid 29307961)