Developmental changes in spider spinning fields: A comparison between Mimetus and Araneus (Araneae: Mimetidae, Araneidae)

Biological Journal of the Linnean Society

Volumn 98, Issue 2, 2009, Pages 343-383

  Townley, Mark A ^a   Tillinghast, Edward K ^a  

^a UNIVERSITY OF NEW HAMPSHIRE   (United States)

Author keywords

Aciniform silk gland; Araneoidea; Australomimetus; Ero; Molt; Palpimanoidea; Phylogeny; Piriform silk gland; Sexual dimorphism; Tartipore

Indexed keywords

BIOLOGICAL DEVELOPMENT; COMPARATIVE STUDY; LIFE CYCLE; MOLT; ONTOGENY; PHYLOGENY; SCANNING ELECTRON MICROSCOPY; SEXUAL DIMORPHISM; SPIDER;

ARANEAE; ARANEIDAE; ARANEOIDEA; ARANEUS; ERO; MIMETIDAE; MIMETUS; PALPIMANOIDEA;

EID: 70349214920     PISSN: 00244066     EISSN: 10958312     Source Type: Journal    
DOI: 10.1111/j.1095-8312.2009.01297.x     Document Type: Article

References (141)

1. Agnarsson I. 2003. The phylogenetic placement and circumscription of the genus Synotaxus (Araneae: Synotaxidae), a new species from Guyana, and notes on theridioid phylogeny. Invertebrate Systematics 17 : 719 734.
2. Agnarsson I. 2004. Morphological phylogeny of cobweb spiders and their relatives (Araneae, Araneoidea, Theridiidae). Zoological Journal of the Linnean Society 141 : 447 626.
3. Agnarsson I. 2005. Revision and phylogenetic analysis of American ethicus and rupununi groups of Anelosimus (Araneae, Theridiidae). Zoologica Scripta 34 : 389 413.
4. Agnarsson I. 2006a. A revision of the New World eximius lineage of Anelosimus (Araneae, Theridiidae) and a phylogenetic analysis using worldwide exemplars. Zoological Journal of the Linnean Society 146 : 453 593.
5. Agnarsson I. 2006b. Asymmetric female genitalia and other remarkable morphology in a new genus of cobweb spiders (Theridiidae, Araneae) from Madagascar. Biological Journal of the Linnean Society 87 : 211 232.
6. Agnarsson I, Zhang J-X. 2006. New species of Anelosimus (Araneae: Theridiidae) from Africa and Southeast Asia, with notes on sociality and color polymorphism. Zootaxa 1147 : 1 34.
7. Álvarez-Padilla F. 2007. Systematics of the spider genus Metabus O.P.-Cambridge, 1899 (Araneoidea: Tetragnathidae) with additions to the tetragnathid fauna of Chile and comments on the phylogeny of Tetragnathidae. Zoological Journal of the Linnean Society 151 : 285 335.
8. Arnedo MA, Oromí P, Ribera C. 1996. Radiation of the genus Dysdera (Araneae, Haplogynae, Dysderidae) in the Canary Islands: the western islands. Zoologica Scripta 25 : 241 274.
9. Arnedo MA, Oromí P, Ribera C. 2000. Systematics of the genus Dysdera (Araneae, Dysderidae) in the eastern Canary Islands. Journal of Arachnology 28 : 261 292.
10. Arnedo MA, Ribera C. 1996. Dysdera ratonensis Wunderlich, 1991 (Arachnida, Araneae), a troglomorphic species from La Palma, Canary Islands: description of the male and redescription of the female. Revue Arachnologique 11 : 109 122.
11. Arnedo MA, Ribera C. 1997. Radiation of the genus Dysdera (Araneae, Haplogynae, Dysderidae) in the Canary Islands: the island of Gran Canaria. Zoologica Scripta 26 : 205 243.
12. Arnedo MA, Ribera C. 1999. Radiation of the genus Dysdera (Araneae, Dysderidae) in the Canary Islands: the island of Tenerife. Journal of Arachnology 27 : 604 662.
13. Barth FG. 2002. A spider's world: senses and behavior. Berlin : Springer-Verlag.
14. Blackledge TA, Scharff N, Coddington JA, Szüts T, Wenzel JW, Hayashi CY, Agnarsson I. 2009. Reconstructing web evolution and spider diversification in the molecular era. Proceedings of the National Academy of Sciences of the United States of America 106 : 5229 5234.
15. Bleher B. 2000. Development of web-building and spinning apparatus in the early ontogeny of Nephila madagascariensis (Vinson, 1863) (Araneae: Tetragnathidae). Bulletin of the British Arachnological Society 11 : 275 283.
16. Brignoli PM. 1979. Recherches en Afrique de l'Institut de Zoologie de l'Aquila (Italie). II. Reo latro nov. gen., nov. sp. du Kenya (Araneae: Mimetidae). Revue de Zoologie Africaine 93 : 919 928.
17. Brignoli PM. 1984. On some West Indian Mimetus and Lyssomanes (Araneae: Mimetidae, Salticidae). Bulletin of the British Arachnological Society 6 : 200 204.
18. Coddington JA. 1986. The genera of the spider family Theridiosomatidae. Smithsonian Contributions to Zoology 422 : i iv, 1 96.
19. Coddington JA. 1989. Spinneret silk spigot morphology: evidence for the monophyly of orbweaving spiders, Cyrtophorinae (Araneidae), and the group Theridiidae plus Nesticidae. Journal of Arachnology 17 : 71 95.
20. Coddington JA, Levi HW. 1991. Systematics and evolution of spiders (Araneae). Annual Review of Ecology and Systematics 22 : 565 592.
21. Dimitrov D, Ribera C. 2007. The genus Pholcus (Araneae, Pholcidae) in the Canary Islands. Zoological Journal of the Linnean Society 151 : 59 114.
22. Downes MF. 1987. A proposal for standardization of the terms used to describe the early development of spiders, based on a study of Theridion rufipes Lucas (Araneae: Theridiidae). Bulletin of the British Arachnological Society 7 : 187 193.
23. Eberhard WG. 1976. Physical properties of sticky spirals and their connections: sliding connections in orb webs. Journal of Natural History 10 : 481 488.
24. Fannes W, Jocqué R. 2008. Ultrastructure of Antoonops, a new, ant-mimicking genus of Afrotropical Oonopidae (Araneae) with complex internal genitalia. American Museum Novitates 3614 : 1 30.
25. Foelix RF. 1996. Biology of spiders, 2nd edn. New York, NY : Oxford University Press.
26. Forster RR, Platnick NI. 1984. A review of the archaeid spiders and their relatives, with notes on the limits of the superfamily Palpimanoidea (Arachnida, Araneae). Bulletin of the American Museum of Natural History 178 : 1 106.
27. Forster RR, Platnick NI, Coddington J. 1990. A proposal and review of the spider family Synotaxidae (Araneae, Araneoidea), with notes on theridiid interrelationships. Bulletin of the American Museum of Natural History 193 : 1 116.
28. Glatz L. 1973. Der Spinnapparat der Orthognatha (Arachnida, Araneae). Zeitschrift für Morphologie der Tiere 75 : 1 50.
29. Goloboff PA. 1995. A revision of the South American spiders of the family Nemesiidae (Araneae, Mygalomorphae). Part I: Species from Peru, Chile, Argentina, and Uruguay. Bulletin of the American Museum of Natural History 224 : 1 189.
30. Gorb SN, Barth FG. 1996. A new mechanosensory organ on the anterior spinnerets of the spider Cupiennius salei (Araneae, Ctenidae). Zoomorphology 116 : 7 14.
31. Gray MR. 2005. A revision of the spider genus Taurongia (Araneae, Stiphidioidea) from south-eastern Australia. Journal of Arachnology 33 : 490 500.
32. Griswold CE. 1997. The spider family Cyatholipidae in Madagascar (Araneae, Araneoidea). Journal of Arachnology 25 : 53 83.
33. Griswold CE. 2001. A monograph of the living world genera and Afrotropical species of cyatholipid spiders (Araneae, Orbiculariae, Araneoidea, Cyatholipidae). Memoirs of the California Academy of Sciences 26 : 1 251.
34. Griswold CE, Coddington JA, Hormiga G, Scharff N. 1998. Phylogeny of the orb-web building spiders (Araneae, Orbiculariae: Deinopoidea, Araneoidea). Zoological Journal of the Linnean Society 123 : 1 99.
35. Griswold CE, Coddington JA, Platnick NI, Forster RR. 1999. Towards a phylogeny of entelegyne spiders (Araneae, Araneomorphae, Entelegynae). Journal of Arachnology 27 : 53 63.
36. Griswold CE, Ramírez MJ, Coddington JA, Platnick NI. 2005. Atlas of phylogenetic data for entelegyne spiders (Araneae: Araneomorphae: Entelegynae) with comments on their phylogeny. Proceedings of the California Academy of Sciences, Fourth Series 56 Suppl. II : 1 324.
37. Harms D, Harvey MS. 2009a. A review of the pirate spiders of Tasmania (Arachnida, Mimetidae, Australomimetus) with description of a new species. Journal of Arachnology 37 : 188 205.
38. Harms D, Harvey MS. 2009b. Australian pirates: systematics and phylogeny of the Australasian pirate spiders (Araneae: Mimetidae), with a description of the Western Australian fauna. Invertebrate Systematics 23 : 231 280.
39. Heimer S. 1986. Notes on the spider family Mimetidae with description of a new genus from Australia (Arachnida, Araneae). Entomologische Abhandlungen, Staatliches Museum für Tierkunde, Dresden 49 : 113 137.
40. Hormiga G. 1994a. Cladistics and the comparative morphology of linyphiid spiders and their relatives (Araneae, Araneoidea, Linyphiidae). Zoological Journal of the Linnean Society 111 : 1 71.
41. Hormiga G. 1994b. A revision and cladistic analysis of the spider family Pimoidae (Araneoidea: Araneae). Smithsonian Contributions to Zoology 549 : i iii. 1-104.
42. Hormiga G. 2000. Higher level phylogenetics of erigonine spiders (Araneae, Linyphiidae, Erigoninae). Smithsonian Contributions to Zoology 609 : i iii. 1-160.
43. Hormiga G. 2002. Orsonwelles, a new genus of giant linyphiid spiders (Araneae) from the Hawaiian Islands. Invertebrate Systematics 16 : 369 448.
44. Hormiga G. 2003. Weintrauboa, a new genus of pimoid spiders from Japan and adjacent islands, with comments on the monophyly and diagnosis of the family Pimoidae and the genus Pimoa (Araneoidea, Araneae). Zoological Journal of the Linnean Society 139 : 261 281.
45. Hormiga G, Alvarez-Padilla F, Benjamin SP. 2007. First records of extant Hispaniolan spiders of the families Mysmenidae, Symphytognathidae, and Ochyroceratidae (Araneae), including a new species of Ochyrocera. American Museum Novitates 3577 : 1 21.
46. Hormiga G, Eberhard WG, Coddington JA. 1995. Web-construction behaviour in Australian Phonognatha and the phylogeny of nephiline and tetragnathid spiders (Araneae: Tetragnathidae). Australian Journal of Zoology 43 : 313 364.
47. Hormiga G, Scharff N. 2005. Monophyly and phylogenetic placement of the spider genus Labulla Simon, 1884 (Araneae, Linyphiidae) and description of the new genus Pecado. Zoological Journal of the Linnean Society 143 : 359 404.
48. Huber BA. 2000. New world pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History 254 : 1 348.
49. Huber BA. 2001. The pholcids of Australia (Araneae; Pholcidae): Taxonomy, biogeography, and relationships. Bulletin of the American Museum of Natural History 260 : 1 144.
50. Huber BA. 2003a. Cladistic analysis of Malagasy pholcid spiders reveals generic level endemism: revision of Zatavua n. gen. and Paramicromerys Millot (Pholcidae, Araneae). Zoological Journal of the Linnean Society 137 : 261 318.
51. Huber BA. 2003b. High species diversity in one of the dominant groups of spiders in East African montane forests (Araneae: Pholcidae: Buitinga n. gen., Spermophora Hentz). Zoological Journal of the Linnean Society 137 : 555 619.
52. Huber BA. 2003c. Southern African pholcid spiders: revision and cladistic analysis of Quamtana gen. nov. and Spermophora Hentz (Araneae: Pholcidae), with notes on male-female covariation. Zoological Journal of the Linnean Society 139 : 477 527.
53. Huber BA. 2004. Evolutionary transformation from muscular to hydraulic movements in spider (Arachnida, Araneae) genitalia: a study based on histological serial sections. Journal of Morphology 261 : 364 376.
54. Huber BA. 2005a. Revision and cladistic analysis of the spider genus Carapoia González-Sponga (Araneae: Pholcidae), with descriptions of new species from Brazil's Atlantic forest. Invertebrate Systematics 19 : 541 556.
55. Huber BA. 2005b. High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica 155 : 1 126.
56. Huber BA. 2005c. Revision of the genus Spermophora Hentz in Southeast Asia and on the Pacific Islands, with descriptions of three new genera (Araneae: Pholcidae). Zoologische Mededelingen Leiden 79 : 61 114.
57. Huber BA. 2007. Two new genera of small, six-eyed pholcid spiders from West Africa, and first record of Spermophorides for mainland Africa (Araneae: Pholcidae). Zootaxa 1635 : 23 43.
58. Huber BA, Benjamin SP. 2005. The pholcid spiders from Sri Lanka: redescription of Pholcus ceylonicus and description of a new genus (Araneae: Pholcidae). Journal of Natural History 39 : 3305 3319.
59. Huber BA, Brescovit AD, Rheims CA. 2005. Exaggerated female genitalia in two new spider species (Araneae: Pholcidae), with comments on genital evolution by female choice versus antagonistic coevolution. Insect Systematics and Evolution 36 : 285 292.
60. Huber BA, Deeleman-Reinhold CL, Pérez-González A. 1999. The spider genus Crossopriza (Araneae, Pholcidae) in the New World. American Museum Novitates 3262 : 1 10.
61. Huber BA, Rheims CA, Brescovit AD. 2005. Two new species of litter-dwelling Metagonia spiders (Araneae, Pholcidae) document both rapid and slow genital evolution. Acta Zoologica 86 : 33 40.
62. Huber BA, Wunderlich J. 2006. Fossil and extant species of the genus Leptopholcus in the Dominican Republic, with the first cases of egg-parasitism in pholcid spiders (Araneae: Pholcidae). Journal of Natural History 40 : 2341 2360.
63. Jackson RR. 1971. Fine structure of the thread connections in the orb web of Araneus diadematus. Psyche 78 : 12 31.
64. Kavanagh EJ, Tillinghast EK. 1979. Fibrous and adhesive components of the orb webs of Araneus trifolium and Argiope trifasciata. Journal of Morphology 160 : 17 31.
65. e Série 14 : 1 40.
66. e Série 19 : 63 87.
67. Kovoor J. 1986. L'appareil séricigène dans les genres Nephila Leach et Nephilengys Koch: anatomie microscopique, histochimie, affinités avec d'autres Araneidae. Revue Arachnologique 7 : 15 34.
68. Kovoor J. 1987. Comparative structure and histochemistry of silk-producing organs in arachnids. In : Nentwig W, ed. Ecophysiology of spiders. Berlin : Springer-Verlag, 160 186.
69. Kovoor J. 1990. The silk-gland system in some Tetragnathinae (Araneae: Araneidae). Comparative anatomy and histochemistry. Acta Zoologica Fennica 190 : 215 221.
70. Kovoor J, Lopez A. 1980. Variation de l'appareil séricigène dans la famille des Araneidae: Cas des genres Cyclosa Menge et Nemoscolus Simon. Comptes-Rendus de la Vème Colloque d'Arachnologie d'Expression Française, Septembre 1979, Barcelone 119 127.
71. Kovoor J, Lopez A. 1982. Anatomie et histologie des glandes séricigènes des Cyrtophora (Araneae, Araneidae): affinités et corrélations avec la structure et la composition de la toile. Revue Arachnologique 4 : 1 21.
72. Kovoor J, Lopez A. 1983. Composition et histologie de l'appareil séricigène des Argyrodes, relations avec le comportement de ces araignées (Theridiidae). Revue Arachnologique 5 : 29 43.
73. Kovoor J, Lopez A. 1988. L'appareil séricigène des Mecynogea Simon (Araneae, Araneidae). Revue Arachnologique 7 : 205 212.
74. Kovoor J, Peters HM. 1988. The spinning apparatus of Polenecia producta (Araneae, Uloboridae): structure and histochemistry. Zoomorphology 108 : 47 59.
75. e Série 16 : 5 25.
76. Kullmann E. 1975. Die Produktion und Funktion von Spinnenfäden und Spinnengeweben. In : Bach K, ed. Netze in Natur und Technik, IL8. Stuttgart : Instituts für leichte Flächentragwerke (IL), Universität Stuttgart, 318 378.
77. Kullmann E, Otto F, Braun T, Raccanello R. 1975. Grundlagen und Ordnung - Übersicht der Netzkonstruktionen der Spinnen. In : Bach K, ed. Netze in Natur und Technik, IL8. Stuttgart : Instituts für leichte Flächentragwerke (IL), Universität Stuttgart, 304 316.
78. Kuntner M. 2005. A revision of Herennia (Araneae: Nephilidae: Nephilinae), the Australasian 'coin spiders'. Invertebrate Systematics 19 : 391 436.
79. Kuntner M. 2006. Phylogenetic systematics of the Gondwanan nephilid spider lineage Clitaetrinae (Araneae, Nephilidae). Zoologica Scripta 35 : 19 62.
80. Kuntner M. 2007. A monograph of Nephilengys, the pantropical 'hermit spiders' (Araneae, Nephilidae, Nephilinae). Systematic Entomology 32 : 95 135.
81. Labarque FM, Ramírez MJ. 2007. A new species of Drymusa Simon, 1891 (Araneae: Drymusidae) from Chile. Studies on Neotropical Fauna and Environment 42 : 241 246.
82. Ledford JM. 2004. A revision of the spider genus Calileptoneta Platnick (Araneae, Leptonetidae), with notes on morphology, natural history and biogeography. Journal of Arachnology 32 : 231 269.
83. Lehtinen PT. 1996. The ultrastructure of leg skin in the phylogeny of spiders. Revue Suisse de Zoologie volume hors série II : 399 421.
84. Lew SE, Mott DJ. 2005. Mimetidae. In : Ubick D, Paquin P, Cushing PE, Roth V, eds. Spiders of North America: an identification manual. American Arachnological Society, 171 172.
85. Lopardo L, Hormiga G. 2007. On the synaphrid spider Cepheia longiseta (Simon 1881) (Araneae, Synaphridae). American Museum Novitates 3575 : 1 18.
86. Lopardo L, Hormiga G. 2008. Phylogenetic placement of the Tasmanian spider Acrobleps hygrophilus (Araneae, Anapidae) with comments on the evolution of the capture web in Araneoidea. Cladistics 24 : 1 33.
87. Lopardo L, Hormiga G, Melic A. 2007. Spinneret spigot morphology in synaphrid spiders (Araneae, Synaphridae), with comments on the systematics of the family and description of a new species of Synaphris Simon 1894 from Spain. American Museum Novitates 3556 : 1 26.
88. Lopez A, Juberthie-Jupeau L, Stowe MK. 1986. L'appareil séricigène de Kaira alba (Hentz) (Araneae: Araneidae). Mémoires de la Société Royale Belge d'Entomologie 33 : 119 128.
89. Martin D. 1978. Zum Radnetzbau der Gattung Pachygnatha Sund. (Araneae: Tetragnathidae). Mitteilungen aus dem Zoologischen Museum in Berlin 54 : 83 95.
90. Miller JA. 1999. Revision and cladistic analysis of the erigonine spider genus Sisicottus (Araneae, Linyphiidae, Erigoninae). Journal of Arachnology 27 : 553 603.
91. Miller JA. 2005. A redescription of Porrhomma cavernicola Keyserling (Araneae, Linyphiidae) with notes on Appalachian troglobites. Journal of Arachnology 33 : 426 438.
92. Miller JA. 2007. Synaphridae of Madagascar (Araneae: Araneoidea): A new family record for the Afrotropical region. Proceedings of the California Academy of Sciences, Fourth Series 58 : 21 48.
93. Moon M-J, An J-S. 2006. Microstructure of the silk apparatus of the comb-footed spider, Achaearanea tepidariorum (Araneae: Theridiidae). Entomological Research 36 : 56 63.
94. Moon MJ, Baek SB, Kim WK. 1988. Study on the histochemical characteristics and protein patterns of the spider silk glands in Nephila clavata L. Koch. Korean Arachnology 4 : 127 136.
95. Moon MJ, Kim WK. 1990. Ultrastructure of the aciniform glands in Nephila clavata L. Koch (Araneae: Araneidae). Korean Arachnology 5 : 195 206.
96. Mott DJ. 1989. A revision of the genus Mimetus in North America (Araneae, Mimetidae). DPhil Dissertation, Southern Illinois University at Carbondale.
97. Mullen GR. 1969. Morphology and histology of the silk glands in Araneus sericatus Cl. Transactions of the American Microscopical Society 88 : 232 240.
98. Murphy JA. 2007. Gnaphosid genera of the world, Volume 1: text. St Neots, UK : British Arachnological Society.
99. Müller MC, Westheide W. 1993. Comparative morphology of the sexually dimorphic orb-weaving spider Argiope bruennichi (Araneae: Araneidae). Memoirs of the Queensland Museum 33 : 615 620.
100. Palmer JM. 1990. Comparative morphology of the external silk production apparatus of 'primitive' spiders. DPhil Thesis, Harvard University.
101. Park J-G, Lee E-Y, Moon M-J. 2009. Postembryonic development of the silk spinning apparatus in the spiderlings of Araneus ventricosus (Araneae: Araneidae). American Arachnology 78 : 17 (abstract).
102. Peters HM. 1969. Maturing and coordination of web-building activity. American Zoologist 9 : 223 227.
103. Peters HM. 1993. Functional organization of the spinning apparatus of Cyrtophora citricola with regard to the evolution of the web (Araneae, Araneidae). Zoomorphology 113 : 153 163.
104. Peters HM, Kovoor J. 1991. The silk-producing system of Linyphia triangularis (Araneae, Linyphiidae) and some comparisons with Araneidae: Structure, histochemistry and function. Zoomorphology 111 : 1 17.
105. Platnick NI. 1990. Spinneret morphology and the phylogeny of ground spiders (Araneae, Gnaphosoidea). American Museum Novitates 2978 : 1 42.
106. Platnick NI. 1993. A new genus of the spider family Caponiidae (Araneae, Haplogynae) from California. American Museum Novitates 3063 : 1 8.
107. Platnick NI. 1994a. A revision of the spider genus Caponina (Araneae, Caponiidae). American Museum Novitates 3100 : 1 15.
108. Platnick NI. 1994b. A new spider of the genus Archoleptoneta (Araneae, Leptonetidae) from Panama. American Museum Novitates 3101 : 1 8.
109. Platnick NI. 1994c. A review of the Chilean spiders of the family Caponiidae (Araneae, Haplogynae). American Museum Novitates 3113 : 1 10.
110. Platnick NI. 2009. The world spider catalog, Version 9.5. New York, NY : American Museum of Natural History. Available at: http://research.amnh.org/ entomology/spiders/catalog/index.html
111. Platnick NI, Coddington JA, Forster RR, Griswold CE. 1991. Spinneret morphology and the phylogeny of haplogyne spiders (Araneae, Araneomorphae). American Museum Novitates 3016 : 1 73.
112. Platnick NI, Di Franco F. 1992. On the relationships of the spider genus Cybaeodes (Araneae, Dionycha). American Museum Novitates 3053 : 1 9.
113. Platnick NI, Forster RR. 1986. On Teutoniella, an American genus of the spider family Micropholcommatidae (Araneae, Palpimanoidea). American Museum Novitates 2854 : 1 9.
114. Platnick NI, Jäger P. 2008. On the first Asian spiders of the family Caponiidae (Araneae, Haplogynae), with notes on the African genus Diploglena. American Museum Novitates 3634 : 1 12.
115. Platnick NI, Shadab MU. 1993. A review of the pirate spiders (Araneae, Mimetidae) of Chile. American Museum Novitates 3074 : 1 30.
116. Ramírez MJ, Lopardo L, Platnick NI. 2004. Notes on Chilean anapids and their webs. American Museum Novitates 3428 : 1 13.
117. Rix MG, Harvey MS, Roberts JD. 2008. Molecular phylogenetics of the spider family Micropholcommatidae (Arachnida: Araneae) using nuclear rRNA genes (18S and 28S). Molecular Phylogenetics and Evolution 46 : 1031 1048.
118. Scharff N, Coddington JA. 1997. A phylogenetic analysis of the orb-weaving spider family Araneidae (Arachnida, Araneae). Zoological Journal of the Linnean Society 120 : 355 434.
119. Schütt K. 1995. Drapetisca socialis (Araneae: Linyphiidae): Web reduction - ethological and morphological adaptations. European Journal of Entomology 92 : 553 563.
120. Schütt K. 2000. The limits of the Araneoidea (Arachnida: Araneae). Australian Journal of Zoology 48 : 135 153.
121. Schütt K. 2003. Phylogeny of Symphytognathidae s.l. (Araneae, Araneoidea). Zoologica Scripta 32 : 129 151.
122. Selden PA, Huang D, Ren D. 2008. Palpimanoid spiders from the Jurassic of China. Journal of Arachnology 36 : 306 321.
123. Shear WA. 1986. The evolution of web-building behavior in spiders: a third generation of hypotheses. In : Shear WA, ed. Spiders: webs, behavior, and evolution. Stanford, CA : Stanford University Press, 364 400.
124. Shear WA, Palmer JM, Coddington JA, Bonamo PM. 1989. A Devonian spinneret: early evidence of spiders and silk use. Science 246 : 479 481.
125. Thaler K, van Harten A, Knoflach B. 2004. Pirate spiders of the genus Ero C. L. Koch from southern Europe, Yemen, and Ivory Coast, with two new species (Arachnida, Araneae, Mimetidae). Denisia 13 : 359 368.
126. Tillinghast EK. 1984. The chemical fractionation of the orb web of Argiope spiders. Insect Biochemistry 14 : 115 120.
127. Townley MA. 1993. Silk use during proecdysis in the spider Araneus cavaticus (Araneae, Araneidae). DPhil Dissertation, University of New Hampshire.
128. Townley MA, Horner NV, Cherim NA, Tugmon CR, Tillinghast EK. 1991. Selected aspects of spinning apparatus development in Araneus cavaticus (Araneae, Araneidae). Journal of Morphology 208 : 175 191, C5 C7.
129. Townley MA, Tillinghast EK. 2003. On the use of ampullate gland silks by wolf spiders (Araneae, Lycosidae) for attaching the egg sac to the spinnerets and a proposal for defining nubbins and tartipores. Journal of Arachnology 31 : 209 245.
130. Townley MA, Tillinghast EK, Cherim NA. 1993. Moult-related changes in ampullate silk gland morphology and usage in the araneid spider Araneus cavaticus. Philosophical Transactions of the Royal Society London Series B, Biological Sciences 340 : 25 38.
131. Townley MA, Tillinghast EK, Neefus CD. 2006. Changes in composition of spider orb web sticky droplets with starvation and web removal, and synthesis of sticky droplet compounds. Journal of Experimental Biology 209 : 1463 1486.
132. Wiśniewski H. 1986a. Studies on the spinning apparatus of Clubiona phragmitis C. L. Koch (Araneida, Clubionidae). I. The external spinning apparatus. Zoologica Poloniae 33 : 71 81.
133. Wiśniewski H. 1986b. Studies on the spinning apparatus of Clubiona phragmitis C. L. Koch (Araneida, Clubionidae). II. The structure of the internal spinning apparatus. Zoologica Poloniae 33 : 83 104, Plates I V.
134. Work RW. 1981. Web components associated with the major ampullate silk fibers of orb-web-building spiders. Transactions of the American Microscopical Society 100 : 1 20.
135. Wunderlich J. 1986. Spinnenfauna gestern und heute: Fossile Spinnen in Bernstein und ihre heute lebenden Verwandten. Wiesbaden : Erich Bauer Verlag.
136. Wunderlich J. 2004a. On the relationships of the families of the superfamily Araneoidea (Araneae) and their kin, with cladograms, remarks on the origin of the orb web, and description of the new and extinct families Baltsuccinidae and Protheridiidae in Tertiary Baltic amber. Beiträge zur Araneologie 3B : 1112 1154.
137. Wunderlich J. 2004b. Fossil pirate spiders (Araneae: Araneoidea: Mimetidae s. l.) in Baltic and Dominican amber, with notes on intrafamiliar higher taxa. Beiträge zur Araneologie 3B : 1249 1278.
138. Wa̧sowska S. 1969. Particularités de l'appareil séricigène des mâles Clubiona phragmitis C. L. Koch. Zoologica Poloniae 19 : 505 515.
139. Yoshida M. 1996. Spinnerets and spigots of tetragnathid spiders. Memoirs of Institute of Science and Engineering, Ritsumeikan University, Shiga, Japan 55 : 95 106.
140. Yoshida M. 1999. Spinnerets silk spigots morphology of Araneidae, Tetragnathidae, Theridiidae and Linyphiidae (Araneae: Araneoidea). Acta Arachnologica 48 : 1 22.
141. Yu L, Coddington JA. 1990. Ontogenetic changes in the spinning fields of Nuctenea cornuta and Neoscona theisi (Araneae, Araneidae). Journal of Arachnology 18 : 331 345.