Monophyly and phylogenetic placement of the spider genus Labulla Simon, 1884 (Araneae, Linyphiidae) and description of the new genus Pecado

Zoological Journal of the Linnean Society

Volumn 143, Issue 3, 2005, Pages 359-404

  Hormiga, Gustavo ^a   Scharff, Nikolaj ^b  

^a GEORGE WASHINGTON UNIVERSITY   (United States)

^b UNIVERSITY OF COPENHAGEN   (Denmark)

Author keywords

Anatomy; Cladistics; Genitalia; Morphology; Phylogeny; Systematics; Taxonomy

Indexed keywords

EID: 15744383706     PISSN: 00244082     EISSN: None     Source Type: Journal    
DOI: 10.1111/j.1096-3642.2005.00147.x     Document Type: Article

References (76)

1. Becker L. 1896. Les arachnides de Belgique. Annals du Musée Royal de Histoire Naturelle de Belgique 12: 1-378.
2. Blackwall J. 1841. The difference in the number of eyes with which spiders are provided proposed as the basis of their distribution into tribes; with descriptions of newly discovered species and the characters of a new family and three new genera of spiders. Transactions of the Linnean Society of London 18: 601-670.
3. Blackwall J. 1864. A history of the spiders of Great Britain and Ireland. London, Ray Society 2: 175-384.
4. Blest AD. 1976. The tracheal arrangement and the classification of linyphiid spiders. Journal of Zoology (London) 180: 185-194.
5. Bösenberg W. 1901. Die Spinnen Deutschlands. I. Zoologica (Stuttgart) 14: 1-96.
6. Bosmans R, De Keer R,. 1985. Catalogue des Araignées des Pyrenées. Espèces citées, nouvelles récoltes, bibliographie. Documents de Travail, no. 23. Institute Royal de Sciences naturelles de Belgique.
7. Bosmans R, Maelfait JP, Kimpe A. 1986. Analysis of the spider communities in an altitudinal gradient in the French and Spanish Pyrenees. Bulletin of the British Arachnological Society 7: 69-76.
8. Bremer K. 1988. The limits of amino acid sequence data in angiosperm phylogenetic reconstruction. Evolution 42: 795-803.
9. Bremer K. 1995. Branch support and tree stability. Cladistics 10: 295-304.
10. Bristowe WS. 1958. The world of spiders. London: Collins.
11. Castro A. 2004. Estudio biocenológico y faunístico del Orden Araneae (Arthropoda, Arachnida) en los encinares cantábricos de Guipúzcoa y Navarra (España). PhD thesis. Universidad Autónoma de Madrid.
12. Chyzer C, Kulczynski W. 1894. Araneae Hungariae. Budapest 2: 1-151.
13. Coddington JA. 1983. A temporary slide mount allowing precise manipulation of small structures. In: Kraus O, ed., Taxonomy, biology, and ecology of Araneae and Myriapoda. Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg 26: 291-292.
14. Denis J. 1941. Les araignées des îles Canaries. Annales de la Société Entomologique de France 110: 105-130.
15. Denis J. 1945. Descriptions d'araignées nord-africaines. Bulletin de la Société d'Histoire Naturelle de Toulouse 79: 41-57.
16. Denis J. 1949. Notes sur les érigonides. XVI. Essai sur la détermination des femelles d'érigonides. Bulletin de la Société d'Histoire Naturelle de Toulouse 83: 129-158.
17. Donoghue MJ, Olmstead RG, Smith JF, Palmer JD. 1992. Phylogenetic relationships of Dipsacales based on rbcL sequence data. Annals of the Missouri Botanical Garden 79: 333-345.
18. Fage L. 1935. Description du male de Metella breuili Fage (Araneida) accompagnee de remarques sur la position systematique du genre Metella. Revista Española de Entomología 11: 177-180.
19. Fitch WM. 1971. Toward defining the course of evolution: minimum change for a specific tree topology. Systematic Zoology 20: 406-416.
20. Foelix RF. 1996. Biology of spiders, 2nd edn. Oxford: Oxford University Press.
21. Förster A, Bertkau P. 1883. Beiträge zur Kenntniss der Spinnenfauna der Rheinprovinz. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens 40: 205-278.
22. Gerhardt U. 1921. Vergleichende Studien über die Morphologie des männlichen Tasters und die Biologie der Kopulation der Spinnen. Archiv für Naturgeschichte 87: 78-247.
23. Gerhardt U. 1923. Weitere sexualbiologische Untersuchungen an Spinnen. Archiv für Naturgeschichte 89: 1-225, pl. 1-3.
24. Gerhardt U. 1925. Neue Sexualbiologische Spinnenstudien. Zeitschrift für Morphologie und Ökologie der Tiere 3: 567-618.
25. Gerhardt U. 1928. Biologische Studien an griechischen, corsischen und deutschen Spinnen. Zeitschrift für Morphologie und Ökologie der Tiere 10: 576-675.
26. Goloboff PA. 1993a. NONA, Version 2.0. Program available at http://www.cladistics.com
27. Goloboff PA. 1993b. Pee-Wee, Version 3.0. Program available at http://www.cladistics.com
28. Goloboff PA. 1993c. Estimating character weights during tree search. Cladistics 9: 83-91.
29. Grandjean F. 1949. Observation et conservation des très petits arthropodes. Bulletin du Muséum National d'Histoire Naturelle 21: 363-370.
30. Griswold CE, Coddington JA, Hormiga G, Scharff N. 1998. Phylogeny of the orb-web building spiders (Araneae, Orbiculariae: Deinopoidea, Araneoidea). Zoological Journal of the Linnean Society 123: 1-99.
31. Harvey PR, Nellist DR, Telfer MG. 2002. Provisional atlas of British spiders (Arachnida, Araneae), Vols 1 & 2. Huntingdon: Biological Records Centre.
32. Heimer S, Nentwig W. 1991. Spinnen Mitteleuropas: ein Bestimmungsbuch. Berlin: Verlag Paul Parey.
33. van Helsdingen PJ. 1968. Comparative notes on the species of the Holarctic genus Stemonyphantes Menge (Araneida, Linyphiidae). Zoologische Mededelingen 43: 117-139.
34. van Helsdingen PJ. 1969. A reclassification of the species of Linyphia Latreille, based on the functioning of the genitalia (Araneida, Linyphiidae) I. Zoologische Verhandelingen (Leiden) 105: 1-303.
35. van Helsdingen PJ. 1973. Annotations on two species of linyphiid spiders by the late Wilton Ivie. Psyche 80: 48-61.
36. Holm Å. 1979. A taxonomic study of European and East African species of the genera Pelecopsis and Trichopterna (Araneae, Linyphiidae), with descriptions of a new genus and two new species of Pelecopsis from Kenya. Zoologica Scripta 8: 255-278.
37. Hormiga G. 1994a. A revision and cladistic analysis of the spider family Pimoidae (Araneoidea, Araneae). Smithsonian Contributions to Zoology 549: 1-104.
38. Hormiga G. 1994b. Cladistics and the comparative morphology of linyphiid spiders and their relatives (Araneae, Araneoidea, Linyphiidae). Zoological Journal of the Linnean Society 111: 1-71.
39. Hormiga G. 2000. Higher level phylogenetics of erigonine spiders (Araneae, Linyphiidae, Erigoninae). Smithsonian Contributions to Zoology 609: 1-160.
40. Hormiga G. 2002. Orsonwelles, a new genus of giant linyphiid spiders (Araneae) from the Hawaiian islands. Invertebrate Systematics 16: 1-80.
41. Hormiga G. 2003. Weintrauboa, a new genus of pimoid spiders from Japan and adjacent islands, with comments on the monophyly and diagnosis of the family Pimoidae and the genus Pimoa (Araneoidea, Araneae). Zoological Journal of the Linnean Society 139: 261-281.
42. Hormiga G, Arnedo MA, Gillespie RG. 2003. Speciation on a conveyor belt: sequential colonization of the Hawaiian Islands by Orsonwelles spiders (Araneae, Linyphiidae). Systematic Biology 52: 70-88.
43. Hormiga G, Eberhard WG, Coddington JA. 1995. Webconstruction behaviour in Australian Phonognatha and the phylogeny of nephiline and tetragnathid spiders (Araneae: Tetragnathidae). Australian Journal of Zoology 43: 313-364.
44. Huber BA. 1998. Spider reproductive behaviour: a review of Gerhardt's work from 1911 to 1933, with implications for sexual selection. Bulletin of the British Arachnological Society 11: 81-91.
45. Ledoux JC, Emerit M, Pinault G. 1996. Les araignées et opilions de Nohèdes (Pyrénées Orientales). Office por l'Information Eco-Entomologique du Languedoc-Roussillon.
46. Locket GH, Millidge AF. 1953. British spiders, Vol. 2. London: Ray Society.
47. Maddison DR, Maddison WP. 2001. Macclade 4: analysis of phylogeny and character evolution, Version 4.03. Sunderland, MA: Sinauer Associates.
48. Marusik YM, Eskov KY, Logunov DV, Basarukin AM. 1993. A check-list of spiders (Arachnida Aranei) from Sakhalin and Kurile Islands. Arthropoda Selecta 1: 73-85.
49. Merrett P. 1963. The palpus of male spiders of the family Linyphiidae. Proceedings of the Zoological Society of London 140: 347-467.
50. Miller F. 1971. Pavouci-Araneida. Klíc Zvíreny CSSR 4: 51-306.
51. Millidge AF. 1977. The conformation of the male palpal organs of Linyphiid spiders and its application to the taxonomic and phylogenetic analysis of the family (Araneae: Linyphiidae). Bulletin of the British Arachnological Society 4: 1-60.
52. Millidge AF. 1984. The taxonomy of the Linyphiidae, based chiefly on the epigynal and tracheal characters (Araneae: Linyphiidae). Bulletin of the British Arachnological Society 6: 229-267.
53. Millidge AF. 1993. Further remarks on the taxonomy and relationships of the Linyphiidae, based on the epigynal duct confirmations and other characters (Araneae). Bulletin of the British Arachnological Society 9: 145-156.
54. Nielsen E. 1928. De Danske Edderkoppers Biologi. Copenhagen: Levin & Munksgaards Forlag.
55. Nixon K. 1999. Winclada, Version 1.00.08. Program available at http://www.cladistics.com
56. Page RDM. 2001. Nexus Data Editor, Version 0.4.9. Available at http://taxonomy.zoology.gla.ac.uk/rod/NDE/nde.html
57. Palmgren P. 1975. Die Spinnenfauna Finnlands und Ostfennoskandiens VI: Linyphiidae 1. Fauna Fennica 28: 1-102.
58. Platnick NI. 2004. The world spider catalog, Version 4.5. American Museum of Natural History, online at http://research.amnh.org/entomology/spiders/ catalog/index.html
59. Ribera C, Hormiga G. 1985. Artrópodos epigeos del macizo de San Juan de la Peña (Jaca, Prov. Huesca). XI. Arañas linífidas. Pirineos 126: 163-209.
60. Roberts MJ. 1987. The spiders of Great Britain and Ireland, Vol. 2: Linyphiidae and Check List. Colchester: Harley Books.
61. Roberts MJ. 1995. Collins field guide: spiders of Britain and northern Europe. London: Harper Collins.
62. Roberts MJ. 1998. Spinnengids. Baarn, the Netherlands: Tirion.
63. Saito S. 1935. Further notes on spiders from southern Saghalin, with descriptions of three new species. Annotationes Zoologicae Japonenses 15: 58-61.
64. Scharff N, Coddington JA, Griswold CE, Hormiga G, Bjørn P, de Place. 2003. When to quit? Estimating spider species richness in a northern European deciduous forest. Journal of Arachnology 31: 246-273.
65. Simon E. 1884. Les arachnides de France. Tome 5(2). Paris: Libraire Encyclopédique de Roret.
66. Simon E. 1894. Histoire naturelle des araignées. Tome 1. Paris: Libraire Encyclopédique de Roret.
67. Simon E. 1900. Arachnida. Fauna Hawaiiensis 2: 443-519.
68. Simon E. 1914. Descriptions de plusiers espèces d'arachnides récemment découvertes en France. Bulletin de la Société Entomologique de France 1914: 478-479.
69. Simon E. 1929. Les arachnides de France. Synopsis générale et catalogue des espèces françaises de l'ordre des Araneae; 3e partie. Tome 6. Paris: Libraire Encyclopédique de Roret.
70. Swofford DL. 2001. PAUP*: phylogenetic analysis using parsimony (* and other methods), Version 4.0b 7. Sunderland, MA: Sinauer Associates.
71. Tanasevitch AV, Eskov KY. 1987. Spiders of the genus Lepthyphantes (Aranei, Linyphiidae) in the Siberian and Far-Eastern fauna. Zoologiske Zhurnal 66: 185-197.
72. Tikader BK. 1970. Spider fauna of Sikkim. Records of the Zoological Survey of India 64: 1-83.
73. Townley MA, Tillinghast EK. 2003. On the use of ampullate gland silks by wolf spiders (araneae, lycosidae) for attaching the egg sac to the spinnerets and a proposal for defining nubbins and tartipores. Jounal of Arachnology 31: 209-245.
74. Wider. 1834. Beschreibung der Arachniden. In: Reuss A, ed. Zoologische Miscellen. Museum Senckenbergianum (Abhandlungen) 1: 195-282.
75. Wiehle H. 1952. Eine übersehene deutsche Theridion-Art. Zoolgischer Anzeiger 149: 226-235.
76. Wiehle H. 1956. Spinnentiere oder Arachnoidea (Araneae). 28. Familie Linyphiidae-Baldachinspinnen. Tierwelt Deutschlands 44: i-viii, 1-337.