Anatomy and relationships of the middle permian varanopid Heleosaurus Scholtzi based on a social aggregation from the karoo basin of South Africa

Journal of Vertebrate Paleontology

Volumn 29, Issue 2, 2009, Pages 389-400

  Botha Brink, Jennifer ^a,b   Modesto, Sean P ^c  

^a NATIONAL MUSEUM   (South Africa)

^b UNIVERSITY OF THE FREE STATE   (South Africa)

^c CAPE BRETON UNIVERSITY   (Canada)

Author keywords

[No Author keywords available]

Indexed keywords

AGGREGATION BEHAVIOR; ANATOMY; CLADISTICS; DINOSAUR; PALEOECOLOGY; PALEONTOLOGY; PERMIAN; PHYLOGENY; SKELETAL REMAINS; SOCIAL BEHAVIOR; TAXONOMY;

AFRICA; KAROO; SOUTH AFRICA; SOUTHERN AFRICA; SUB-SAHARAN AFRICA;

EID: 67651096009     PISSN: 02724634     EISSN: None     Source Type: Journal    
DOI: 10.1671/039.029.0209     Document Type: Article

References (25)

1. Anderson, J. S., and R. R. Reisz. 2004. Pyozia mesenensis, a new, small varanopid (Synapsida, Eupelycosauria) from Russia: "pelycosaur" diversity in the Middle Permian. Journal of Vertebrate Paleontology 24:173-179.
2. Berman, D. S., and R. R. Reisz. 1982. Restudy of Mycterosaurus long- iceps (Reptilia, Pelycosauria) from the Lower Permian of Texas. Annals of Carnegie Museum 51:423-453.
3. Botha-Brink, J., and S. P. Modesto. 2007. A mixed-age classed 'pelycosaur' aggregation from South Africa: earliest evidence of parental care in amniotes? Proceedings of the Royal Society B 274: 2829-2834.
4. Broom, R. 1907. On some new fossil reptiles from the Karroo beds of Victoria West, South Africa. Transactions of the South African Philosophical Society 18:31-42.
5. Broom, R. 1937. A further contribution to our knowledge of the fossil reptiles of the Karroo. Proceedings of the Zoological Society of London 1937:299-318. Carroll, R. L. 1976.
6. Eosuchians and the origin of archosaurs; pp. 58-79 in C. S. Churcher (ed.), Essays on Palaeontology in Honour of Loris Shano Russell. Royal Ontario Museum, Toronto.
7. Dias-da-Silva, S., S. P. Modesto, and C. L. Schultz. 2007. New material of Procolophon (Parareptilia: Procolophonoidea) from the Lower Tri- assic of Brazil, with remarks on the ages of the Sanga do Cabral and Buena Vista formations of South America. Canadian Journal of Earth Sciences 43:1685-1693.
8. Dilkes, D. W., and R. R. Reisz. 1996. First record of a basal synapsid ('mammal-like reptile') in Gondwana. Proceedings of the Royal Society of London, Series B 263:1165-1170.
9. Gauthier, J. A., A. G. Kluge, and T. Rowe. 1988. Amniote phylogeny and the importance of fossils. Cladistics 4:105-209.
10. Kemp, T. S. 1982. Mammal-like Reptiles and the Origin of Mammals. Academic Press, London, 363 pp.
11. Kitching, J. W. 1977. The distribution of the Karroo vertebrate fauna. Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, Memoir 1:1-131.
12. Langston, J. W., and R. R. Reisz. 1981. Aerosaurus wellesi, new species, a varanopseid mammal-like reptile (Synapsida: Pelycosauria) from the Lower Permian of New Mexico. Journal of Vertebrate Paleontology 1:73-96.
13. Lucas, S. G. 2006. Global Permian tetrapod biostratigraphy and biochro- nology; pp. 65-93 in S. G. Lucas, G. Cassinis, and J. W. Schneider (eds.), Non-Marine Permian Biostratigraphy and Biochronology. Geological Society of London, Special Publication 265.
14. Maddin, H. C, D. C. Evans, and R. R. Reisz. 2006. An Early Permian varanodontine varanopid (Synapsida: Eupelycosauria) from the Richards Spur Locality, Oklahoma. Journal of Vertebrate Paleontology 26:957-966.
15. Modesto, S. P., C. A. Sidor, B. S. Rubidge, and J. Weiman. 2001. A second varanopseid skull from the Upper Permian of South Africa: implications for Late Permian 'pelycosaur' evolution. Lethaia 34:249-259.
16. Osborn, H. F. 1903. On the primary division of the Reptilia into two subclasses, Synapsida and Diapsida. Science 17:275-276.
17. Pineiro, G., M. Verde, M. Ubilla, and J. Ferigolo. 2003. First basal synapsids ("pelycosaurs") from the Upper Permian-?Lower Triassic of Uruguay, South America. Journal of Paleontology 77:389-392.
18. Reisz, R. R., and D. S Berman. 2001. The skull of Mesenosaurus romeri, a small varanopseid (Synapsida: Eupelycosauria) from the Upper Permian of the Mezen River Basin, northern Russia. Annals of Carnegie Museum 70:113-132.
19. Reisz, R. R., and D. W. Dükes. 2003. Archaeovenator hamiltonensis, a new varanopid (Synapsida: Eupelycosauria) from the Upper Carboniferous of Kansas. Canadian Journal of Earth Sciences 40:667-678.
20. Reisz, R. R., and M. Laurin. 2004. A reevaluation of the enigmatic Permian synapsid Watongia and of its stratigraphic significance. Canadian Journal of Earth Sciences 41:377-386.
21. Reisz, R. R., and S. P. Modesto. 2007. Heleosaunis scholtzi from the Permian of South Africa: a varanopid synapsid, not a diapsid reptile. Journal of Vertebrate Paleontology 27:734-739.
22. Reisz, R. R., D. W. Dükes, and D. S Berman. 1998. Anatomy and relationships of Elliotsmithia longiceps Broom, a small synapsid (Eupelycosauria: Varanopseidae) from the Late Permian of South Africa. Journal of Vertebrate Paleontology 18:602-611.
23. Romer, A. S., and L. I. Price. 1940. Review of the Pelycosauria. Geological Society of America, Special Papers 40:1-538.
24. Smith, R. M. H., and A. W. Keyser. 1995. Biostratigraphy of the Tapino- cephalus Assemblage Zone; pp. 8-12 in B. S. Rubidge (ed.), Biostratigraphy of the Beaufort Group (Karoo Supergroup). South African Committee for StratigraphyBiostratigraphic Series 1.
25. Swofford, D. L. 2002. PAUP*: Phylogenetic analysis Using Parsimony (*and Other Methods). Version 4.0b8. Sinauer Associates: Sunderland, Massachusetts.