Taphonomic perspectives on hominid site use and foraging strategies during Bed II times at Olduvai Gorge, Tanzania

Journal of Human Evolution

Volumn 55, Issue 6, 2008, Pages 1031-1052

  Egeland, Charles P ^a,b   Domínguez Rodrigo, Manuel ^c  

^a UNIVERSITY OF WISCONSIN MADISON   (United States)

^b Wisconsin Historical Society   (United States)

^c UNIVERSIDAD COMPLUTENSE DE MADRID   (Spain)

Author keywords

Bone surface modifications; Breakage patterns; Cutmarks; Taphonomy; Tooth marks

Indexed keywords

FORAGING BEHAVIOR; HOMINID; STONE AGE; TAPHONOMY; TOOL USE;

AFRICA; ARUSHA [TANZANIA]; EAST AFRICA; OLDUVAI GORGE; SUB-SAHARAN AFRICA; TANZANIA;

HOMINIDAE;

EID: 55949086529     PISSN: 00472484     EISSN: None     Source Type: Journal    
DOI: 10.1016/j.jhevol.2008.05.021     Document Type: Article

References (156)

1. Albert R.M., Bamford M.K., and Cabanes D. Taphonomy of phytoliths and macroplants in different soils from Olduvai Gorge (Tanzania) and the application to Plio-Pleistocene paleoanthropological samples. Quatern. Int 148 (2006) 78-94
2. Alcántara García V., Barba R., Barral del Pino J.M., Crespo Ruiz A.B., Eiriz Vidal A.I., Falquina Aparicio A., Herrero Calleja S., Ibara Jiménez A., Megías González M., Pérez Gil M., Pérez Tello V., Rolland Calvo J., Yravedra Sáinz de los Torreros J., Vidal A., and Domínguez-Rodrigo M. Determinación de procesos de fractura sobre huesos frescos: un sistema de anáiysis de los ángulos de los planos de fracturación como discriminar de agentes bióticos. Trabajos de Prehistoria 63 (2006) 25-38
3. Andrews P., and Fernández-Jalvo Y. Surface modifications of the Sima de los Huesos fossil humans. J. Hum. Evol. 33 (1997) 191-217
4. Antón S.C. Natural history of Homo erectus. Yearb. Phys. Anthropol. 46 (2003) 126-169
5. Assefa Z. Faunal remains from Porc-Epic: paleoecological and zooarchaeological investigations from a Middle Stone Age site in southeastern Ethiopia. J. Hum. Evol. 51 (2006) 50-75
6. Bartram L.E., and Marean C.W. Explaining the "Klaises Pattern": Kua ethnoarchaeology, the Die Kelders Middle Stone Age archaeofauna, long bone fragmentation and carnivore ravaging. J. Archaeol. Sci. 26 (1999) 9-29
7. Bamford M.K., Albert R.M., and Cabanes D. Plio-Pleistocene macroplant fossil remains and phytoliths from lowermost Bed II in the eastern paleolake margin of Olduvai Gorge, Tanzania. Quatern. Int 148 (2006) 95-112
8. Behrensmeyer A.K. Taphonomic and ecologic information from bone weathering. Paleobiology 4 (1978) 150-162
9. Behrensmeyer A.K. Time sampling intervals in the vertebrate fossil record. Proc. Third N. Am. Paleon. Convention 1 (1982) 41-45
10. Behrensmeyer A.K. Patterns of natural bone distribution on recent land surfaces: implications for archaeological site formation. In: Clutton-Brock J., and Grigson C. (Eds). Animals and Archaeology I: Hunters and Their Prey. British Archaeological Reports International Series (1983) Oxford, pp. 93-106
11. Behrensmeyer A.K. Taphonomy and hunting. In: Nitecki M.H., and Nitecki D.V. (Eds). The Evolution of Human Hunting (1987), Plenum Press, New York 423-450
12. Behrensmeyer A.K., and Dechant Boaz D.E. The recent bones of Amboseli Park, Kenya, in relation to East African paleoecology. In: Behrensmeyer A.K., and Hill A. (Eds). Fossils in the Making (1980), University of Chicago Press, Chicago 72-92
13. Behrensmeyer A.K., Gordon K.D., and Yanagi G.T. Trampling as a cause of bone surface damage and psuedo-cutmarks. Nature 319 (1986) 768-771
14. Binford L.R. Nunamiut Ethnoarchaeology (1978), Academic Press, New York
15. Binford L.R. Bones: Ancient Men and Modern Myths (1981), Academic Press, New York
16. Binford L.R. In Pursuit of the Past: Decoding the Archaeological Record (1983), Thames and Hudson, New York
17. Binford L.R., and Bertram J. Bone frequencies-and attritional processes. In: Binford L.R. (Ed). For Theory Building in Archaeology (1977), Academic Press, New York 77-153
18. Blumenschine, R.J., 1986. Early Hominid Scavenging Opportunities: Implications of Carcass Availability in the Serengeti and Ngorongoro Ecosystems. British Archaeological Reports International Series Oxford.
19. Blumenschine R.J. An experimental model of the timing of hominid and carnivore influence on archaeological bone assemblages. J. Archaeol. Sci. 15 (1988) 483-502
20. Blumenschine R.J. A landscape taphonomic model of the scale of prehistoric scavenging opportunities. J. Hum. Evol. 18 (1989) 345-371
21. Blumenschine R.J. Percussion marks, tooth marks, and experimental determination of the timing of hominid and carnivore access to long bones at FLK Zinjanthropus, Olduvai Gorge, Tanzania. J. Hum. Evol. 29 (1995) 21-51
22. Blumenschine R.J., and Masao F.T. Living sites at Olduvai Gorge, Tanzania? Preliminary landscape archaeology results in the basal Bed II lake margin zone. J. Hum. Evol. 21 (1991) 451-462
23. Blumenschine R.J., and Selvaggio M.M. Percussion marks on bone surfaces as a new diagnostic of hominid behaviour. Nature 333 (1988) 763-765
24. Blumenschine R.J., and Selvaggio M.M. On the marks of marrow bone processing by hammerstones and hyenas: their anatomical patterning and archaeological implications. In: Clarke J.D. (Ed). Cultural Beginnings: Approaches to Understanding Early Hominid Life-Ways in the African Savanna (1991), Dr. Rudolf Habelt GMBH, Bonn 17-32
25. Blumenschine R.J., and Madrigal T.C. Variability in long bone marrow yields of East African ungulates and its zooarchaeological implications. J. Archaeol. Sci. 20 (1993) 555-587
26. Blumenschine R.J., Marean C.W., and Capaldo S.D. Blind tests of inter-analyst correspondence and accuracy in the identification of cut marks, percussion marks, and carnivore tooth marks on bone surfaces. J. Archaeol. Sci. 23 (1996) 493-507
27. Blumenschine R.J., Peters C.R., Masao F.T., Clarke R.J., Deino A.L., Hay R.L., Swisher III C.C., Stanistreet I.G., Ashely G.M., McHenry L.J., Sikes N.E., Van der Merwe N.J., Tactikos J.C., Cushing A.E., Deocampo D.M., Njau J.K., and Ebert J.I. Late Pliocene Homo and hominid land use from western Olduvai Gorge, Tanzania. Science 299 (2003) 1217-1221
28. Brain C.K. Hottentot food remains and their bearing on the interpretation of fossil bone assemblages. Scien. Pap. Namib Desert Res. Stat. 32 (1967) 1-7
29. Brain C.K. The contribution of Namib Desert Hottentots to an understanding of australopithecine bone accumulations. Scien. Pap. Namib Desert Res. Stat. 39 (1969) 13-22
30. Brain C.K. Some suggested procedures in the analysis of bone accumulations from southern African Quaternary sites. Ann. Transvaal Mus. 29 (1974) 1-8
31. Brain C.K. The Hunters or the Hunted? An Introduction to African Cave Taphonomy (1981), University of Chicago Press, Chicago
32. Bunn H.T. Archaeological evidence for meat-eating by Plio-Pleistocene hominids from Koobi Fora and Olduvai Gorge. Nature 291 (1981) 574-577
33. Bunn, H.T., 1982. Meat-eating and human evolution: Studies on the diet and subsistence patterns of Plio-Pleistocene hominids in East Africa. Ph.D. Dissertation, University of California, Berkeley.
34. Bunn H.T. Comparative analysis of modern bone assemblages from a San hunter-gatherer camp in the Kalahari Desert, Botswana, and from a spotted hyena den near Nairobi, Kenya. In: Clutton-Brock J., and Grigson C. (Eds). Animals and Archaeology I: Hunters and Their Prey. British Archaeological Reports International Series (1983) Oxford, 21--30.
35. Bunn H.T. Patterns of skeletal part representation and hominid subsistence activities at Olduvai Gorge, Tanzania, and Koobi Fora, Kenya. J. Hum. Evol. 15 (1986) 673-690
36. Bunn H.T. Diagnosing Plio-Pleistocene hominid activity with bone fracture evidence. In: Bonnichsen R., and Sorg M.H. (Eds). Bone Modification (1989), Center for the Study of the First Americans, Orono 299-315
37. Bunn H.T. Early Pleistocene hominid foraging strategies along the ancestral Omo River at Koobi Fora, Kenya. J. Hum. Evol. 27 (1994) 247-266
38. Bunn H.T. Hunting, power scavenging, and butchering by Hadza foragers and by Plio-Pleistocene Homo. In: Stanford C.B., and Bunn H.T. (Eds). Meat-Eating and Human Evolution (2001), Oxford University Press, Oxford 199-218
39. Bunn H.T. Meat made us human. In: Ungar P.S. (Ed). Evolution of the Human Diet: The Known, the Unknown, and the Unknowable (2006), Oxford University Press, Oxford 191-211
40. Bunn H.T. Butchering backstraps and bearing backbones: insights from Hadza foragers and implications for Paleolithic archaeology. In: Pickering T.R., Schick K., and Toth N. (Eds). Breathing Life into Fossils: Taphonomic Studies in Honor of C.K. (Bob) Brain (2007), Stone Age Institute Press, Bloomington 269-280
41. Bunn H.T., and Kroll E.M. Systematic butchery by Plio Pleistocene hominids at Olduvai-Gorge, Tanzania. Curr. Anthropol. 27 (1986) 431-452
42. Bunn H.T., Kroll E.M., and Bartram L.E. Bone distribution on a modern East African landscape and its archaeological implications. In: Clark J.D. (Ed). Cultural Beginnings: Approaches to Understanding Early Hominid Life-Ways in the African Savanna (1991), Dr. Rudolf Habelt GMBH, Bonn 33-54
43. Bunn H.T., Kroll E.M., Kaufulu Z.M., and Isaac G.L. FxJj 50. In: Isaac G.L. (Ed). Koobi Fora Research Project, vol. 5: Plio-Pleistocene Archaeology (1997), Oxford University Press, Oxford 192-211
44. Bunn H.T., Harris J.W.K., Isaac G.L., Kaufulu Z., Kroll E.M., Schick K., Toth N., and Behrensmeyer A.K. FxJj50: an early Pleistocene site in northern Kenya. World Archaeol. 12 (1980) 109-139
45. Capaldo, S.D., 1995. Inferring hominid and carnivore behavior from dual-patterned archaeofaunal assemblages. Ph.D. Dissertation, Rutgers University, New Brunswick.
46. Capaldo S.D. Experimental determinations of carcass processing by Plio-Pleistocene hominids and carnivores at FLK 22 (Zinjanthropus), Olduvai Gorge, Tanzania. J. Hum. Evol. 33 (1997) 555-597
47. Capaldo S.D. Simulating the formation of dual-patterned archaeofaunal assemblages with experimental control samples. J. Archaeol. Sci. 25 (1998) 311-330
48. Capaldo S.D., and Blumenschine R.J. A quantitative diagnosis of notches made by hammerstone percussion and carnivore gnawing on bovid long bones. Am. Antiq. 59 (1994) 724-748
49. Cerling T.E., and Hay R.L. An isotopic study of paleosol carbonates from Olduvai Gorge. Quatern. Res. 25 (1986) 63-78
50. Cleghorn N., and Marean C.W. Distinguishing selective transport and in situ attrition: a critical review of analytical approaches. J. Taphonomy 2 (2004) 43-67
51. Cruz-Uribe K. Distinguishing hyena from hominid bone accumulations. J. Field Archaeol. 18 (1991) 467-486
52. Cushing, A.E., 2002. The landscape zooarchaeology and paleontology of Plio-Pleistocene Olduvai, Tanzania and their implications for early hominid ecology. Ph.D. Dissertation, Rutgers University, New Brunswick.
53. Domínguez-Rodrigo M. La analítica espacial micro como herramienta tafonomica: aplicación al entorno de una charca africana. Procesos postdeposicionales. Arquelologia Espacial 16-17 (1993) 41-56
54. Domínguez-Rodrigo M. A landscape study of bone preservation in the Galana and Kulalu (Kenya) ecosystem. Origini 20 (1996) 17-38
55. Domínguez-Rodrigo M. Meat-eating by early hominids at the FLK 22 Zinjanthropus site, Olduvai Gorge (Tanzania): an experimental approach using cut-mark data. J. Hum. Evol. 33 (1997) 669-690
56. Domínguez-Rodrigo M. Flesh availability and bone modifications in carcasses consumed by lions: palaeoecological relevance in hominid foraging patterns. Palaeogeogr. Palaeoclimatol. Palaeoecol. 149 (1999) 373-388
57. Domínguez-Rodrigo M. Meat-eating and carcass procurement at the FLK Zinj 22 site, Olduvai Gorge (Tanzania): a new experimental approach to the old hunting-versus-scavenging debate. In: Ullrich H. (Ed). Lifestyles and Survival Strategies in Pliocene and Pleistocene Hominids (1999), Edition Archaea, Schwelm 89-111
58. Domínguez-Rodrigo M. Hunting and scavenging by early humans: the state of the debate. World Archaeol. 16 (2002) 1-54
59. Domínguez-Rodrigo M., and Pickering T.R. Early hominid hunting and scavenging: a zooarchaeological review. Evol. Anthropol. 12 (2003) 275-282
60. Domínguez-Rodrigo M., and Piqueras A. The use of tooth pits to identify carnivore taxa in tooth-marked archaeofaunas and their relevance to reconstruct hominid carcass processing behaviours. J. Archaeol. Sci. 30 (2003) 1385-1391
61. Domínguez-Rodrigo M., and Barba R. A study of cut marks on small-sized carcasses and its application to the study of cut-marked bones from small mammals at the FLK Zinj site. J. Taphonomy 3 (2005) 121-134
62. Domínguez-Rodrigo M., and Barba R. The behavioral meaning of cutmarks at FLK Level 22 Zinjanthropus: the carnivore-hominid-carnivore hypothesis falsified (II). In: Domínguez-Rodrigo M., Barba R., and Egeland C.P. (Eds). Deconstructing Olduvai: A Taphonomic Study of the Bed I Sites (2007), Springer Verlag, New York 75-100
63. In: Domínguez-Rodrigo M., Barba R., and Egeland C.P. (Eds). Deconstructing Olduvai: A Taphonomic Study of the Bed I Sites (2007), Springer Verlag, New York
64. Domínguez-Rodrigo M., Egeland C.P., and Pickering T.R. Equifinality in carnivore tooth marks and the extended concept of archaeological palimpsests: implications for models of passive scavenging in early hominids. In: Pickering T.R., Schick K., and Toth N. (Eds). Breathing Life into Fossils: Taphonomic Studies in Honor of C.K. (Bob) Brain (2007), Stone Age Institute Press, Bloomington 255-268
65. Domínguez-Rodrigo M., de la Torre I., de Luque L., Alcalá L., Serrallonga J., and Medina V. The ST Site Complex at Peninj, West Lake Natron, Tanzania: implications for early hominid behavioural models. J. Archaeol. Sci. 29 (2002) 639-665
66. Egeland, C.P., 2007. Zooarchaeological and taphonomic perspectives on hominid and carnivore interactions at Olduvai Gorge, Tanzania. Ph.D. Dissertation, Indiana University, Bloomington.
67. Egeland C.P., Pickering T.R., Dominguez-Rodrigo M., and Brain C.K. Disentangling Early Stone Age palimpsests: determining the functional independence of hominid- and carnivore-derived portions of archaeofaunas. J. Hum. Evol. 47 (2004) 343-357
68. Faith J.T., and Behrensmeyer A.K. Changing patterns of carnivore modification in a landscape bone assemblage, Amboseli Park, Kenya. J. Archaeol. Sci. 33 (2006) 1718-1733
69. Faith J.T., and Gordon A.D. Skeletal element abundances in archaeofaunal assemblages: economic utility, sample size, and assessment of carcass transport strategies. J. Archaeol. Sci. 34 (2007) 872-882
70. Faith J.T., Marean C.W., and Behrensmeyer A.K. Carnivore competition, bone destruction, and bone density. J. Archaeol. Sci. 34 (2007) 2025-2034
71. Fiorillo A. An experimental study of trampling: implications for the fossil record. In: Bonnichsen R., and Sorg M.H. (Eds). Bone Modification (1989), Center for the Study of the First Americans, Orono 61-71
72. Foley R. Off-site archaeology: an alternative approach for the short-sited. In: Hodder I., Isaac G.L., and Hammond N. (Eds). Pattern of the Past: Studies in Honour of David Clarke (1981), Cambridge University Press, Cambridge 157-183
73. Hay R.L. Geology of the Olduvai Gorge (1976), University of California Press, Berkeley
74. Haynes G. On watering holes, mineral licks, death, and predation. In: Mead J.I., and Meltzer D.J. (Eds). Environments and Extinction: Man in Late Glacial North America (1985), Center for the Study of Early Man, Orono 53-71
75. Haynes G. Mass deaths and serial predation: comparative taphonomic studies of modern large mammal death sites. J. Archaeol. Sci. 15 (1988) 219-235
76. Henschel J.R., Tilson R., and von Blottnitz F. Implications of a spotted hyaena bone assemblage in the Namib Desert. S. Afr. Archaeol. Bull. 34 (1979) 127-131
77. Hill A. Bone modification by modern spotted hyenas. In: Bonnichsen R., and Sorg M.H. (Eds). Bone Modification (1989), Center for the Study of the First Americans, Orono 169-178
78. Isaac G.L. Archaeological tests of alternative models of early hominid behaviour: excavation and experiments. Phil. Trans. Roy. Soc. Lond. 292B (1981) 177-188
79. Isaac G.L. Bones in contention: competing explanations for the juxtaposition of early Pleistocene artefacts and faunal remains. In: Clutton-Brock J., and Grigson C. (Eds). Animals and Archaeology I: Hunters and Their Prey. British Archaeological Reports International Series (1983) Oxford, pp. 3-19
80. Isaac G.L., and Crader D.C. To what extent were early hominids carnivorous? An archaeological perspective. In: Harding R.S.O., and Teleki G. (Eds). Omnivorous Primates: Gathering and Hunting in Human Evolution (1981), Columbia University Press, New York 37-108
81. Isaac G.L., Harris J.W.K., and Marshall F. Small is informative: the application of the study of mini-sites and least effort criteria in the interpretation of the early Pleistocene archaeological record at Koobi Fora, Kenya. In: Clark J.D., and Isaac G.L. (Eds). Las Industrias Mas Antiguas (1981), X Congreso Union Internacional de Ciencias Prehistoricas y Protohistoricas, Mexico City 101-119
82. Johnson E. Current developments in bone technology. Adv. Archaeol. Meth. Theory 8 (1985) 157-235
83. Kappelman J. Plio-Pleistocene marine-continental correlation using habitat indicators from Olduvai Gorge, Tanzania. Quatern. Res. 25 (1986) 141-149
84. Kimura Y. Tool-using strategies by early hominids at Bed II, Olduvai Gorge, Tanzania. J. Hum. Evol. 37 (1999) 807-831
85. Kimura Y. Examining time trends in the Oldowan technology at Beds I and II, Olduvai Gorge. J. Hum. Evol. 43 (2002) 291-321
86. Lam Y.M. Variability in the behavior of spotted hyaenas as taphonomic agents. J. Archaeol. Sci. 19 (1992) 389-406
87. Lam Y.M., Chen X.B., and Pearson O.M. Intertaxonomic variability in patterns of bone density and the differential representation of bovid, cervid, and equid elements in the archaeological record. Am. Antiq. 64 (1999) 343-362
88. Leakey L.S.B. Preliminary report on a Chellean 1 living site at BK II, Olduvai. In: Clark J.D., and Cole S. (Eds). Proceedings of the 3rd Pan-African Congress on Prehistory (1957), Chatto and Windus, London 217-218
89. Leakey M.D. Olduvai Gorge, Vol. 3: Excavations in Beds I and II, 1960-1963 (1971), Cambridge University Press, Cambridge
90. Lupo K.D. Archaeological skeletal part profiles and differential transport: an ethnoarchaeological example from Hadza bone assemblages. J. Anthropol. Archaeol. 20 (2001) 361-378
91. Lyman R.L. Faunal analysis: an outline of method and theory with some suggestions. Northwest Anthropol. Res. Notes 13 (1979) 22-35
92. Lupo K.D., and O'Connell J.F. Cut and tooth mark distributions on large animal bones: ethnoarchaeological data from the Hadza and their implications for current ideas about early human carnivory. J. Archaeol. Sci. 29 (2002) 85-109
93. Lyman R.L. Vertebrate Taphonomy (1994), Cambridge University Press, Cambridge
94. Manega, P.C., 1993. Geochronology, geochemistry and isotopic study of the Plio-Pleistocene hominid sites and the Ngorongoro volcanic highland in northern Tanzania. Ph.D. Dissertation, University of Colorado, Boulder.
95. Marean C.W. A critique of the evidence for scavenging by Neandertals and early modern humans: new data from Kobeh Cave (Zagros Mountains, Iran) and Die Kelders Cave 1 Layer 10 (South Africa). J. Hum. Evol. 35 (1998) 111-136
96. Marean C.W., and Spencer L.M. Impact of carnivore ravaging on zooarchaeological measures of element abundance. Am. Antiq. 56 (1991) 645-658
97. Marean C.W., and Cleghorn N. Large mammal skeletal element transport: applying foraging theory in a complex taphonomic system. J. Taphonomy 1 (2003) 15-42
98. Marean C.W., Domínguez-Rodrigo M., and Pickering T.R. Skeletal element equifinality in zooarchaeology begins with method: the evolution and status of the "shaft critique.". J. Taphonomy 2 (2004) 69-98
99. Marean C.W., Spencer L.M., Blumenschine R.J., and Capaldo S.D. Captive hyaena bone choice and destruction, the schlepp effect and Olduvai archaeofaunas. J. Archaeol. Sci. 19 (1992) 101-121
100. Marean C.W., Abe Y., Frey C.J., and Randall R.C. Zooarchaeological and taphonomic analysis of the Die Kelders Cave 1 Layers 10 and 11 Middle Stone Age larger mammal fauna. J. Hum. Evol. 38 (2000) 197-233
101. McHenry H.M., and Coffing K. Australopithecus to Homo: transformations in body and mind. Annu. Rev. Anthropol. 29 (2000) 125-146
102. Metcalfe D., and Jones K.T. A reconsideration of animal body-part utility indices. Am. Antiq. 53 (1988) 486-504
103. Mills M.G.L., and Mills M.E.J. An analysis of bones collected at hyaena breeding dens in the Gemsbok National Parks. Ann. Transvaal Mus. 30 (1977) 145-155
104. Monahan C.M. New zooarchaeological data from Bed II, Olduvai Gorge, Tanzania: implications for hominid behavior in the early Pleistocene. J. Hum. Evol. 31 (1996) 93-128
105. Monahan, C.M., 1996b. Variability in the foraging behavior of early Homo: A taphonomic perspective from Bed II, Olduvai Gorge, Tanzania. Ph.D. Dissertation, University of Wisconsin, Madison.
106. Monahan C.M. The Hadza carcass transport debate and its archaeological implications. J. Archaeol. Sci. 25 (1998) 405-424
107. Mora R., and de la Torre I. Percussion tools in Olduvai Beds I and II (Tanzania): implications for early human activities. J. Anthropol. Archaeol. 24 (2005) 179-192
108. Morlan R.E. Toward the definition of criteria for the recognition of artificial bone alteration. Quatern. Res. 22 (1984) 160-171
109. Morlan R.E. Bison bone fragmentation and survivorship: a comparative approach. J. Archaeol. Sci. 21 (1994) 797-807
110. Oliver J.S. Analogues and site context: bone damage from Shield Trap Cave (24CB91), Carbon County, Montana, USA. In: Bonnichsen R., and Sorg M.H. (Eds). Center for the Study of the First Americans (1989), Bone Modification 61-71 Orono
111. Oliver J.S. Estimates of hominid and carnivore involvement in the FLK Zinjanthropus fossil assemblage: some socioecological implications. J. Hum. Evol. 27 (1994) 267-294
112. Peters C.R. Nut-like oil seeds: food for monkeys, chimpanzees, humans, and probably ape-men. Am. J. Phys. Anthropol. 73 (1987) 333-363
113. Peters C.R., and Blumenschine R.J. Landscape perspectives on possible land use patterns for early Pleistocene hominids in the Olduvai Basin, Tanzania. J. Hum. Evol. 29 (1995) 321-362
114. Peters C.R., Obrien E.M., and Box E.O. Plant types and seasonality of wild-plant foods, Tanzania to southwestern Africa: resources for models of the natural environment. J. Hum. Evol. 13 (1984) 397-414
115. Petraglia M.D., and Potts R. Water flow and the formation of early Pleistocene artifact sites in Olduvai Gorge, Tanzania. J. Anthropol. Archaeol 13 (1994) 228-254
116. Pickering T.R. Taphonomy of the Swartkrans hominid postcrania and its bearing on issues of meat-eating and fire management. In: Stanford C.B., and Bunn H.T. (Eds). Meat-Eating and Human Evolution (2001), Oxford University Press, Oxford 33-51
117. Pickering T.R. Carnivore voiding: a taphonomic process with the potential for the deposition of forensic evidence. J. Forensic Sci. 26 (2001) 406-411
118. Pickering T.R. Reconsideration of criteria for differentiating faunal assemblages accumulated by hyenas and hominids. Int. J. Osteoarchaeol 12 (2002) 127-141
119. Pickering T.R., and Egeland C.P. Experimental patterns of hammerstone percussion damage on bones: implications for inferences of carcass processing by humans. J. Archaeol. Sci. 33 (2006) 459-469
120. Pickering T.R., Marean C.W., and Domínguez-Rodrigo M. Importance of limb bone shaft fragments in zooarchaeology: a response to "On in situ attrition and vertebrate body part profiles" (2002) by M.C. Stiner. J. Archaeol. Sci. 30 (2003) 1469-1482
121. Pickering T.R., Domínguez-Rodrigo M., Egeland C.P., and Brain C.K. Beyond leopards: tooth marks and the contribution of multiple carnivore taxa to the accumulation of the Swartkrans Member 3 fossil assemblage. J. Hum. Evol. 46 (2004) 595-604
122. Pickering T.R., Domínguez-Rodrigo M., Egeland C.P., and Brain C.K. New data and ideas on the foraging behaviour of Early Stone Age hominids at Swartkrans Cave, South Africa. S. Afr. J. Sci. 100 (2004) 215-219
123. Pickering T.R., Domínguez-Rodrigo M., Egeland C.P., and Brain C.K. The contribution of limb bone fracture patterns to reconstructing early hominid behaviour at Swartkrans Cave (South Africa): archaeological application of a new analytical method. Int. J. Osteoarchaeol. 15 (2005) 247-260
124. Pickering T.R., Domínguez-Rodrigo M., Egeland C.P., and Brain C.K. Carcass foraging by early hominids at Swartkrans Cave (South Africa): a new investigation of the zooarchaeology and taphonomy of Member 3. In: Pickering T.R., Schick K., and Toth N. (Eds). Breathing Life into Fossils: Taphonomic Studies in Honor of C.K. (Bob) Brain (2007), Stone Age Institute Press, Bloomington 231-254
125. Pickering T.R., Egeland C.P., Domínguez-Rodrigo M., Brain C.K., and Schnell A.G. Testing the "shift in the balance of power" hypothesis at Swartkrans, South Africa: hominid cave use and subsistence behavior in the early Pleistocene. J. Anthropol. Archaeol. 27 (2008) 30-45
126. Pobiner, B.L., 2007. Hominin-carnivore interactions: evidence from modern carnivore bone modification and early Pleistocene archaeofaunas (Koobi Fora, Kenya; Olduvai Gorge, Tanzania). Ph.D. Dissertation, Rutgers University, New Brunswick.
127. Pobiner B.L., and Braun D.R. Strengthening the inferential link between cutmark frequency data and Oldowan hominid behavior: results from modern butchery experiments. J. Taphonomy 3 (2005) 107-119
128. Pokines J.T., and Kerbis Peterhans J.C. Spotted hyena (Crocuta crocuta) den use and taphonomy in the Masai Mara National Reserve, Kenya. J. Archaeol. Sci. 34 (2007) 1914-1931
129. Potts, R., 1982. Lower Pleistocene site formation and hominid activities at Olduvai Gorge, Tanzania. Ph.D. Dissertation, Harvard University, Cambridge.
130. Potts R. Early Hominid Activities at Olduvai (1988), Aldine de Gruyter, New York
131. Richardson P.R.K. Carnivore damage to antelope bones and its archaeological implications. Palaeontol. Afr. 23 (1980) 109-125
132. Schick K.D., and Toth N. Making Silent Stones Speak (1993), Simon and Schuster, New York
133. Schick K.D., and Toth N. Early Stone Age technology in Africa: a review and case study into the nature and function of spheroids and subspheroids. In: Corruchini R.S., and Ciochon R.L. (Eds). Integrative Paths to the Past: Paleoanthropological Advances in Honor of F. Clark Howell (1994), Prentice Hall, Englewood Cliffs 429-449
134. Selvaggio, M.M., 1994. Evidence from carnivore tooth marks and stone-tool-butchery marks for scavenging by hominids at FLK Zinjanthropus, Olduvai Gorge, Tanzania. Ph.D. Dissertation, Rutgers University, New Brunswick.
135. Selvaggio M.M. Evidence for a three-stage sequence of hominid and carnivore involvement with long bones at FLK Zinjanthropus, Olduvai Gorge, Tanzania. J. Archaeol. Sci. 25 (1998) 191-202
136. Selvaggio M.M., and Wilder J. Identifying the involvement of multiple carnivore taxa with archaeological bone assemblages. J. Archaeol. Sci. 28 (2001) 465-470
137. Sept J.M. Archaeological evidence and ecological perspectives for reconstructing early hominid subsistence behavior. J. Archaeol. Meth. Theory 4 (1992) 1-56
138. Sept J.M. Bone distribution in a semi-arid riverine habitat in eastern Zaire: implications for the interpretation of faunal assemblages at early archaeological sites. J. Archaeol. Sci. 21 (1994) 217-235
139. Sept J.M. Modeling the edible landscape. In: Stanford C.B., and Bunn H.T. (Eds). Meat-Eating and Human Evolution (2001), Oxford University Press, Oxford 73-98
140. Shipman P. Life History of a Fossil: An Introduction to Taphonomy and Paleoecology (1981), Harvard University Press, Cambridge
141. Sikes N.E. Early hominid habitat preferences in East Africa: paleosol carbonate isotopic evidence. J. Hum. Evol. 27 (1994) 25-45
142. Sikes N.E. Plio-Pleistocene floral context and habitat preferences of sympatric hominid species in East Africa. In: Bromage T.G., and Schrenk F. (Eds). African Biogeography, Climate Change and Human Evolution (1999), Oxford University Press, Oxford 301-315
143. Snyder L.M. A controlled feeding study involving gray wolf (Canus lupus) and white-tailed deer (Odocoileus virginianus). Tennessee Anthropological Association Newsletter 13 (1988) 1-9
144. Tacktikos, J.C., 2005. A landscape perspective on the Oldowan from Olduvai Gorge, Tanzania. Ph.D. Dissertation, Rutgers University, New Brunswick.
145. Tappen M. Savanna ecology and natural bone deposition: implications for early hominid site formation, hunting and scavenging. Curr. Anthropol. 36 (1995) 223-260
146. Tappen M., Lordkipanidze D., Bukshianidze M., Ferring R., and Vekua A. Are you in or out (of Africa)? Site formation at Dmanisi and actualistic studies in Africa. In: Pickering T.R., Schick K., and Toth N. (Eds). Breathing Life into Fossils: Taphonomic Studies in Honor of C.K. (Bob) Brain (2007), Stone Age Institute Press, Bloomington 119-135
147. Todd L.C., and Rapson D.J. Long bone fragmentation and interpretation of faunal assemblages: approaches to comparative analysis. J. Archaeol. Sci. 15 (1988) 307-325
148. de la Torre, I., 2004. Estrategias technológicas en el Pleistoceno inferior de África oriental (Olduvai y Peninj, norte de Tanzania). Ph.D. Dissertation, Universidad Complutense, Madrid.
149. Toth N. The Oldowan reassessed: a close look at early stone artifacts. J. Archaeol. Sci. 12 (1985) 101-120
150. Turner A., and Antón M. Climate and evolution: implications of some extinction patterns in African and European machairodontine cats of the Plio-Pleistocene. Estudios Geológicos 54 (1998) 209-230
151. Villa P., and Mathieu E. Breakage patterns of human long bones. J. Hum. Evol. 21 (1991) 27-48
152. Walker A., and Leakey R. The postcranial bones. In: Walker A., and Leakey R. (Eds). The Nariokotame Homo erectus Skeleton (1993), Harvard University Press, Cambridge 95-160
153. 39Ar dating to calibrating biological and climatic change. Quatern. Int. 13/14 (1992) 37-46
154. 39Ar dating of Bed I, Olduvai Gorge, Tanzania. Nature (1991) 145-149
155. Werdelin L., and Lewis M.E. Plio-Pleistocene Carnivora of eastern Africa: species richness and turnover patterns. Zool. J. Linn. Soc. 144 (2005) 121-144
156. Willoughby, P.R., 1987. Spheroids and Battered Stone in African Early Stone Age. British Archaeological Reports International Series, Oxford.