Revision and phylogenetic analysis of the Hadronema group (miridae: Orthotylinae: Orthotylini), with descriptions of new genera and new species, and comments on the neotropical genus Tupimiris

Bulletin of the American Museum of Natural History

Volumn , Issue 312, 2008, Pages 1-172

  Forero, Dimitri ^a  

^a Department of Molecular Biology and Genetics

Author keywords

[No Author keywords available]

Indexed keywords

AOPLONEMA; HADRONEMA; MIRIDAE; ORTHOTYLINAE; ORTHOTYLINI;

EID: 48549098236     PISSN: 00030090     EISSN: None     Source Type: Book Series    
DOI: 10.1206/312.1     Document Type: Article

References (142)

1. Agrawal, A.A. 2001. Phenotypic plasticity in the interactions and evolution of species. Science 294: 321-326.
2. Amarillo-Suárez, A.R., and C.W. Fox. 2006. Population differences in host use by a seed-beetle: local adaptation, phenotypic plasticity and maternal effects. Oecologia 150(2): 247-258.
3. Asquith, A. 1990. Taxonomy and variation of the Lopidea nigridea complex of western North America (Heteroptera: Miridae: Orthotylinae). Great Basin Naturalist 50(2): 135-154.
4. Asquith, A. 1991. Revision of the genus Lopidea in America north of Mexico (Heteroptera: Miridae: Orthotylinae). Theses Zoologicae 16: 1-280.
5. Barneby, R.C. 1977. Daleae imagines: an illustrated revision of Errazuriza Philippi, Psorothamnus Rydberg, Marina Leibmann, and Dalea Lucanus emend. Barneby, including all species of Leguminosae tribe Amorpheae Borissova ever referred to Dalea. Memoirs of the New York Botanical Garden 27: 1-891.
6. Barneby, R.C. 1990. Two new taxa in Dalea (Fabaceae: Amorpheae) from southern Mexico and northern Chile. Brittonia 42(2): 89-91.
7. Benhke, H.D. 1997. Sarcobataceae: a new family of Caryophyllales. Taxon 46(3): 495-507.
8. Betz, O., and G. Kölsh. 2004. The role of adhesion in prey capture and predator defense in arthropods. Arthropod Structure and Development 33: 3-30.
9. Beutel, R.G., and S.N. Gorb. 2001. Ultrastructure of attachment specializations of hexapods (Arthropoda): evolutionary patterns inferred from a revised ordinal phylogeny. Journal of Zoological Systematics and Evolutionary Research 39: 177-207.
10. Blatchley, W.S. 1926. Heteroptera or true bugs of eastern North America, with especial reference to the faunas of Indiana and Florida. Indianapolis: Nature Publishing Co., 1116 pp.
11. Braby, M.F., and J.W.H. Trueman. 2006. Evolution of larval host plant associations and adaptive radiation in pierid butterflies. Journal of Evolutionary Biology 19: 1677-1690.
12. Carayon, J. 1971. Notes et documents sur l'appareil odorant métathoracique des Hémiptères. Annales de la Société Entomologique de France 7(4): 737-770.
13. Carpenter, F.M. 1992. Treatise on invertebrate paleontology. Part R. Arthropoda 4. Vol. 3: Superclass Hexapoda. Peace: Geological Society of America, and University of Kansas, xxii + 277 pp.
14. Carpenter, J.M. 1989. Testing scenarios: wasp social behavior. Cladistics 5: 131-144.
15. Carrel, J.E., and T. Eisner. 1974. Cantharidin: potent feeding deterrent to insects. Science 183(4126): 755-757.
16. Carrol, S.P., and C. Boyd. 1992. Host race radiation in the soapberry bug: natural history with the history. Evolution 46(4): 1052-1069.
17. Carvalho, J.C.M. 1958. A catalogue of the Miridae of the world. Part 3. Arquivos do Museu Nacional Rio de Janeiro 47: 1-161.
18. Carvalho, J.C.M. 1961. Mirídeos neotropicais, 89: dois gèneros novos do Peru e Terra do Fogo. Revista Brasileira de Biologia 21: 461-466.
19. Carvalho, J.C.M. 1977. Mirídeos neotropicais, 209: descrições de quatro espécies novas e do macho de Hadronemisca corcovadensis Carvalho and Gomes (Hemiptera). Revista Brasileira de Biologia 37: 17-22.
20. Carvalho, J.C.M. 1983. Mirídeos neotropicais, 244: gênero Araucanocoris n. gen. com descrição de oito espécies novas (Hemiptera). Revista Brasileira de Biologia 43: 413-426.
21. Carvalho, J.C.M. 1984. Mirídeos neotropicais, 241: cinco gêneros e sete espécies novos (Hemiptera). Revista Brasileira de Biologia 44: 45-57.
22. Carvalho, J.C.M. 1985. Mirídeos neotropicais, 253: descrições de novos gêneros e espécies da tribo Orthotylini Van Duzee (Hemiptera). Revista Brasileira de Biologia 45: 249-298.
23. Carvalho, J.C.M., and C.R.S. Afonso. 1977. Mirídeos neotropicais, 208: sobre uma coleção enviada para estudo pela Academia de Ciências da Califórnia (Hemiptera). Revista Brasileira de Biologia 37(1): 7-16.
24. Carvalho, J.C.M., and D.L. Carpintero. 1992. Miridos neotropicales, 337: descripción de doce especies nuevas de Orthotylinae (Hemiptera). Revista Brasileira de Biologia 51: 763-780 (1991).
25. Carvalho, J.C.M., and L.A.A. Costa. 1993. Mirídeos neotropicais, 338: novo gênero e espécie de Mirinae e novas espécies de Orthotylinae (Hemiptera). Revista Brasileira de Biologia 52: 199-207 (1992).
26. Carvalho, J.C.M., and J.C. Schaffner. 1973. Neotropical Miridae, 156: Adsaileria, Cuneocoris and Tupimiris, new genera of Orthotylini (Hemiptera). Revista Brasileira de Biologia 33(supl.): 31-37.
27. Church, N.W., and G.H. Gerber. 1977. Observations on the ontogeny and habits of Lytta nuttalli, L. viridana, and L. cyanipennis (Coleoptera: Meloidae): the adults and eggs. Canadian Entomologist 109: 565-573.
28. Claridge, M.F.T., and M.C. Gilhham. 1992. Variation in populations of leafhoppers and planthoppers (Auchenorrhyncha): biotypes and biological species. In J.T. Sorensen and R. Foottit (editors), Ordination in the study of morphology: evolution and systematics of insects, 241-259. Amsterdam: Elsevier Science Publishers.
29. Clissold, F.J., G.D. Sanson, and J. Read. 2006. The paradoxical effects of nutrient ratios and supply rates on an outbreaking insect herbivore, the Australian plague locust. Journal of Animal Ecology 75: 1000-1013.
30. Cockerell, T.D.A. 1893. The entomology of the mid-Alpine zone of Custer County, Colorado. Transactions of the American Entomological Society 20: 305-370.
31. Crevecoeur, F.F. 1905. Additions to the list of the hemipterous fauna of Kansas. Transactions of the Kansas Academy of Science 19: 232-237.
32. Cuénoud, P., P. Savolainen, L.W. Chatrou, M. Powell, R.J. Grayer, and M.W. Chase. 2002. Molecular phylogenetics of Caryophyllales based on nuclear 18S rDNA and plastid rbcL, atpB, and matK DNA sequences. American Journal of Botany 89: 132-144.
33. Davis, N.T. 1955. Morphology of the female organs of reproduction in the Miridae (Hemiptera). Annals of the Entomological Society of America 48: 132-150.
34. Distant, W.L. 1883 (1880-1893). Insecta. Rhynchota. Hemiptera- Heteroptera. In F.D. Godman and O. Salvin (editors), Biologia Centrali-Americana Vol. 1, xx + 1-462, 39 pls. London: Taylor and Francis.
35. ESRI (Environmental Systems Research Institute), ArcMap: Release 9.1 [software]. Redlands, CA: ESRI Inc.
36. Farris, J. 1970. Methods for computing Wagner trees. Systematic Zoology 34: 21-24.
37. Fitch, W.M. 1971. Towards defining the course of evolution: minimum change for a specific tree topology. Systematic Zoology 20(4): 406-416.
38. Froeschner, R.C. 1949. Contributions to a synopsis of the Hemiptera of Missouri. Part 4. Hebridae, Mesoveliidae, Cimicidae, Anthocoridae, Cryptostemmatidae, Isometopidae, Meridae (sic). American Midland Naturalist 42(1): 123-188.
39. Gibson, E.H. 1918. The genus Hadronema Uhl. (Miridae, Heteroptera). Canadian Entomologist 50: 81-85.
40. Goloboff, P.A. 1993. Estimating character weights during tree search. Cladistics 9: 83-91.
41. Goloboff, P.A. 1997. Self-weighted optimization: tree searches and character state reconstructions under implied transformation costs. Cladistics 13: 225-245.
42. Goloboff, P.A. 1999. NONA (No Name), ver. 2. [software] Tucumán, Argentina: Published by the author.
43. Gruetzmacher, M.C., and J.C. Schaffner. 1977. Modification of the concept of the genus Hadronema Uhler as necessitated by a new species from Texas (Heteroptera, Miridae). Southwestern Entomologist 2: 53-56.
44. Haas, F., and S. Gorb. 2004. Evolution of locomotory attachment pads in the Dermaptera (Insecta). Arthropod Structure and Development 33: 45-66.
45. Hammer, Ø., D.A.T. Harper, and P.D. Ryan. 2001. PAST. Paleontological Statistics: software package for education and data analysis. Paleontologica electronica 4(1): 1-9, Available at:
46. Henry, T.J. 1995. Proboscidotylus carvalhoi, a new genus and species of sexually dimorphic plant bug from Mexico (Heteroptera: Miridae: Orthotylinae). Proceedings of the Entomological Society of Washington 97(2): 340-345.
47. Henry, T.J., and A.G. Wheeler, Jr. 1988. Family Miridae Hahn, 1833 (= Capsidae Burmeister, 1835).. In T.J. Henry and R.C. Froeschner (editors), Catalog of the Heteroptera, or True bugs of Canada and the continental United States: 251-507. Leiden: E.J. Brill.
48. ICZN (International Commission on Zoological Nomenclature). 1999. International Code of Zoological Nomenclature. 4th ed. London: International Trust for Zoological Nomenclature, xxix + 306 pp.
49. ICZN (International Commission on Zoological Nomenclature). 2007. Declaration 44. Available at .
50. James, L.F., D.B. Nielsen, and K.E. Pater. 1992. Impact of poisonous plants on the livestock industry. Journal of Range Management 45(1): 3-8.
51. Janz, N., and S. Nylin. 1998. Butterflies and plants: a phylogenetic study. Evolution 52(2): 486-502.
52. 4 photosynthesis. International Journal of Plant Sciences 164: 959-986.
53. Kelton, L.A. 1959. Male genitalia as taxonomic characters in the Miridae (Hemiptera). Canadian Entomologist, suppl 11: 1-72.
54. Kelton, L.A. 1968. Revision of the North American species of Slaterocoris (Heteroptera: Miridae). Canadian Entomologist 100(11): 1121-1137.
55. Kelton, L. A. 1980. The plant bugs of the prairie provinces of Canada. Heteroptera: Miridae. Part 8. The insects and arachnids of Canada. Ottawa: Agriculture Canada Research Branch Publication 1703, 1-408.
56. Kerzhner, I.M., and R.T. Schuh. 1995. Homonymy, synonymy, and new combinations in the Miridae (Heteroptera). American Museum Novitates 3137: 1-11.
57. Kirk, W.D. 1991. The size relationship between insects and their hosts. Ecological Entomology 16: 351-359.
58. Knight, H.H. 1918. Synoptic key to the subfamilies of Miridae (Hemiptera - Heteroptera). Journal of the New York Entomological Society 26: 40-44 +1 pl.
59. Knight, H.H. 1923. Family Miridae (Capsidae). In W.E. Britton (editor), Guide to the insects of Connecticut. Part 4. The Hemiptera or sucking insects of Connecticut. State of Connecticut Geological and Natural History Survey Bulletin 34: 422-658.
60. Knight, H.H. 1925. A list of Miridae and Anthocoridae from Alberta (Canada). Canadian Entomologist 57: 181-182.
61. Knight, H.H. 1928. Key to the species of Hadronema Uhler with descriptions of five new species (Hemiptera, Miridae). Canadian Entomologist 60: 177-182.
62. Knight, H.H. 1938. Strongylocoris Blanchard: six new species from North America (Hemiptera, Miridae). Iowa St. Coll. J. Sci. 13: 1-7, 1 pl.
63. Knight, H.H. 1941. The plant bugs, or Miridae, of Illinois. Bulletin of the Illinois Natural History Survey 22(1): 1-234.
64. Knight, H.H. 1962. Ten new and six old species of Lopidea from North America (Hemiptera, Miridae). Iowa State Journal of Science 37: 29-41.
65. Knight, H.H. 1968. Taxonomic review: Miridae of the Nevada test site and the western United States. Brigham Young University Science Bulletin Biological Series 9(3): 1-282.
66. Konstantinov, F.D. 2003. Male genitalia in Miridae (Heteroptera) and their significance for suprageneric classification of the family. Part 1: general review, Isometopinae and Psallopinae. Belgian Journal of Entomology 5: 3-36.
67. Lewinsohn, T.M., V. Novotny, and I. Basset. 2005. Insects on plants: diversity of herbivore assemblages revisited. Annual Review of Ecology and Systematics 36: 597-620.
68. Lindberg, H. 1956. Über einige Miriden in E. de Bergevins Sammlung. Notulae Entomologicae 36: 53-64.
69. Linnavuori, R.E. 1992. The lateralis group of the genus Dimorphocoris Reuter of Greece and the Middle East (Heteroptera, Miridae, Halticini). Entomologica Fennica 3: 215-222.
70. Maw, H.E.L., R.G. Foottit, K.G.A. Hamilton, and G.G.E. Scudder. 2000. Checklist of the Hemiptera of Canada and Alaska. Ottawa: NRC Research Press, 220 pp.
71. Mayhew, P.J. 1997. Adaptive patterns of host-plant selection by phytophagous insects. Oikos 79: 417-428.
72. McMahon, M.M. 2005. Phylogenetic relationships and floral evolution in the papilionoid legume clade Amorpheae. Brittonia 57(4): 397-411.
73. Mopper, S. 1996. Adaptive genetic structure in phytophagous insect populations. Trends in Ecology and Evolution 11(6): 235-238.
74. Nixon, K. 2002. WinClada. ver. 1.00.08. [software] Ithaca, NY: published by the author.
75. Pagola-Carte, S. 2006. Hypsitylus arberlaitz sp. nov. (Hemiptera: Heteroptera: Miridae) from Aiako Harria (Gipuzkoa, Basque Country, northern Iberian Peninsula). Heteropterus Revista de Entomologia 6: 75-81.
76. Pagola-Carte, S., and I. Zabalegui. 2006. Orthotylus (Pachylops) jordii sp. nov. (Hemiptera: Heteroptera: Miridae) from the Basque Country, northern Iberian Peninsula. Heteropterus Revista de Entomologia 6: 67-74.
77. Paskevich, V. 2005. srtm30plus-na_pctshade.tif - SRTM30PLUS color-encoded shaded relief image of North America (approximately 1 km) - GeoTIFF image: Open-File Report 2005-1001, U.S. Geological Survey, Coastal and Marine Geology Program, Woods Hole Science Center, Woods Hole, MA. Available at: .
78. Peppe, F.B., and C. Lomônaco. 2003. Phenotypic plasticity of Myzus persicae (Hemiptera: Aphididae) raised on Brassica oleracea L. var. acephala (kale) and Raphanus sativus L. (radish). Genetics and Molecular Biology 26(2): 189-194.
79. Pimentel, R.A. 1992. An introduction to ordination, principal components analysis and discriminant analysis. In J.T. Sorensen and R. Foottit (editors), Ordination in the study of morphology, evolution and systematics of insects, 11-28. Amsterdam: Elsevier Science Publishers.
80. Pinto, J.D. 1972. A synopsis of the bionomics of Phodaga alticeps (Coleoptera: Meloidae) with special reference to sexual behavior. Canadian Entomologist 104: 577-595.
81. Pinto, J.D. 1978. The parasitization of blister beetles by species of Miridae (Coleoptera: Meloidae; Hemiptera: Miridae). Pan-Pacific Entomologist 54: 57-60.
82. Pinto, J.D. 1984. A taxonomic review of Cysteodemus LeConte, Phodaga LeConte and Pleur opasta Wellman (Coleoptera: Meloidae: Eupomphina) with a new generic synonymy. Proceedings of the Entomological Society of Washington 86: 127-143.
83. Pinto, J.D. 1991. The taxonomy of North American Epicauta (Coleoptera: Meloidae), with a revision of the nominate subgenus and a survey of courtship behavior. University of California Publications in Entomology 110: i-x, 1-372, 40 pls.
84. Pluot-Sigwalt, D., and A. Matocq. 2006. On some particular sclerotized structures associated with the vulvar area and the vestibulum in Orthotylinae and Phylinae (Heteroptera, Miridae). Denisia 19: 557-570.
85. Polhemus, D.A., and J.T. Polhemus. 1984. Ephedrodoma, a new genus of orthotyline Miridae (Hemiptera) from western United States. Proceedings of the Entomological Society of Washington 86(3): 550-554.
86. Pulawski, W., I.M. Kerzhner, D.J. Brothers, and N.L. Evenhuis. 2002. Draft proposal to emend Article 74.7.3: request for comments from the Commission and zoological community. Bulletin of Zoological Nomenclature 59(4): 278-280.
87. Reuter, O.M. 1883. Hemiptera Gymnocerata Europae. Hémiptè res Gymnocérates d'Europe, du bassin de la Méditerrranée et de l'Asie Russe. III. Acta Societatis Seientiarum Fennicae 13: 313-496, 5 pls.
88. Reuter, O.M. 1908. Capsidae Mexicanae, a Do Bilimek collectae in Museo i.r. Vindobonensi asservatae. Annalen des Naturhistorischen Museums in Wien 22: 150-179.
89. Rios, N.E., and H.L. Bart, Jr. 2005. GEOLocate. Georeferencing software for natural history collections. Version 2.13 [software]. Available at: .
90. Schaffner, J.C., and P.S.F. Ferreira. 1995. Rolstonocoris, a new genus of Neotropical Miridae (Heteroptera: Orthotylinae). Journal of the New York Entomological Society 103: 374-385.
91. Schuh, R.T. 1974. The Orthotylinae and Phylinae (Hemiptera: Miridae) of South Africa with a phylogenetic analysis of the ant-mimetic tribes of the two subfamilies for the world. Entomologica Americana 47: 1-332.
92. Schuh, R.T. 1989. Review of Daleapidea Knight (Heteroptera: Miridae: Orthotylinae: Orthotylini). Journal of the New York Entomological Society 97(2): 159-166.
93. Schuh, R.T. 1995. Plant bugs of the world (Insecta: Heteroptera: Miridae). Systematic catalog: distributions, host list, and bibliography. New York: New York Entomological Society, 1329 pp.
94. Schuh, R.T. 2000a. Revision of Oligotylus Van Duzee with descriptions of ten new species from western North America and comments on Lepidargyrus in the Nearctic (Heteroptera: Miridae: Phylinae: Phylini). American Museum Novitates 3300: 1-44.
95. Schuh, R.T. 2000b. Biological systematics: Principles and applications. Ithaca, NY: Cornell University Press, 236 pp.
96. Schuh, R.T. 2001. Revision of New World Plagiognathus Fieber: with comments on the Palearctic fauna and the description of a new genus (Heteroptera, Miridae, Phylinae). Bulletin of the American Museum of Natural History 266: 1-267.
97. Schuh, R.T. 2004. Revision of Europiella Reuter in North America, with the description of a new genus (Heteroptera: Miridae: Phylinae). American Museum Novitates 3463: 1-58.
98. Schuh, R.T. 2006. Revision, phylogenetic, biogeographic and host analyses of the endemic Western North American Phymatopsallus group, with the description of 9 new genera and 15 new species (Insecta: Hemiptera: Miridae: Phylinae). Bulletin of the American Museum of Natural History 301: 1-115.
99. Schuh, R.T., and M.D. Schwartz. 1988. Revision of the New World Pilophorini (Heteroptera: Miridae: Phylinae). Bulletin of the American Museum of Natural History 187(2): 102-201.
100. Schwartz, M.D. 1987. Phylogenetic revision of the Stenodemini with a review of the Mirinae (Heteroptera: Miridae). PhD dissertation City University of New York, 383 pp.
101. Schwartz, M.D. 2004. Melymacra, a new plant bug genus (Heteroptera: Miridae: Orthotylinae) inhabiting manzanita, Arctostaphylos Adanson (Ericaceae). Pan-Pacific Entomologist 80(1-4): 29-41.
102. Scudder, G.G.E. 1959. The female genitalia of the Heteroptera: morphology and bearing on classification. Transactions of the Royal Entomological Society of London 111: 405-467.
103. Scudder, S.H. 1890. The Tertiary iInsects of North America. Report of the United States Geological Survey of the Territories 13: 1-734.
104. Seigler, D.S. 2004. Fabaceae. In N. Smith, S.A. Mori, A. Henderson, D.W. Stevenson, and S.V. Heald (editors), Flowering plants of the Neotropics, 151-156. Princeton, NJ: Princeton University Press and New York Botanical Garden.
105. Selander, R.B. 1960. Bionomics, systematics, and phylogeny of Lytta, a genus of blister beetles (Coleoptera, Meloidae). Illinois Biological Monographs 28: i-vi, 1-295.
106. Selander, R.B., and J.C. Downey. 1963. Distributional and food plant records in the Cyanipennis subgroup of the genus Lytta (Coleoptera: Meloidae). Pan-Pacific Entomologist 39(2): 124-129.
107. Silva-Brandão, K.L., and V.N. Solferini. 2007. Use of host plants by Troidini butterflies (Papilionidae, Papilioninae): constraints on host shift. Biological Journal of the Linnean Society 90: 247-261.
108. Slater, J.A. 1950. An investigation of the female genitalia as taxonomic characters in the Miridae (Hemiptera). Iowa State College Journal of Science 25: 1-81, 7 pls.
109. Soulier-Perkins, A., D. Ouvrard, M. Attié, and T. Bourgoin. 2007. Evolutionary patterns in biogeography and host plant association: 'taxonomic conservatism' in Lophopidae (Hemiptera, Fulgoromorpha). Systematic Entomology 32: 305-311.
110. Southwood, T.R.E. 1953. The morphology and taxonomy of the genus Orthotylus Fieber (Hem., Miridae), with special reference to the British species, e Transactions of the Royal Entomological Society of London 104: 415-449.
111. Steyskal, G. 1973. The grammar of the names in the Catalogue of the Miridae (Heteroptera) of the World by Carvalho, 1957-1960. Studia Entomologica 16(1-4): 203-208.
112. Stonedahl, G.M. 1990. Revision and cladistic analysis of the Holarctic genus Atractotomus Fieber (Heteroptera, Miridae, Phylinae). Bulletin of the American Museum of Natural History 198: 1-88.
113. Stonedahl, G.M., and R.T. Schuh. 1986. Squamocoris Knight and Ramentomiris, new genus (Heteroptera: Miridae: Orthotylinae): a cladistic analysis and description of seven new species from Mexico and the western United States. American Museum Novitates 2852: 1-26.
114. Stonedahl, G.M., and M.D. Schwartz. 1986. Revision of the plant bug genus Pseudopsallus Van Duzee (Heteroptera: Miridae). American Museum Novitates 2842: 1-58.
115. Stonedahl, G.M., and M.D. Schwartz. 1988. New species of Oaxacacoris Schwartz and Stonedahl and Pseudopsallus Van Duzee, and a new genus, Presidiomiris, from Texas (Heteroptera: Miridae: Orthotylini). American Museum Novitates 2928: 1-18.
116. Stork, N.E. 1981. The structure and function of the adhesive organs on the antennae of male Harpocera thoracica (Fallén) (Miridae; Hemiptera). Journal of Natural History 15: 639-644.
117. Swofford, D.L., G.J. Olsen, P.J. Waddell, and D.M. Hillis. 1996. Phylogenetic inference. In D.M. Hillis, C. Moritz, and B. Mable (editors), Molecular systematics. 2nd ed., 407-514. Sunderland, MA: Sinauer Associates.
118. Townsend, C.H.T. 1891. Hemiptera collected in southern Michigan. Proceedings of the Entomological Society of Washington 2(1): 52-56.
119. Townsend, C.H.T. 1892. Biologic notes in New Mexico insects. Canadian Entomologist 24(1): 193-200.
120. Tucker, E.S. 1907. Some results of desultory collecting of insects in Kansas and Colorado: list of Hemiptera Heteroptera. Kansas University Science Bulletin 4: 51-112.
121. Uhler, P.R. 1861. Descriptions of a few new species of Hemiptera, and observations upon some already described. Proceedings of the Entomological Society of Philadelphia 1: 21-24.
122. Uhler, P.R. 1872. Notices on the Hemiptera of the western territories of the United States, chiefly from the surveys of Dr. F.V. Hayden In E.V. Hayden (editor), Preliminary report of the United States Geological Survey of Montana and portions of adjacent territories, 392-423.
123. Uhler, P.R. 1886. Checklist of the Hemiptera Heteroptera of North America. Brooklyn, NY: Brooklyn Entomological Society, 29 pp.
124. Uhler, P.R. 1894. Observations upon the heteropterous Hemiptera of Lower California, with descriptions of new species. Proceedings of the California Academy of Sciences 2nd series 4: 223-295.
125. Uhler, P.R. 1895. [Descriptions of new species]. In O.P. Gillette and C.F. Baker (editors), A preliminary list of Hemiptera of Colorado. Bulletin of the Colorado Agricultural Experimental Station 31: 1-137.
126. USDA-NRCS. 2006. The PLANTS Database. National Plant Data Center, Baton Rouge, LA 70874490 USA. Accessed: 12 September 2006. Available at: .
127. USGS [United States Geological Survey]. 2007. Geographic Names Information System (GNIS). Available at: .
128. Van Duzee, E.P. 1910. Descriptions of some new or unfamiliar North American Hemiptera. Transactions of the American Entomological Society 36: 73-88.
129. Van Duzee, E.P. 1917. Catalogue of the Hemiptera of America north of Mexico (excepting the Aphididae, Coccidae, and Aleurodidae). University of California Publications in Entomology 2: i-xiv, 1-902.
130. Van Duzee, E.P. 1918. New species of Hemiptera, chiefly from California. Proceedings of the California Academy of Sciences 4th series 8(7): 271-308.
131. Van Duzee, E.P. 1928. A misidentified Hadronema (Hemiptera). Pan-Pacific Entomologist 4: 182 pp.
132. Verdon, A., N. Margraf, A.C. Davison, M. Rahier, and R.S. Naisbit. 2007. Conserved oviposition preferences in alpine leaf beetle populations despite host shifts and isolation. Ecological Entomology 32: 62-69.
133. Visser, J.H. 1986. Host odor perception in phytophagous insects. Annual Review of Entomology 31: 121-144.
134. Wall, M.A. 2007. A new genus and new species of Austro-Papuan Orthotylinae (Heteroptera: Miridae). American Museum Novitates 3558: 1-17.
135. Weirauch, C. 2006. New genera and species of oak-associated Phylini (Heteroptera: Miridae: Phylinae) from western North America. American Museum Novitates 3522: 1-54.
136. Weirauch, C. 2007. Hairy attachment structures in Reduviidae (Cimicomorpha, Heteroptera), with observations on the fossula spongiosa in some other Cimicomorpha. Zoologischer Anzeiger 246: 155-175.
137. Wheeler, A.G., Jr. 2001. Biology of the plant bugs (Hemiptera: Miridae): pests, predators, opportunists. Ithaca, NY: Cornell University Press, 507 pp.
138. Wheeler, W., L. Aagesen, C.P. Arango, J. Faivovich, T. Grant, C. D'Haese, D. Janies, W.M. Smith, A. Varón, and G. Giribet. 2006. Dynamic homology and phylogenetic systematics: a unified approach using POY. New York: American Museum of Natural History, 365 pp.
139. Wink, M. 1992. The role of quinolizidine alkaloids in plant-insect interactions. In E. Bernays (editor), Insect-plant interactions. Vol. 4, 131-166. Boca Raton, FL: CRC Press.
140. Wyniger, D., and D. Burckhardt. 2003. Die Landwanzenfauna (Hemiptera, Heteroptera) von Basel (Schweiz) und Umgebung. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 76: 1-136.
141. Young, D.K. 1984a. Cantharidin and insects: an historical review. Great Lakes Entomologist 17(4): 187-194.
142. Young, D.K. 1984b. Field records and observations of insects associated with cantharidin. Great Lakes Entomologist 17(4): 195-199.