Cambrian naraoiids (Arthropoda): Morphology, ontogeny, systematics, and evolutionary relationships

Journal of Paleontology

Volumn 81, Issue 5 SUPPL. 2, 2007, Pages 1-52

  Zhang, X L ^a   Shu, D G ^a   Erwin, D H ^b  

^a NORTHWEST UNIVERSITY   (China)

^b DEPARTMENT OF PALEOBIOLOGY   (United States)

Author keywords

[No Author keywords available]

Indexed keywords

ARTHROPOD; BURGESS THEORY; CAMBRIAN; DIMORPHISM; EVOLUTION; FUNCTIONAL MORPHOLOGY; GEOGRAPHICAL VARIATION; LAGERSTATTE; NEW RECORD; ONTOGENY; SILURIAN; SPATIAL DISTRIBUTION; SPECIES OCCURRENCE; SUPERCONTINENT; TAXONOMY;

ASIA; CHENGJIANG; CHINA; EURASIA; FAR EAST; YUNNAN;

ARTHROPODA; NARAOIA COMPACTA; PROTASPIS; TRILOBITA;

EID: 34948814518     PISSN: 00223360     EISSN: None     Source Type: Journal    
DOI: 10.1666/06-082.1     Document Type: Article

References (170)

1. ALLISON, P. A. AND D. E. G. BRIGGS. 1991. Taphonomy of non-mineralized tissues, p. 231-262. In P. A. Allison and D. E. G. Briggs (eds.), Taphonomy: Releasing the Data Locked in the Fossil Record. Plenum Press, New York and London.
2. ANDERSON, M. 1994. Sexual Selection. Princeton University Press, Princeton, 599 p.
3. ARTHUR, W. AND M. FARROW. 1999. The pattern of variation in centipede segment number as an example of developmental constraint in evolution. Journal of Theoretical Biology, 200:183-191.
4. ARTHUR, W. AND C. KETTLE. 2001. Geographic patterning of variation in segment number in geophilomorph centipedes: Clines and speciation. Evolution & Development, 3:34-40.
5. BABCOCK, L. E. 1993. Trilobite malformations and the fossil record of behavioral asymmetry. Journal of Paleontology, 67:217-229.
6. BABCOCK, L. E. 2003. Trilobites in Paleozoic predator-prey systems, and their role in reorganization of Early Paleozoic ecosystems, p. 55-92. In P. H. Kelley, M. Kowalewski, and T. A. Hansen (eds.), Predator-Prey Interactions in the Fossil Record. Kluwer Academic/Plenum Publishers, New York.
7. BABCOCK, L. E. AND W.-T. ZHANG. 1997. Comparative taphonomy of two nonmineralized arthropods: Naraoia (Nektaspida; Early Cambrian, Cheng-jiang biota, China) and Limulus (Xiphosauida; Holocene, Atlantic Ocean). Bulletin of the National Museum of Natural Science, 10:233-250.
8. BABCOCK, L. E. AND W.-T. ZHANG. 2001. Stratigraphy, paleontology and depositional setting of the Chengjiang Lagerstätte (Lower Cambrian), Yunnan, China, p. 66-86. In S.-C. Peng, L. E. Babcock, and M.-Y. Zhu (eds.), Cambrian System of South China. University of Science and Technology of China Press, Hefei.
9. BABCOCK, L. E., W.-T. ZHANG, AND S. A. LESLIE. 2001. The Chengjiang Biota: Record of the Early Cambrian diversification of life and clues to exceptional preservation of fossils. GSA Today, 11:4-9.
10. BENGTSON, S. AND X.-G. HOU. 2001. The integument of Cambrian chancelloriids. Acta Palaeontologica Polonica, 46:1-22.
11. BENGTSON, S., S. CONWAY MORRIS, B. J. COOPER, P. A. JELL, AND B. N. RUNNEGAR. 1990. Early Cambrian fossils from South Australia. Memoir of the Association of Australasian Palaeontologists, 9:364.
12. BERGSTRÖM, J. 2001. Chengjiang, p. 337-340. In D. E. G. Briggs and P. R. Crowther (eds.), Paleobiology II. Blackwell Science, Oxford.
13. BERGSTRÖM, J. AND X.-G. HOU. 1998. Chengjiang arthropods and their bearing on early arthropod evolution, p. 151-184. In G. D. Edgecombe (ed.). Arthropod Fossils and Phylogeny. Columbia University Press, New York.
14. BERGSTRÖM, J. AND X.-G. HOU. 2003. Arthropod origins. Bulletin of Geosciences, 78:323-334.
15. BRIGGS, D. E. G. 1976. The arthropod Branchiocaris n. gen., Middle Cambrian, Burgess Shale, British Columbia. Geological Survey of Canada Bulletin, 264:1-29.
16. BRIGGS, D. E. G. AND D. COLLINS. 1988. A Middle Cambrian chelicerate from Mount Stephen, British Columbia. Palaeontology, 31:779-798.
17. BRIGGS, D. E. G. AND D. COLLINS. 1999. The arthropod Alalcomenaeus cambricus Simonetta, from the Middle Cambrian Burgess Shale of British Columbia. Palaeontology, 42:953-977.
18. BRIGGS, D. E. G. AND A. J. KEAR. 1993. Decay and preservation of polychaetes: Taphonomic thresholds in soft-bodied organisms. Paleobiology, 19: 107-135.
19. BRIGGS, D. E. G. AND A. J. KEAR. 1994. Decay and mineralization of shrimps. Palaios, 9:431-456.
20. BRIGGS, D. E. G., D. H. ERWIN, AND F. J. COLLIER. 1994. The Fossils of the Burgess Shale. Smithsonian Institution Press, Washington, DC, 238 p.
21. BROGNIART, A. 1822. Les Trilobites, p. 1-65. In A. Brogniart and A. G. Desmarest (eds.), Histoire naturelle des crustacés fossils, sous les repports zoologiques et géologiques. Paris.
22. BRUSCA, R. C. AND G. J. BRUSCA. 2003. Invertebrates. Sinauer Associates, Sunderland, Massachusetts, 936 p.
23. BRUTON, D. L. 1981. The arthropod Sidneyia inexpectans. Middle Cambrian, Burgess Shale, British Columbia. Philosophical Transactions of the Royal Society, London B, 295:619-656.
24. BUDD, G. E. 1999. A nektaspid arthropod from the Early Cambrian Sirius Passet fauna, with a description of retrodeformation based on functional morphology. Palaeontology, 42:99-122.
25. BUDD, G. E. AND S. JENSEN. 2000. A critical reappraisal of the fossil record of bilaterian phyla. Biological Reviews, 75:253-295.
26. BUDIL, P., O. FATKA, AND J. BRUTHANSOVÁ . 2003. Trilobite fauna of the Šárka Formation at Praha-Červený vrch Hill (Ordovician, Barrandian area, Czech Republic). Bulletin of Geosciences, 78:113-117.
27. BUTTERFIELD, N. J. 2002. Leanchoilia guts and the interpretation of three-dimensional structures in Burgess Shale-type fossils. Paleobiology, 28:155-171.
28. CARON, J.-B., S. MILLIKEN, AND D. M. RUDKIN. 2002. A Late Silurian (Pridolian) naraoiid arthropod from the Bertie Formation of Southern Ontario - delayed fall-out from the Cambrian Explosion. Canadian Paleontology Conference, Ottawa, Programs and Abstracts, 12:3-6.
29. CARON, J.-B., D. M. RUDKIN, AND S. MILLIKEN. 2004. A new Late Silurian (Pridolian) naraoiid (Euarthrapoda: Nektaspida) from the Bertie Formation of southern Ontario, Canada - delayed fallout from the Cambrian explosion. Journal of Paleontology, 78:1138-1145.
30. CHATTERTON, B. D. E. AND S. E. SPEYER. 1997. Ontogeny, p. 173-247. In R. L. Kaesler (ed.), Treatise on Invertebrate Paleontology, Pt. O, Arthropoda 1. Trilobita. Revised. Geological Society of America and University of Kansas Press, Lawrence.
31. CHEN, J.-Y. 2004. The Dawn of the Animal World. Jiangsu Science and Technology Press, Nanjing, 366 p.
32. CHEN, J.-Y. AND B.-D. ERDTMANN. 1991. Lower Cambrian lagerstätte from Chengjiang, Yunnan, China: Insights for reconstructing early metazoan life, p. 57-76. In A. M. Simonetta and S. Conway Morris (eds.), The Early Evolution of Metazoa and the Significance of Problematic Taxa. Cambridge University Press, Cambridge.
33. CHEN, J.-Y. AND D.-Y. HUANG. 2002. A possible Lower Cambrian chaeto-gnath (arrow worm). Science, 298:187.
34. CHEN, J.-Y. AND G.-Q. ZHOU. 1997. The biology of the Chengjiang fauna. Bulletin of the National Museum of Natural Science, 10:11-106.
35. CHEN, J.-Y., G. D. EDGECOMBE, AND L. RAMSKÖLD. 1997. Morphological and ecological disparity in naraoiids (Arthropoda) from the Early Cambrian Chengjiang fauna, China. Records of the Australian Museum, 49:1-24.
36. CHEN, J.-Y., G.-Q. ZHOU, AND L. RAMSKÖLD. 1995. A new Early Cambrian onychophoran-like animal, Paucipodia gen. nov., from the Chengjiang fauna, China. Transactions of the Royal Society of Edinburgh: Earth Sciences, 85:275-282.
37. CHEN, J.-Y., J. BERGSTRÖM, J. LINDSTRÖM, AND X.-G. HOU. 1991. Fossilized soft-bodied fauna. National Geographic Research & Exploration, 7(1):8-19.
38. CHEN, J.-Y., G.-Q. ZHOU, M.-Y. ZHU, AND K. YEH. 1996. The Chengjiang Biota - A Unique Window of the Cambrian Explosion. The National Museum of Natural Science, Taichung, Taiwan, 222 p. (In Chinese)
39. CHEN, L.-Z., H.-L. LUO, S.-X. HU, G.-Y. YIN, Z.-W. JIANG, Z.-L. WU, F. LI, AND A.-L. CHEN. 2002. Early Cambrian Chengjiang fauna in Eastern Yunnan, China. Yunnan Science and Technology Press, Kunming, 199 p., 28 pls. (In Chinese, with English summary)
40. COLLINS, D. 2000. The geology and biology of the Middle Cambrian Burgess Shale, Canada's most important fossil fauna. Canadian Society of Exploration Geophysicists, Conference Abstracts: 141.
41. CONWAY MORRIS, S. 1977. Fossil priapulid worms. Special Papers in Palaeontology, 20:1-95.
42. CONWAY MORRIS, S. 1985. The Middle Cambrian metazoan Wiwaxia corrugata (Matthew) from the Burgess Shale and Ogygopsis Shale, British Columbia, Canada. Philosophical Transactions of the Royal Society, London, B, 307:507-582.
43. CONWAY MORRIS, S. 1998. The Crucible of Creation. Oxford University Press, Oxford, 242 p.
44. CONWAY MORRIS, S. AND J. PEEL. 1995. Articulated halkieriids from the Lower Cambrian of north Greenland and their role in early protostome evolution. Philosophical Transactions of the Royal Society, London, B, 347: 305-358.
45. COTTON, T. J. AND S. J. BRADDY. 2004. Phylogeny of arachnomorph arthropods and the origin of the Chelicerata. Transactions of the Royal Society of Edinburgh, Earth Science, 94:169-193.
46. DALZIEL, I. W. D. 1997. Neoproterozoic-Paleozoic geography and tectonics: Review, hypothesis, environmental speculation. Geological Society of America Bulletin, 09:16-42.
47. DELLE CAVE, L. AND A. M. SIMONETTA. 1991. Early Paleozoic arthropods and problems of arthropod phylogeny: With some notes on taxa of doubtful affinities, p. 189-244. In A. M. Simonetta and S. Conway Morris (eds.), The Early Evolution of Metazoa and the Significance of Problematic Taxa. Cambridge University Press, Cambridge.
48. DZIK, J. AND K. LENDZION. 1988. The oldest arthropods of the east European platform. Lethaia, 21:29-38.
49. EDGECOMBE, G. D. AND L. RAMSKÖLD. 1999. Relationships of Cambrian Arachnata and the systematic position of Trilobita. Journal of Paleontology, 73:263-287.
50. ERDTMANN, B.-D., M. STEINER, AND H. SIEGMUND. 1994. Entwicklungsgeschichte des höheren Lebens an der Wende vom Präkambrium zum Kambrium. Ausfschluss, 45:26-35.
51. FELGENHAUER, B. E., L. W. ABELE, AND D. L. FELDER. 1992. Remipedia, p. 225-247. In F. W. Harrison and A. G. Humes (eds.), Microscopic Anatomy of Invertebrates. Volume 9. Crustacea. Wiley-Liss, New York.
52. FORTEY, R. A. 1997. Classification, p. 289-302. In R. L. Kaesler (ed.), Treatise on Invertebrate Paleontology, Pt. O, Arthropoda 1. Trilobita 1 Revised. The Geological Society of America and the University of Kansas Press, Lawrence.
53. FORTEY, R. A. AND R. M. OWENS. 1990. Evolutionary radiations in the Trilobita, p. 139-164. In P. D. Taylor and G. Larwood (eds.), Major Evolutionary Radiations. Clarendon Press, Oxford.
54. FORTEY, R. A. AND J. N. THERON. 1994. A new Ordovician arthropod, Soomaspis, and the agnostid problem. Palaeontology, 37:841-861.
55. FORTEY, R. A., D. E. G. BRIGGS, AND M. A. WILLS. 1996. The Cambrian evolutionary 'explosion': Decoupling cladogenesis from morphological disparity. Biological Journal of the Linnean Society, 57:13-33.
56. GABBOTT, S. E., X.-G. HOU, M. J. NORRY, AND D. J. SIVETER. 2004. Preservation of Early Cambrian animals of the Chengjiang biota. Geology, 32: 901-904.
57. GEYER, G. 1998. Intercontinental, trilobite-based correlation of the Moroccan Early Middle Cambrian. Canadian Journal of Earth Sciences, 35:374-401.
58. GEYER, G. AND J. SHERGOLD. 2000. The quest for internationally recognized divisions of Cambrian time. Episodes, 23:188-195.
59. GOULD, S. J. 1989. Wonderful Life: The Burgess Shale and the Nature of History. W. W. Norton, New York, 347 p.
60. GRÜNER, H. E. 1993. Lehrbuch der speziellen Zoologie, Bd 1, Wirbellose, 4. Teil: Arthropoda (ohne Insecta). Fischer, Stuttgart, 666 p.
61. HAGADORN, J. W. 2002. Chengjiang: Early record of the Cambrian Explosion, p. 35-60. In D. J. Bottjer, W. Etter, J. W. Hagadorn, and C. M. Tang (eds.), Exceptional Fossil Preservation: A Unique View on the Evolution of Marine Life. Columbia University Press, New York.
62. HAMMANN, W., R. LASKE, AND G. L. PILLOLA. 1990. Tarricoia arrusensis n. g. n. sp., an unusual trilobite-like arthropod. Rediscovery of the 'Phyllocarid' Beds of Taricoo (1922) in the Ordovician 'Puddinga' sequence of Sardinia. Bolletino della Societa Palaeontologica Italiana, 29:163-178.
63. HOU, X.-G. 1987. Three new large arthropods from Lower Cambrian, Chengjiang, eastern Yunnan. Acta Palaeontologica Sinica, 26:272-85.
64. HOU, X.-G. 1993. The arthropod Naraoia from the Lower Cambrian Chengjiang fauna. Lundadagarna I Historisk Geologi och Paleontologi, III, Abstract: 12.
65. HOU, X.-G. AND J. BERGSTRÖM. 1997. Arthropods of the Lower Cambrian Chengjiang fauna, southwest China. Fossils and Strata, 45:1-115.
66. HOU, X.-G., J.-Y. CHEN, AND H.-Z. LU. 1989. Early Cambrian new arthropods from Chengjiang, Yunnan. Acta Palaeontologica Sinica, 28:42-57.
67. HOU, X.-G., L. RAMSKÖLD, AND J. BERGSTRÖM. 1991. Composition and preservation of the Chengjiang fauna - a Lower Cambrian soft-bodied biota. Zoologica Scripta, 20:395-411.
68. HOU, X.-G., D. J. SIVETER, M. WILLIAMS, AND X.-H. FENG. 2002. A monograph of the bradoriid arthropods from the Lower Cambrian of southwest China. Transactions of the Royal Society of Edinburgh: Earth Sciences, 92: 347-409.
69. HOU, X.-G., R. J. ALDRIDGE, J. BERGSTRÖM, D. J. SIVETER, AND X.-H. FENG. 2004. The Cambrian Fossils of Chengjiang, China: The Flowering of Early Animal Life. Blackwell Science, Oxford, 233 p.
70. HOU, X.-G., J. BERGSTRÖM, H.-F. WANG, X.-F. FENG, AND A.-L. CHEN. 1999. The Chengjiang Fauna - Exceptionally Weil-Preserved Animals from 530 Million Years Ago. Yunnan Science and Technology Press, Kunming, 170 p. (In Chinese with English summary)
71. HUANG, D.-Y., J. VANNIER, AND J.-Y. CHEN. 2003. Anatomy and life styles of Early Cambrian priapulid worms exemplified by Corynetis and Anningella from the Early Cambrian Maotianshan Shale (southwest China). Lethaia, 37:21-33.
72. HUANG, D.-Y., J. VANNIER, AND J.-Y. CHEN. 2004a. Recent Priapulidae and their early Cambrian ancestors: Comparisons and evolutionary significance. Geobios, 37:217-228.
73. HUANG, D.-Y., J.-Y. CHEN, J. VANNIER, AND J. I. SAIZ-SALINAS. 2004b. Early Cambrian sipunculan worms from southwest China. Proceedings of the Royal Society of London, B, 271:1671-1676.
74. HUGHES, C. P. 1975. Redescription of Burgessia bella from the Middle Cambrian Burgess Shale, British Columbia. Fossils and Strata, 4:415-436.
75. HUGHES, N. C. 2003. Trilobite tagmosis and body patterning from morphological and developmental perspectives. Integrative and Comparative Biology, 43:185-206.
76. JAGO, J. B. AND P. W. HAINES. 2002. Repairs to an injured early Middle Cambrian trilobite, Elkedra Area, Northern Territory. Alcheringa, 26:19-21.
77. JELL, P. A. AND J. M. ADRAIN. 2003. Available generic names for trilobites. Memoirs of the Queensland Museum, 48:331-553.
78. KETTLE, C., J. JOHNSTONE, T. JOWETT, H. ARTHUR, AND W. ARTHUR. 2003. The pattern of segment Formation, as revealed by engrailed expression, in a centipede with variable number of segments. Evolution & Development, 5:198-207.
79. KLINGENBERG, C. P. 1998. Heterochrony and allometry: The analysis of evolutionary change in ontogeny. Biological Reviews, 73:79-123.
80. LANDING, E. 1994. Precambrian-Cambrian boundary global stratotype ratified and a new perspective of Cambrian time. Geology, 22:179-184.
81. LESLIE, S. A., L. E. BABCOCK, J. C. MITCHELL, AND W.-T. ZHANG. 1996. Phosphatization and its relationship to exceptional preservation of fossils in the Chengjiang Lagerstätte (Lower Cambrian, China). Geological Society of America Abstracts with Programs, 27:294.
82. LI, Y.-W. 1975. On the Cambrian Ostracoda with new material from Sichuan, Yunnan and southern Shaanxi, China. Professional Papers on Stratigraphy and Paleontology, 2:37-72.
83. LI, Z.-X., L. ZHANG, AND C. M. POWELL. 1995. South China in Rodinia: Part of the missing link between Australia-East Antarctica and Laurentia? Geology, 23:407-410.
84. LIEBERMAN, B. S. 2003. Biogeography of Trilobita during the Cambrian radiation: Deducing geological processes from trilobite evolution. In P. D. Lane, D. J. Siveter, and R. A. Fortey (eds.), Trilobites and Their Relatives. Special Papers in Palaeontology, 70:59-72.
85. LIN, J.-P. 2003. Systematics and Taphonomy of Naraoia and Skania (Arthropoda) from Guizhou, China, and western North America. Unpublished M.S. thesis, Ohio State University, Columbus, 84 p.
86. LIN, J.-P., L. E. BABCOCK, Y.-L. ZHAO, AND X.-L. ZHANG. In press. Two Cambrian arthropods (Naraoia and Skania) from Guizhou, China, and western North America. Journal of Paleontology.
87. LIN, J.P., S. M. GON III, J. G. GEHLING, L. E. BABCOCK, Y.-L. ZHAO, X.-L. ZHANG, S.-X. HU, J.-L. YUAN, M.-Y. YU, AND J. PENG. 2006. A Parvancorina-like arthropod from the Cambrian of South China. Historical Geology, 18:33-45.
88. LINNAEUS, C. 1758 [1757] Systema naturae per regna tri naturae, sedundum classes, ordines, genera, species cum characteribus differentiis, synonymis, locis. Edito decimal, reformata. Laauretii Salvii, Holmiae, 824 pp.
89. LU, Y.-H. 1940. On the ontogeny and phylogeny of Redlichia intermedia Lu (sp. nov.). Bulletin of the Geological Society of China, 20:333-342.
90. LU, Y.-H. 1950. New species of Redlichia. Geological Review, 15:4-6.
91. LUO, H.-L., Z.-W. JIANG, AND L. TANG. 1996. Stages, zones and correlations of the Lower Cambrian in China. In The Geological Society of China (ed.), Progress in Geology of China - Papers to 30th IGC. China Ocean Press, Beijing.
92. LOU, H.-L., S.-X. HU, S.-S. ZHANG, AND Y.-H. TAO. 1997. New occurrence of the Early Cambrian Chengjiang fauna from Haikou, Kunming, Yunnan Province. Acta Geologica Sinica, 71:97-104. (In Chinese with English summary)
93. LOU, H.-L., S.-X. HU, L.-Z. CHEN, S.-S. ZHANG, AND Y.-H. TAO. 1999. Early Cambrian Chengjiang Fauna from the Kunming Region, China. Yunnan Science and Technology Press, Kunming, 129 p.
94. MATTHEW, G. F. 1899. Studies on Cambrian faunas, no. 3: Upper Cambrian faunas of Mt. Stephen, British Columbia. Transactions of the Royal Society of Canada, Series 2, 5:39-66.
95. MCKINNEY, M. L. AND K. J. MCNAMARA. 1991. Heterochrony: The Evolution of Ontogeny. New York, Plenum Press, 437 p.
96. MELNIKOVA, L. M., D. J. SIVETER, AND M. WILLIAMS. 1997. Cambrian Bradoriida and Phosphatocopida (Arthropoda) of the former Soviet Union. Journal of Micropalaeontology, 16:179-191.
97. MÜLLER, K. J. AND D. WALOSZEK. 1987. Morphology, ontogeny, and life habit of Agnostus pisiformis from the Upper Cambrian of Sweden. Fossils and Strata, 19:1-124.
98. NEDIN, C. 1995. The Emu Bay Shale, a Lower Cambrian fossil Lagerstätte, Kangaroo Island, South Australia. In P. A. Jell (ed.), APC94: Papers from the First Australian Palaeontological Convention. Association of Australasian Palaeontologists Memoir, 18:31-40.
99. NEDIN, C. 1999. Anomalocaris predation on nonmineralized and mineralized trilobites. Geology, 27:987-990.
100. PELECHATY, S. M. 1996. Stratigraphic evidence for the Siberia-Laurentia connection and Early Cambrian rifting. Geology, 24:719-722.
101. PENG, S.-C. AND R. A. ROBISON. 2000. Agnostoid biostratigraphy across the Middle-Upper Cambrian boundary in Hunan, China. Journal of Paleontology, 74(supp. 4):1-104.
102. PETROVICH, R. 2001. Mechanisms of fossilization of the soft-bodied and lightly armored faunas of the Burgess Shale and some other classical localities. American Journal of Science, 301:683-726.
103. POCOCK, K. J. 1964. Estaingia, a new trilobite genus from Lower Cambrian of South Australia. Palaeontology, 7:458-471.
104. POULSON, C. 1967. Fossils from the Lower Cambrian of Bornholm. Matematisk-fysiske Meddelelser, 36:1-48.
105. QIAN, Y. AND S. BENGTSON. 1989. Palaeontology and biostratigraphy of the Early Cambrian Meishucunian Stage in Yunnan Province, South China. Fossils and Strata, 24:1-156.
106. RAFF, R. A. 1996. The Shape of Life: Genes, Development, and the Evolution of Animal Form. University of Chicago Press, Chicago, 544 p.
107. RAMSKÖLD, L. AND G. D. EDGECOMBE. 1996. Trilobite appendage structure - Eoredlichia reconsidered. Alcheringa, 20:269-276.
108. RAMSKÖLD, L., J.-Y. CHEN, G. D. EDGECOMBE, AND G.-Q. ZHOU. 1996. Preservational folds simulating tergite junctions in tegopeltid and naraoiid arthropods. Lethaia, 29:15-20.
109. RAMSKÖLD, L., J.-Y. CHEN, G. D. EDGECOMBE, AND G.-Q. ZHOU. 1997. Cindarella and the arachnate clade Xandarellida (Arthropoda, Early Cambrian) from China. Transactions of the Royal Society of Edinburgh: Earth Science, 88:19-38.
110. RAYMOND, P. E. 1920. Phylogeny of the Arthropoda with especial reference to the trilobites. American Naturalist, 54:398-413.
111. REPINA, L. N. AND O. G. OKUNEVA. 1969. Cambrian arthropods of the Maritime Territory. Palaeontologischeskii Zhurnal, 1969(1):95-103.
112. ROBISON, R. B. 1984. New occurrence of the unusual trilobite Naraoia from the Cambrian of Idaho and Utah. University of Kansas Paleontological Contributions, 112:1-8.
113. ROMINGER, C. 1887. Description of primordial fossils from Mt. Stephens, N. W. Territory of Canada. Proceedings of the Academy of Natural Sciences at Philadelphia, 1887:12-19.
114. RUDKIN, D. M., G. A. YOUNG, R. J. ELIAS, AND E. P. DOBRZANSKI. 2003. The world's biggest trilobite - Isotelus rex new species from the Upper Ordovician of Northern Manitoba, Canada. Journal of Paleontology, 77:99-112.
115. SCHOLTZ, G. 1997. Cleavage, germ band formation and head segmentation: the ground pattern of the Euarthropoda. In R. A. Fortey and R. H. Thomas (eds.), Arthropod Relationships. The Systematics Association, Special Volume, Series 55:317-332. Chapman and Hall, London.
116. SCHRAM, F. R. 1986. Crustacea. Oxford University Press, Oxford, 620 p.
117. SCHWIMMER, D. R. AND W. M. MONTANTE. 2002. Exceptional fossil preservation in the Conasauga Formation, Middle Cambrian of western Georgia. Geological Society of America, Abstracts with Program, 31:65.
118. SHU, D.-G., L. CHEN, J. HAN, AND X.-L. ZHANG. 2001a. An Early Cambrian tunicate from China. Nature, 411:472-473.
119. SHU, D.-G., G. GEYER, L. CHEN, AND X.-L. ZHANG. 1995. Redlichiacean trilobites with preserved soft parts from the Lower Cambrian Chengjiang fauna (South China). In G. Geyer and E. Landing (eds.), Morocco '95 - the Lower Cambrian standard of western Gondwana. Beringeria Special Issue, 2:203-241.
120. SHU, D.-G., S. CONWAY MORRIS, J. HAN, L. CHEN, X.-L. ZHANG, Z.-F. ZHANG, H.-Q. LIU, AND Y. LI, J.-N. LIU. 2001b. Primitive deuterostomes from the Chengjiang Lagerstätte (Lower Cambrian, China). Nature, 414: 419-424.
121. SIEBOLD, C. T. E. AND H. STANNIUS. 1848. Lehruch der verliechenden Anatomie der wirbellosen Tiere. Viet, Berlin.
122. SIMONETTA, A. M. 1970. Studies on non-trilobite arthropods of the Burgess Shale (Middle Cambrian). Palaeontographica Italica, 66:35-45.
123. SIMONETTA, A. M. AND L. DELLE CAVE. 1975. The Cambrian non-trilobite arthropods from the Burgess Shale of British Columbia. A study of their comparative morphology, taxonomy and evolutionary significance. Palaeontographica Italica, 69:1-37.
124. SPEYER, S. E. AND B. D. E. CHATTERTON. 1989. Trilobite larvae and larval ecology. Historical Biology, 3:27-60.
125. STEINER, M., M.-Y. ZHU, Y.-L. ZHAO, AND B.-D. ERDTMANN, 2005. Lower Cambrian Burgess Shale-type fossil associations of South China. Palaeogeography, Palaeoclimatology, Palaeoecology, 220:129-152.
126. STRAUSS, R. E. 1987. On allometry and relative growth in evolutionary studies. Systematic Zoology, 36:72-75.
127. SUN, W.-G. AND X.-G. HOU. 1987a. Early Cambrian medusae from Chengjiang, Yunnan, China. Acta Palaeontologica Sinica, 26:257-271.
128. SUN, W.-G. AND X.-G. HOU. 1987b. Early Cambrian worms from Chengjiang, Yunnan, China. Acta Palaeontologica Sinica, 26:300-305.
129. VANNIER, J. AND J.-Y. CHEN. 2002. Digestive system and feeding mode in Cambrian naraoiid arthropods. Lethaia, 35:107-120.
130. VINTHER, J. AND C. NIELSEN. 2005. The Early Cambrian Halkieria is a molluscan. Zoologica Scripta, 34:81-89.
131. WALCOTT, C. D. 1911. Middle Cambrian Merostomata. Cambrian Geology and Paleontology II. Smithsonian Miscellaneous Collections, 57:17-40.
132. WALCOTT, C.D. 1912. Cambrian geology and paleontology, II, Middle Cambrian Branchiopoda, Malacostraca, Trilobita and Merostomata. Smithsonian Miscellaneous Collections, 57:146-228.
133. WALCOTT, C. D. 1931. Addenda to descriptions of Burgess Shale fossils. Smithsonian Miscellaneous Collections, 85:1-46.
134. WALOSZEK, D. 1993. The Upper Cambrian Rehbachiella and the phylogeny of Branchiopoda and Crustacea. Fossils and Strata, 32:1-202.
135. WALOSZEK, D. 1995. The Upper Cambrian Rehbachiella, its larval development, morphology and significance for the phylogeny of Branchiopoda and Crustacea. Hydrobiologia, 298:1-13.
136. WALOSZEK, D. 2003a. Cambrian 'Orsten'-type preserved arthropods and the phylogeny of Crustacea, p. 69-87. In A. Legakis, S. Sfenthourakis, R. Polymeni, and M. Thessalou-Legaki (eds.), The New Panorama of Animal Evolution. Proceedings of 18th International Congress of Zoology. Pensoft Publishers, Sofia and Moscow.
137. WALOSZEK, D. 2003b. The 'Orsten' window - a three-dimensionally preserved Upper Cambrian meiofauna and its contribution to our understanding of the evolution of Arthropoda. Paleontological Research, 7:71-88.
138. WALOSZEK, D. AND K. J. MÜLLER. 1990. Stem-lineage crustaceans from the Upper Cambrian of Sweden and their bearing upon the monophyletic origin of Crustacea and the position of Agnostus. Lethaia, 23:409-427.
139. WALOSZEK, D. AND K. J. MÜLLER. 1997. Cambrian 'Orsten'-type arthropods and the phylogeny of Crustacea, p. 139-154. In R. A. Fortey and R. H. Thomas (eds.), Arthropod Relationships. The Systematics Association Special Volume Series 55. Chapman and Hall, London.
140. WALOSZEK, D. AND K. J. MÜLLER. 1998. Early arthropod phylogeny in light of the Cambrian "Orsten" fossils, p. 185-231. In G. D. Edgecombe (ed.), Arthropod Fossils and Phylogeny. Columbia University Press, New York.
141. WESTHEIDE, W. AND R. RIEGER. 2004. Spezielle Zoologie, Teil 1: Einzeller und Wirbellose Tiere. Spektrum Akademischer Verlag, Heidelberg and Berlin, 919 p.
142. WHITTINGTON, H. B. 1959. Ontogeny of Trilobita, p. 127-144. In R. C. Moore (ed.), Treatise on Invertebrate Paleontology, Pt. O, Arthropoda 1. Geological Society of America and University of Kansas Press, Lawrence.
143. WHITTINGTON, H. B. 1971. Redescription of Marrella splendens (Trilobitoidea) from the Burgess Shale, Middle Cambrian, British Columbia. Bulletin of the Geological Survey, Canada, 209:1-24.
144. WHITTINGTON, H. B. 1975. Trilobites with appendages from the Middle Cambrian, Burgess Shale, British Columbia. Fossils and Strata, 4:97-136.
145. WHITTINGTON, H. B. 1977. The Middle Cambrian trilobite Naraoia, Burgess Shale, British Columbia. Philosophical Transactions of the Royal Society of London, B, 280:409-443.
146. WHITTINGTON, H. B. 1985a. The Burgess Shale. Yale University Press, New Haven, 151 p.
147. WHITTINGTON, H. B. 1985b. Tegopelte gigas, a second soft-bodied trilobite from the Burgess Shale, Middle Cambrian, British Columbia. Journal of Paleontology, 59:1251-1274.
148. WHITTINGTON, H. B. 1992. Trilobites. The Boydell Press, Woodbridge, 145 p., 120 pls.
149. WHITTINGTON, H. B. 1997. Morphology of the exoskeleton, p. 1-86. In R. L. Kaesler (ed.), Treatise on Invertebrate Paleontology, Pt. O, Arthropoda 1. Trilobita (Revised). Geological Society of America and University of Kansas Press, Lawrence.
150. WILLS, M. A. 1997. A phylogeny of recent and fossil Crustacea derived from morphological characters, p. 189-209. In R. A. Fortey and R. H. Thomas (eds.), Arthropod Relationships. The Systematics Association Special Volume Series 55. Chapman and Hall, London.
151. WILLS, M. A., D. E. G. BRIGGS, R. A. FORTEY, M. WILKINSON, AND P. H. A. SNEATH. 1998. An arthropod phylogeny based on fossil and recent taxa, p. 33-105. In G. D. Edgecombe (ed.), Arthropod Fossils and Phylogeny. Columbia University Press, New York.
152. YAGER, J. AND J. H. CARPENTER. 1999. Speleonectes epilimnius new species (Remipedia, Speleonectidae) from surface water of an Anchialine Cave on San Salvador Island, Bahamas. Crustaceana, 72:965-977.
153. YUAN, J.-L. AND Y.-L. ZHAO. 1999. Subdivision and correlation of Lower Cambrian in southwest China, with a discussion of the age of Early Cambrian series biota. Acta Palaeontologica Sinica, 38(supplement):116-131. (In Chinese with English summary)
154. YUAN, J.-L., Y.-L. ZHAO, Y. LI, AND Y.-Z. HUANG. 2002. Trilobite fauna of the Kaili Formation (uppermost Lower Cambrian-lower Middle Cambrian) from southeastern Guizhou, South China. Shanghai Science and Technology Press, Shanghai, 423 p.
155. ZHANG, W.-T. 1951. Trilobites from the Shipai Shale and their stratigraphical significance. Huai Shuan (Newsletter of the Geological Society of China), 2:10.
156. ZHANG, W.-T. AND X.-G. HOU. 1985. Preliminary notes on the occurrence of the unusual trilobite Naraoia in Asia. Acta Palaeontologica Sinica, 24:591-595.
157. ZHANG, W.-T., Y.-H. LU, Z.-L. ZHU, Y. QIAN, H.-Z. LIN, Z.-Y. ZHOU, S.-G. ZHANG, AND J.-L. YUAN. 1980. Cambrian trilobite faunas of Southwest China. Palaeontographica Sinica, 159(16):1-497. (In Chinese with English summary)
158. ZHANG, X.-L. 1995. Naraoiids from the Chengjiang Lagerstätte. Unpublished MS thesis, Northwest University at Xian (China), 51 p., 20 pls. (In Chinese with English abstract)
159. ZHANG, X.-L. AND D.-G. SHU. 1996. Preservation and post-mortem decay of naraoiids from the Chengjiang Lagerstätte. Journal of Northwest University, 26(supp.):454-458. (In Chinese with English abstract)
160. ZHANG, X.-L., D.-G. SHU, AND G. GEYER. 1995. A new genus of naraoiids, p. 30. In J.-Y. Chen, G. Edgecombe, and L. Ramsköld (eds.), International Cambrian Explosion Symposium. (April, 1995, Nanjing, Program and Abstract).
161. ZHANG, X.-L., D.-G. SHU, Y. LI, AND J. HAN. 2001. New sites of Chengjiang fossils: Crucial windows on the Cambrian explosion. Journal of the Geological Society of London, 158:211-218.
162. ZHANG, X.-L., J. HAN, Z.-F. ZHANG, H.-Q. LIU, AND D.-G. SHU. 2003. Reconsideration of the supposed naraoiid larva from the Early Cambrian Chengjiang Lagerstätte, South China. Palaeontology, 46:447-465.
163. ZHANG, X.-L., J. HAN, Z.-F. ZHANG, H.-Q. LIU, AND D.-G. SHU. 2004. Redescription of the Chengjiang arthropod Squamacula clypeata Hou and Bergström from the Lower Cambrian, southwest China. Palaeontology, 47: 1-13.
164. ZHAO, Y.-L., J.-L. YUAN, M.-Y. ZHU, X.-L. YANG, AND J. PENG. 2003. The occurrence of genus Marrella (Trilobitoidae) in Asia. Progress in Natural Science, 13:708-701.
165. ZHAO, Y.-L., J.-L. YUAN, M.-Y. ZHU, R.-G. YANG, Q.-J. GUO, Y. QIAN, Y.-Z. HUANG, AND Y. PAN. 1999. A progress report on research on the Early Middle Cambrian Kaili biota, Guizhou, P. R. C. Acta Palaeontologica Sinica, 38(supp.):1-15. (In Chinese with English summary)
166. ZHU, M.-Y. 1997. Trace fossils of Yunnan. Bulletin of National Museum of Natural Science, 10:275-312.
167. ZHU, M.-Y., L. E. BABCOCK, AND M. STEINER. 2005. Fossilization modes in the Chengjiang Lagerstatte (Cambrian of China): Testing the roles of organic preservation and diagenetic alteration in exceptional preservation. Palaeogeography, Palaeoclimatology, Palaeoecology, 220:31-46.
168. ZHU, M.-Y., J.-M. ZHANG, AND G.-X. LI. 2001. Sedimentary environments of the Early Cambrian Chengjiang biota: Sedimentology of the Yu'anshan Formation in Chengjiang County, eastern Yunnan, p. 80-105. In M.-Y. Zhu, H. Van Iten, S.-C. Peng, and G.-X. Li (eds.), The Cambrian of South China. Acta Palaeontologica Sinica, 40 (supplement).
169. ZHU, M.-Y., H. VAN ITEN, R. S. COX, Y.-L. ZHAO, AND B. D. ERDTMANN. 2000. Occurrence of Byronia Matthew and Sphenothallus Hall in the Lower Cambrian of China. Paläontologische Zeitschrift, 74:227-238.
170. ZHURAVLEV, A. Y. 1995. Preliminary suggestions on the global Early Cambrian Zonation, p. 147-160. In G. Geyer and E. Landing (eds.), Morocco '95: The Lower-Middle Cambrian standard of western Gondwana. Beringeria, Special Issue 2.