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Volumn 36, Issue 3, 2007, Pages 271-280

Molecular phylogeny of pontobdelline leeches and their place in the descent of fish leeches (Hirudinea, Piscicolidae)

Author keywords

[No Author keywords available]

Indexed keywords

CHONDRICHTHYES; HIRUDINEA; HIRUDINIDA; OSTEICHTHYES; OXYTONOSTOMA; PISCICOLIDAE; PONTOBDELLA; PONTOBDELLA MURICATA; PONTOBDELLINAE;

EID: 34147183890     PISSN: 03003256     EISSN: 14636409     Source Type: Journal    
DOI: 10.1111/j.1463-6409.2007.00279.x     Document Type: Article
Times cited : (31)

References (53)
  • 1
    • 0033174344 scopus 로고    scopus 로고
    • Higher level relationships of leeches (Annelida: Clitellata: Euhirudinea) based on morphology and gene sequences
    • Apakupakul, K., Siddall, M. E. & Burreson, E. M. (1999). Higher level relationships of leeches (Annelida: Clitellata: Euhirudinea) based on morphology and gene sequences. Molecular Phylogenetics and Evolution, 12, 350-359.
    • (1999) Molecular Phylogenetics and Evolution , vol.12 , pp. 350-359
    • Apakupakul, K.1    Siddall, M.E.2    Burreson, E.M.3
  • 2
    • 0347300499 scopus 로고    scopus 로고
    • Arhynchobdellida (Annelida: Oligochaeta: Hirudinida): phylogenetic relationships and evolution
    • Borda, E. & Siddall, M. E. (2004a). Arhynchobdellida (Annelida: Oligochaeta: Hirudinida): phylogenetic relationships and evolution. Molecular Phylogenetics and Evolution, 30, 213-225.
    • (2004) Molecular Phylogenetics and Evolution , vol.30 , pp. 213-225
    • Borda, E.1    Siddall, M.E.2
  • 3
    • 22544459513 scopus 로고    scopus 로고
    • Review of the evolution of life history strategies and phylogeny of the Hirudinida (Annelida: Oligochaeta)
    • Borda, E. & Siddall, M. E. (2004b). Review of the evolution of life history strategies and phylogeny of the Hirudinida (Annelida: Oligochaeta). Lauterbornia, 52, 5-25.
    • (2004) Lauterbornia , vol.52 , pp. 5-25
    • Borda, E.1    Siddall, M.E.2
  • 4
    • 34147137414 scopus 로고    scopus 로고
    • Edgar, R. C, 2004a August, MUSCLE version 3.5. Available via
    • Edgar, R. C. (2004a August). MUSCLE version 3.5. Available via http:// www.drive5.com/muscle/.
  • 5
    • 3042666256 scopus 로고    scopus 로고
    • MUSCLE: Multiple sequence alignment with high accuracy and high throughput
    • Edgar, R. C. (2004b). MUSCLE: multiple sequence alignment with high accuracy and high throughput. Nucleic Acids Research, 32, 1792-1797.
    • (2004) Nucleic Acids Research , vol.32 , pp. 1792-1797
    • Edgar, R.C.1
  • 7
    • 3242688934 scopus 로고
    • Neravnomernost' tempov evolyutsii system organov i printsipy sistematiki ryb'ikh piyavok (Hirudinea, Piscicolidae). [Unevenness of rates of evolution of organ systems and principles of the systematics of fish leeches (Hirudinea, Piscicolidae)]
    • Kiev
    • Epshtein, V. M. (1970). Neravnomernost' tempov evolyutsii system organov i printsipy sistematiki ryb'ikh piyavok (Hirudinea, Piscicolidae). [Unevenness of rates of evolution of organ systems and principles of the systematics of fish leeches (Hirudinea, Piscicolidae)]. In Voprosy Morskoy Parazitologii. Materialy Vsesoyuznogo Simpoziuma Po Parazitam I Boleznyam Morskikh Zhivotnykh Sevastopol' 1970 (pp. 140-142). Kiev.
    • (1970) Voprosy Morskoy Parazitologii. Materialy Vsesoyuznogo Simpoziuma Po Parazitam I Boleznyam Morskikh Zhivotnykh Sevastopol' 1970 , pp. 140-142
    • Epshtein, V.M.1
  • 8
    • 34147166860 scopus 로고
    • Sistematicheskoe polozhenie antarkticheskoy piyavki Pontobdella rugosa Moore (Piscicolidae). [Systematic status of the Antarctic leech Pontobdella rugosa Moore (Piscicolidae)]
    • Epshtein, V. M. (1972). Sistematicheskoe polozhenie antarkticheskoy piyavki Pontobdella rugosa Moore (Piscicolidae). [Systematic status of the Antarctic leech Pontobdella rugosa Moore (Piscicolidae)]. Zoologicheskiy Zhurn, 51, 1142-1146.
    • (1972) Zoologicheskiy Zhurn , vol.51 , pp. 1142-1146
    • Epshtein, V.M.1
  • 9
    • 0015627177 scopus 로고
    • O sistematicheskom polozhenii, obraze zhizni i rasprostranenii Levinsenia rectangulata. [On the taxonomic position, mode of life and geographic distribution of Levinsenia rectangulata]
    • Epshtein, V. M. (1973). O sistematicheskom polozhenii, obraze zhizni i rasprostranenii Levinsenia rectangulata. [On the taxonomic position, mode of life and geographic distribution of Levinsenia rectangulata]. Parazitologiya, 7, 286-292.
    • (1973) Parazitologiya , vol.7 , pp. 286-292
    • Epshtein, V.M.1
  • 10
    • 34147177372 scopus 로고
    • Novye svedeniya ob elementakh tropicheskogo proiskhozhdeniya v faune ryb'ikh piyavok Antarktiki. [New data on tropical origin elements in Antarctic piscicolid leech fauna]
    • Moscow
    • Epshtein, V. M. (1974). Novye svedeniya ob elementakh tropicheskogo proiskhozhdeniya v faune ryb'ikh piyavok Antarktiki. [New data on tropical origin elements in Antarctic piscicolid leech fauna]. In The Sixth All-Union Symposium Diseases and Parasites of Marine Animals (pp. 301-305). Moscow.
    • (1974) The Sixth All-Union Symposium Diseases and Parasites of Marine Animals , pp. 301-305
    • Epshtein, V.M.1
  • 11
    • 34147142684 scopus 로고    scopus 로고
    • Epshtein, V. M. (1984). Shchetinkonosnye, cherepash'i i ryb'i piyavki mirovoy fauny (sistemnyi podhod k klassifikatsii i filogenii). Avtoreferat Dissertatsii Na Soiskanie Uchenoy Stepeni Doktora Biologicheskikh Nauk. [Chaetiferous, turtle and fish leeches of the world fauna (system approach to classification and phylogeny). Abstract of the Dissertation for the degree of Doctor of Biological Sciences]. Leningrad: Zoologicheskiy Institut AN SSSR.
    • Epshtein, V. M. (1984). Shchetinkonosnye, cherepash'i i ryb'i piyavki mirovoy fauny (sistemnyi podhod k klassifikatsii i filogenii). Avtoreferat Dissertatsii Na Soiskanie Uchenoy Stepeni Doktora Biologicheskikh Nauk. [Chaetiferous, turtle and fish leeches of the world fauna (system approach to classification and phylogeny). Abstract of the Dissertation for the degree of Doctor of Biological Sciences]. Leningrad: Zoologicheskiy Institut AN SSSR.
  • 12
    • 34147164780 scopus 로고    scopus 로고
    • Epshtein, V. M. (1985). Infrastruktura piyavok sem. Piscicolidae. [Infrastructure of the leeches of the family Piscicolidae]. In VIII Vsesoyuznoe Soveshchanie Po Parazitam I Boleznyam Ryb. Tezisy Dokladov. Astrahan', Aprel' 1985 G (pp. 156-157). Leningrad: Nauka.
    • Epshtein, V. M. (1985). Infrastruktura piyavok sem. Piscicolidae. [Infrastructure of the leeches of the family Piscicolidae]. In VIII Vsesoyuznoe Soveshchanie Po Parazitam I Boleznyam Ryb. Tezisy Dokladov. Astrahan', Aprel' 1985 G (pp. 156-157). Leningrad: Nauka.
  • 13
    • 84879391058 scopus 로고
    • Tip kol 'chatye chervi - Annelida. [The Phylum Annelids - Annelida]
    • O. N. Bauer Ed, Identification Key to Parasites of the Freshwater Fishes of the USSR, Leningrad: Nauka
    • Epshtein, V. M. (1987). Tip kol 'chatye chervi - Annelida. [The Phylum Annelids - Annelida]. In O. N. Bauer (Ed.) Opredelitel' Parazitov Presnovodnykh Ryb Fauny SSSR. [Identification Key to Parasites of the Freshwater Fishes of the USSR], Vol. 3 (pp. 340-372). Leningrad: Nauka.
    • (1987) Opredelitel' Parazitov Presnovodnykh Ryb Fauny SSSR , vol.3 , pp. 340-372
    • Epshtein, V.M.1
  • 14
    • 34147154551 scopus 로고
    • O stroenii, sistematicheskom polozhenii i rasprostranenii antarkticheskoy piyavki Pontobdella biannulata Moore, 1957. [On the structure, systematic status and distribution of the Antarctic leech Pontobdella biannulata Moore, 1957]
    • Epshtein, V. M. & Okun', S. V. (1991). O stroenii, sistematicheskom polozhenii i rasprostranenii antarkticheskoy piyavki Pontobdella biannulata Moore, 1957. [On the structure, systematic status and distribution of the Antarctic leech Pontobdella biannulata Moore, 1957]. Informatsionnyi Byulleten' Sovetskoi Antarcticheskoi Ekspedicii, 116, 99-103.
    • (1991) Informatsionnyi Byulleten' Sovetskoi Antarcticheskoi Ekspedicii , vol.116 , pp. 99-103
    • Epshtein, V.M.1    Okun', S.V.2
  • 15
    • 34147165799 scopus 로고    scopus 로고
    • Geograficheskoe rasprostranenie i khozyaeva piyavok roda Notostomum (Hirudinea: Piscicolidae). [The geographical distribution and hosts of the Notostomum leeches (Hirudinea: Piscicolidae)]
    • Epshtein, V. M. & Utevsky, S. Y. (1996). Geograficheskoe rasprostranenie i khozyaeva piyavok roda Notostomum (Hirudinea: Piscicolidae). [The geographical distribution and hosts of the Notostomum leeches (Hirudinea: Piscicolidae)]. Vestnik Zoologii, 30, 26-31.
    • (1996) Vestnik Zoologii , vol.30 , pp. 26-31
    • Epshtein, V.M.1    Utevsky, S.Y.2
  • 16
    • 3242673453 scopus 로고
    • The system of fish leeches (Hirudinea: Piscicolidae)
    • Epshtein, V. M., Utevsky, A. Y. & Utevsky, S. Y. (1994). The system of fish leeches (Hirudinea: Piscicolidae). Genus, 5, 401-409.
    • (1994) Genus , vol.5 , pp. 401-409
    • Epshtein, V.M.1    Utevsky, A.Y.2    Utevsky, S.Y.3
  • 18
    • 34147098217 scopus 로고
    • Notice regarding the ova of the Pontobdella muricata, Lam
    • Grant, R. E. (1827). Notice regarding the ova of the Pontobdella muricata, Lam. Edinburgh Journal of Science, 7 (1), 160-161.
    • (1827) Edinburgh Journal of Science , vol.7 , Issue.1 , pp. 160-161
    • Grant, R.E.1
  • 19
    • 0242578620 scopus 로고    scopus 로고
    • A simple, fast, and accurate algorithm to estimate large phylogenies by maximum likelihood
    • Guindon, S. & Gascuel, O. (2003). A simple, fast, and accurate algorithm to estimate large phylogenies by maximum likelihood. Systematic Biology, 52, 696-704.
    • (2003) Systematic Biology , vol.52 , pp. 696-704
    • Guindon, S.1    Gascuel, O.2
  • 20
    • 1642356002 scopus 로고    scopus 로고
    • Congruence versus phylogenetic accuracy: Revisiting the incongruence length difference test
    • Hipp, A. L., Hall, J. C. & Sytsma, K. J. (2004). Congruence versus phylogenetic accuracy: revisiting the incongruence length difference test. Systematic Biology, 53, 81-89.
    • (2004) Systematic Biology , vol.53 , pp. 81-89
    • Hipp, A.L.1    Hall, J.C.2    Sytsma, K.J.3
  • 21
    • 0034849408 scopus 로고    scopus 로고
    • MRBAYES: Bayesian inference of phylogeny
    • Huelsenbeck, J. P. & Ronquist, F. R. (2001). MRBAYES: Bayesian inference of phylogeny. Bioinformatics, 17, 754-755.
    • (2001) Bioinformatics , vol.17 , pp. 754-755
    • Huelsenbeck, J.P.1    Ronquist, F.R.2
  • 22
    • 34147093947 scopus 로고    scopus 로고
    • Keane, T, December, 2004, MODELGENERATOR version 0.83. Available via
    • Keane, T. (December, 2004). MODELGENERATOR version 0.83. Available via http://bioinf.nuim.ie/software/modelgenerator/.
  • 23
    • 33645816587 scopus 로고    scopus 로고
    • Assessment of methods for amino acid matrix selection and their use on empirical data shows that ad hoc assumptions for choice of matrix are not justified
    • Keane, T. M., Creevey, C. J., Pentony, M. M., Naughton, T. J. & McInerney, O. J. (2006). Assessment of methods for amino acid matrix selection and their use on empirical data shows that ad hoc assumptions for choice of matrix are not justified. BMC Evolutionary Biology, 6, 29.
    • (2006) BMC Evolutionary Biology , vol.6 , pp. 29
    • Keane, T.M.1    Creevey, C.J.2    Pentony, M.M.3    Naughton, T.J.4    McInerney, O.J.5
  • 25
    • 0032730391 scopus 로고    scopus 로고
    • Phylogeny of the leech family Glossiphoniidae based on mitochondrial gene sequences and morphological data
    • Light, J. E. & Siddall, M. E. (1999). Phylogeny of the leech family Glossiphoniidae based on mitochondrial gene sequences and morphological data. Journal of Parasitology, 85, 815-823.
    • (1999) Journal of Parasitology , vol.85 , pp. 815-823
    • Light, J.E.1    Siddall, M.E.2
  • 27
    • 30744470239 scopus 로고
    • Pontobdellinae (Hirudinea: Piscicolidae) in the British Museum (Natural History) with a revue of the subfamily
    • Llewellyn, L. C. (1966). Pontobdellinae (Hirudinea: Piscicolidae) in the British Museum (Natural History) with a revue of the subfamily. Bulletin of the British Museum (Natural History). Zoology, 14, 341-439.
    • (1966) Bulletin of the British Museum (Natural History). Zoology , vol.14 , pp. 341-439
    • Llewellyn, L.C.1
  • 28
    • 34147126658 scopus 로고    scopus 로고
    • Maddison, W. P. & Maddison, D. R, 2006, MESQUITE: A Modular System for Evolutionary Analysis, Version 1.11. Available via
    • Maddison, W. P. & Maddison, D. R. (2006). MESQUITE: A Modular System for Evolutionary Analysis, Version 1.11. Available via http:// mesquiteproject.org.
  • 31
    • 34147166344 scopus 로고
    • Australasian Antarctic Expedition 1911-14. Scientific Reports (C), 10
    • Moore, J. P. (1938). Leeches (Hirudinea). Australasian Antarctic Expedition 1911-14. Scientific Reports (C), 10, 5-16.
    • (1938) , pp. 5-16
    • Moore, J.P.1
  • 32
    • 34147152499 scopus 로고    scopus 로고
    • Moore, J. P. (1957). Hirudinea. B.A.N.Z. Antarctic Research expedition. Reports - Series B. (Zoology and Botany), 6, 102-105.
    • Moore, J. P. (1957). Hirudinea. B.A.N.Z. Antarctic Research expedition. Reports - Series B. (Zoology and Botany), 6, 102-105.
  • 33
    • 0011080676 scopus 로고
    • Leeches (Hirudinea) from Alaskan and adjacent waters
    • Moore, J. P. & Meyer, M. C. (1951). Leeches (Hirudinea) from Alaskan and adjacent waters. The Wasmann Journal of Biology, 9, 11-77.
    • (1951) The Wasmann Journal of Biology , vol.9 , pp. 11-77
    • Moore, J.P.1    Meyer, M.C.2
  • 34
    • 2142651467 scopus 로고
    • A rare heavy infestation of Levinsenia rectangulata (Hirudinea) on a Walleye Pollock Theragra chalcogramma in Hokkaido waters
    • Nagasawa, K. (1988). A rare heavy infestation of Levinsenia rectangulata (Hirudinea) on a Walleye Pollock Theragra chalcogramma in Hokkaido waters. Nippon Suisan Gakkaishi, 54, 319.
    • (1988) Nippon Suisan Gakkaishi , vol.54 , pp. 319
    • Nagasawa, K.1
  • 35
    • 33646428355 scopus 로고    scopus 로고
    • Higher phylogeny of zygaenid moths (Insecta: Lepidoptera) inferred from nuclear and mitochondrial sequence data and the evolution of larval cuticular cavities for chemical defense
    • Niehuis, O., Yen, S.-H., Naumann, C. M. & Misof, B. (2006). Higher phylogeny of zygaenid moths (Insecta: Lepidoptera) inferred from nuclear and mitochondrial sequence data and the evolution of larval cuticular cavities for chemical defense. Molecular Phylogenetics and Evolution, 39, 812-829.
    • (2006) Molecular Phylogenetics and Evolution , vol.39 , pp. 812-829
    • Niehuis, O.1    Yen, S.-H.2    Naumann, C.M.3    Misof, B.4
  • 36
    • 33745032085 scopus 로고    scopus 로고
    • Further problems with the incongruence length difference test: 'hypercongruence' effect and multiple comparisons
    • Ramírez, M. J. (2006). Further problems with the incongruence length difference test: 'hypercongruence' effect and multiple comparisons. Cladistics, 22, 289-295.
    • (2006) Cladistics , vol.22 , pp. 289-295
    • Ramírez, M.J.1
  • 40
    • 0036623258 scopus 로고    scopus 로고
    • An approximately unbiased test of phylogenetic tree selection
    • Shimodaira, H. (2002). An approximately unbiased test of phylogenetic tree selection. Systematic Biology, 46, 492-508.
    • (2002) Systematic Biology , vol.46 , pp. 492-508
    • Shimodaira, H.1
  • 41
    • 85142141485 scopus 로고    scopus 로고
    • Shimodaira, H. & Hasegawa, M. (2001). CONSEL: for assessing the confidence of phylogenetic tree selection. Bioinformatics, 17, 1246-1247.
    • Shimodaira, H. & Hasegawa, M. (2001). CONSEL: for assessing the confidence of phylogenetic tree selection. Bioinformatics, 17, 1246-1247.
  • 42
    • 0031389775 scopus 로고    scopus 로고
    • Prior agreement: Arbitration or arbitrary?
    • Siddall, M. E. (1997). Prior agreement: arbitration or arbitrary? Systematic Biology, 46, 765-769.
    • (1997) Systematic Biology , vol.46 , pp. 765-769
    • Siddall, M.E.1
  • 44
    • 0037227822 scopus 로고    scopus 로고
    • Phylogeny and revision of the leech genus Helobdella (Glossiphoniidae) based on mitochondrial gene sequences and morphological data and a special consideration of the triserialis complex
    • Siddall, M. E. & Borda, E. (2003). Phylogeny and revision of the leech genus Helobdella (Glossiphoniidae) based on mitochondrial gene sequences and morphological data and a special consideration of the triserialis complex. Zoologica Scripta, 32, 23-33.
    • (2003) Zoologica Scripta , vol.32 , pp. 23-33
    • Siddall, M.E.1    Borda, E.2
  • 45
    • 0031989263 scopus 로고    scopus 로고
    • Phylogeny of leeches (Hirudinea) based on mitochondrial cytochrome c oxidase subunit
    • Siddall, M. E. & Burreson, E. M. (1998). Phylogeny of leeches (Hirudinea) based on mitochondrial cytochrome c oxidase subunit. Molecular Phylogenetics and Evolution, 9, 156-162.
    • (1998) Molecular Phylogenetics and Evolution , vol.9 , pp. 156-162
    • Siddall, M.E.1    Burreson, E.M.2
  • 46
    • 34147125643 scopus 로고    scopus 로고
    • Swofford, D. L. (2003). PAUP* - Phylogenetic Analysis Using Parsimony (*and other methods), Version 4. [Computer software and manual]. Champaign, Illinois: Illinois Natural History Survey. Sinauer Associates, Sunderland, MA.
    • Swofford, D. L. (2003). PAUP* - Phylogenetic Analysis Using Parsimony (*and other methods), Version 4. [Computer software and manual]. Champaign, Illinois: Illinois Natural History Survey. Sinauer Associates, Sunderland, MA.
  • 47
    • 0000927082 scopus 로고    scopus 로고
    • Molecular phylogeny of leeches: Congruence of nuclear and mitochondrial rDNA data sets and the origin of bloodsucking
    • Trontelj, P., Sket, B. & Steinbrück, G. (1999). Molecular phylogeny of leeches: congruence of nuclear and mitochondrial rDNA data sets and the origin of bloodsucking. Journal of Zoological Systematics and Evolutionary Research, 37, 141-147.
    • (1999) Journal of Zoological Systematics and Evolutionary Research , vol.37 , pp. 141-147
    • Trontelj, P.1    Sket, B.2    Steinbrück, G.3
  • 48
    • 15944393218 scopus 로고    scopus 로고
    • Celebrity with a neglected taxonomy: Molecular systematics of the medicinal leech (genus Hirudo)
    • Trontelj, P. & Utevsky, S. Y. (2005). Celebrity with a neglected taxonomy: molecular systematics of the medicinal leech (genus Hirudo). Molecular Phylogenetics and Evolution, 34, 616-624.
    • (2005) Molecular Phylogenetics and Evolution , vol.34 , pp. 616-624
    • Trontelj, P.1    Utevsky, S.Y.2
  • 49
    • 34147139163 scopus 로고    scopus 로고
    • A new Species of Piscicolid Leeches (Hirudinea, Piscicolidae) from Antarctic Seas
    • Utevsky, A. Y. (1997). A new Species of Piscicolid Leeches (Hirudinea, Piscicolidae) from Antarctic Seas. Vestnik zoologii, 31 (1-2), 17-24.
    • (1997) Vestnik zoologii , vol.31 , Issue.1-2 , pp. 17-24
    • Utevsky, A.Y.1
  • 50
    • 3242702554 scopus 로고    scopus 로고
    • Phylogenetic relationships of fish leeches (Hirudinea, Piscicolidae) based on mitochondrial DNA sequences and morphological data
    • Utevsky, S. Y. & Trontelj, P. (2004). Phylogenetic relationships of fish leeches (Hirudinea, Piscicolidae) based on mitochondrial DNA sequences and morphological data. Zoologica Scripta, 33, 375-385.
    • (2004) Zoologica Scripta , vol.33 , pp. 375-385
    • Utevsky, S.Y.1    Trontelj, P.2
  • 51
    • 33750288604 scopus 로고    scopus 로고
    • Phylogeny of the fish leeches (Oligochaeta, Hirudinida, Piscicolidae) based on nuclear and mitochondrial genes and morphology
    • Williams, J. I. & Burreson, E. M. (2006). Phylogeny of the fish leeches (Oligochaeta, Hirudinida, Piscicolidae) based on nuclear and mitochondrial genes and morphology. Zoologica Scripta, 35, 627-639.
    • (2006) Zoologica Scripta , vol.35 , pp. 627-639
    • Williams, J.I.1    Burreson, E.M.2
  • 52
    • 0035527350 scopus 로고    scopus 로고
    • Failure of the ILD to determine data combinability for slow loris phylogeny
    • Yoder, A. D., Irwin, J. A. & Payseur, B. A. (2001). Failure of the ILD to determine data combinability for slow loris phylogeny. Systematic Biology, 50, 408-424.
    • (2001) Systematic Biology , vol.50 , pp. 408-424
    • Yoder, A.D.1    Irwin, J.A.2    Payseur, B.A.3
  • 53
    • 33750980115 scopus 로고    scopus 로고
    • Phylogeny of the cave shrimp Troglocaris: Evidence of a young connection between Balkans and Caucasus
    • Zakšek, V., Sket, B. & Trontelj, P. (2007). Phylogeny of the cave shrimp Troglocaris: evidence of a young connection between Balkans and Caucasus. Molecular Phylogenetics and Evolution, 42, 223-235.
    • (2007) Molecular Phylogenetics and Evolution , vol.42 , pp. 223-235
    • Zakšek, V.1    Sket, B.2    Trontelj, P.3


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