Taxonomic affinities and evolutionary history of the Early Pleistocene Hominids of Java: Dentognathic evidence

American Journal of Physical Anthropology

Volumn 128, Issue 4, 2005, Pages 709-726

  Kaifu, Yousuke ^a   Baba, Hisao ^a,b   Aziz, Fachroel ^c   Indriati, Etty ^d   Schrenk, Friedemann ^e   Jacob, Teuku ^d  

^a DEPARTMENT OF BOTANY   (Japan)

^b UNIVERSITY OF TOKYO   (Japan)

^c Geological Research and Development Centre Bandung   (Indonesia)

^d GADJAH MADA UNIVERSITY   (Indonesia)

^e SENCKENBERG RESEARCH INSTITUTE   (Germany)

Author keywords

H. erectus; Hominid evolution; Java; Mandible; Meganthropus; Pleistocene; Teeth

Indexed keywords

AFRICA; ARTICLE; CHRONOLOGY; DENTISTRY; EVOLUTION; GENETIC VARIABILITY; HISTORY OF MEDICINE; HOMINID; HUMAN; INDONESIA; LOWER PLEISTOCENE; PLEISTOCENE; TAXONOMIC IDENTIFICATION;

ADULT; ANIMALS; CHILD; EVOLUTION; FEMALE; FOSSILS; HISTORY, ANCIENT; HOMINIDAE; HUMANS; INDONESIA; MALE; MANDIBLE; PALEODONTOLOGY; SEX CHARACTERISTICS; TOOTH;

EID: 27144511731     PISSN: 00029483     EISSN: None     Source Type: Journal    
DOI: 10.1002/ajpa.10425     Document Type: Article

References (142)

1. Andrews P. 1984. An alternative interpretation of characters used to define Homo erectus. Courier Forsch Senckenberg 69:167-175.
2. Antón SC. 2002. Evolutionary significance of cranial variation in Asian Homo erectus. Am J Phys Anthropol 118:301-323.
3. Anton SC, Franzen JL. 1997. The occipital torus and developmental age of Sangiran-3. J Hum Evol 33:599-610.
4. Antón SC, Indriati E. 2002. Earliest Pleistocene Homo in Asia: craniodental comparisons of Dmanisi and Sangiran. Am J Phys Anthropol [Suppl] 34:38.
5. Arambourg C. 1963. Le gisement de Ternifine II. L'Atlanthropus mauritanicus. Arch Inst Paleontol Hum Mem 32:37-190.
6. Arif J, Kaifu Y, Baba H, Suparka ME, Zaim Y, Setoguchi T. 2002. Preliminary observation of a new cranium of Homo erectus (Tjg-1993.05) from Sangiran, Central Jawa. Anthropol Sci 110:165-177.
7. Asfaw B, Beyene Y, Suwa G, Walter RC, White TD, WoldeGabriel G, Yemane T. 1992. The earliest Acheulean from Konso-Gardula. Nature 360:732-735.
8. Asfaw B, Gilbert WH, Beyene Y, Hart WK, Renne PR, WoldeGabriel G, Vrba ES, White TD. 2002. Remains of Homo erectus from Bouri, Middle Awash, Ethiopia. Nature 416:317-320.
9. Aziz F. 2001. Hominid fossils housed at the Geological Research and Development Center, Bandung, Indonesia. In: Indriati E, editor. A scientific life: papers in honor of Prof. Dr. T. Jacob. Yogyakarta: Bigraf Publishing, p 53-66.
10. Baba H, Aziz F. 1991. Hominid skull fragments found in 1979 from Sangiran, Central Java. Bull Natl Sci Mus Tokyo Ser D 17:1-8.
11. Baba H, Aziz F, Narasaki S, Sudijono, Kaifu Y, Suprijo A, Hyodo M, Susanto E E, Jacob T. 2000. A new hominid incisor from Sangiran, Central Java. J Hum Evol 38:855-862.
12. Baba H, Aziz F, Kaifu Y, Suwa G, Kono RT, Jacob T. 2003. New Homo erectus calvarium from the Pleistocene of Java. Science 299:1384-1388.
13. Behrents RG. 1985. Growth in the aging craniofacial skeleton. In: Carlson DS, Ribbons KA, editors. Monograph 7. Craniofacial growth series. Ann Arbor: University of Michigan, Center for Human Growth and Development.
14. Bermúdez de Castro JM, Nicolas ME. 1995. Posterior dental size reduction in hominids: the Atapuerca evidence. Am J Phys Anthropol 96:335-356.
15. Bermúdez de Castro JM, Rosas A, Nicolás ME. 1999. Dental remains from Atapuerca-TD6 (Gran Dolina site, Burgos, Spain). J Hum Evol 37:523-566.
16. Blumenschine RJ, Peters CR, Masao FT, Clarke RJ, Deino AL, Hay RL, Swisher CC, Stanistreet IG, Ashley GM, McHenry LJ, Sikes NE, Van der Merwe NJ, Tactikos JC, Cushing AE, Deocampo DM, Njau JK, Ebert JI. 2003. Late Pliocene Homo and hominid land use from western Olduvai Gorge, Tanzania. Science 299:1217-1221.
17. Bräuer G. 1994. How different are Asian and African Homo erectus? Courier Forsch Senckenberg 171:301-318.
18. Bräuer G, Mbua E. 1992. Homo erectus features used in cladistics and their variability in Asian and African hominids. J Hum Evol 22:79-108.
19. Bräuer G, Schultz M. 1996. The morphological affinities of the Plio-Pleistocene mandible from Dmanisi, Georgia. J Hum Evol 30:445-481.
20. Brown B, Walker A. 1993. The dentition. In: Walker A, Leakey R, editors. The Nariokotome Homo erectus skeleton. Cambridge, MA: Harvard Univesity Press, p 161-192.
21. Brown B, Brown F, Walker A. 2001. New hominids from the Lake Turkana basin, Kenya. J Hum Evol 41:29-44.
22. Brown P. 2001. Chinese Middle Pleistocene hominids and modern human origins in east Asia. In: Barham L, Brown KR, editors. Human roots-Africa and Asia in the Middle Pleistocene. Bristol: Western Academic & Specialist Press, p 135-145.
23. Clarke RJ. 1994. The significance of Swartkrans Homo to the Homo erectus problem. Courier Forsch Senckenberg 171:185-193.
24. Day MH, Leakey REF. 1973. New evidence of the genus Homo from East Rudolf, Kenya. Am J Phys Anthropol. 39:341-354.
25. Day MH, Leakey REF, Walker A, Wood BA. 1976. New hominids from East Turkana, Kenya. Am J Phys Anthropol 45:369-436.
26. Delson E, Harvati K, Reddy D, Marcus LF, Mowbray K, Sawyer GJ, Jacob T, Márquez S. 2001. The Sambungmacan 3 Homo erectus calvaria: a comparative morphometric and morphological analysis. Anat Rec 262:380-397.
27. Dennell RW, Rendell H, Hailwood E. 1988. Late Pliocene artifacts from northern Pakistan. Curr Anthropol 29:495-498.
28. De Vos J, Sondaar PY. 1994. Dating hominid sites in Indonesia. Science 266:1726-1727.
29. De Vos J, Sondaar PY. Van den Bergh GD, Aziz F. 1994. The Homo bearing deposits of Java and its ecological context. Courier Forsch Senckenberg 171:129-140.
30. Donnelly SM, Kramer A. 1999. Testing for multiple species in fossil samples: an evaluation and comparison of tests for equal relative variation. Am J Phys Anthropol 108:507-529.
31. Durband AC. 2003. A re-examination of purported "Meganthropus" cranial fragments. Am J Phys Anthropol [Suppl] 36:91.
32. Etler DA, Crummett TL, Wolpoff MH. 2001. Longuppo: early Homo colonizer or Late Pliocene Lufengpithecus survivor in south China? Hum Evol 16:1-12.
33. Franzen JL. 1985a. What is "Pithucanthropus dubius Koenigswald, 1950"? In: Delson E, editor. Ancestors: the hard evidence. New York: Alan R. Liss. p 221-226.
34. Franzen JL. 1985b. Asian australopithecines? In: Tobias PV, editor. Hominid evolution: past, present, and future. New York: Alan R. Liss. p 255-263.
35. Gabunia L, Vekua A. 1995. A Plio-Pleistocene hominid from Dmanisi, East Georgia, Caucasus. Nature 373:509-512.
36. Gabunia L, Vekua A, Lordkipanidze D, Swisher CC III, Ferring R, Justus A, Nioradze M, Tvalchrelidze M, Antón S, Bosinski G, Jöris O, De Lumley M-A, Majsuradze G, Mouskhelishvili A. 2000. Earliest Pleistocene hominid cranial remains from Dmanisi, Republic of Georgia: taxonomy, geological setting, and age. Science 288:1019-1025.
37. Gabunia L, De Lemley M-A, Vekua A, Lordkipanidze D, De Lumley H. 2002. Découverte d'un nouvel hominidé à Dmanissi (Transcaucasie, Géorgie). C R Palevol 1:243-253.
38. Grimaud-Hervé D, Widianto H, Jacob T, 2000. Two new human fossil remains discovered in Sangira (Central Java, Indonesia). Acta Anthropol Sin [Suppl] 19:41-45.
39. Grine FE. 1984. Comparison of the deciduous dentitions of Africa and Asian hominids. Courier Forsch Senckenberg 69:69-82.
40. Grine FE, Franzen JL. 1994. Fossil hominid teeth from the Sangiran Dome (Java, Indonesia). Courier Forsch Senckenberg 171:75-103.
41. Groves CP. 1989. A theory of human and primate evolution. Oxford: Oxford University Press.
42. Hou Y, Potts R, Yuan B, Guo Z, Deino A, Wang W, Clark J, Xie G, Huang W. 2000. Mid-Pleistocene Acheulean-like stone technology of the Bose Basin, South China. Science 287:1622-1626.
43. Howell FC. 1978. Hominidae. In: Maglio VJ, Cooke HBS, editors. Evolution of African mammals. Cambrage: Harvard University Press, p 154-248.
44. Huang W, Ciochon R, Gu Y, Lnrick R, Fang Q, Schwarcz H, Yonge C, De Vos J, Rink W. 1995. Early Homo and associated artifacts from Asia. Nature 378:275-278.
45. Huffman OF. 2001. Geologic context and age of the Perning/Mojokerto Homo erectus, East Java. J Hum Evol 40:353-362.
46. Hyodo M, Watanabe N, Sunata W, Susanto EE, Wahyono H. 1993. Magnetostratigraphy of hominid fossil bearing formations in Sangiran and Mojokerto, Java. Anthropol Sci 101:157-186.
47. Hyodo M, Nakaya H, Urabe A, Saegusa H, Shunrong X, Jiyun Y, Xuepin J. 2002. Paleomagnetic dates of hominid remains from Yuanmou, China, and other Asian sites. J Hum Evol 43:27-41.
48. Indriati E. 2004. Indonesian fossil hominid discoveries from 1889-2002: problems and catalogue. Natn Sci Mus Monogr. 24:163-177.
49. Itihara M, Sudijono, Kadar D, Shibasaki T, Kuniai H, Yoshikawa S, Aziz F, Soeradi T, Wikarno, Kadar AP, Hashibuan F, Kagemori Y. 1985. Geology and stratigraphy of the Sangiran area. In: Watanabe N, Kadar D, editors. Quaternary geology of the hominid fossil bearing formations in Java. Geo Res Dev Centre. Spec Publ. 4:11-43.
50. Jacob T. 1973. Palaeoanthropological discoveries in Indonesia with special reference to the finds of the last two decades. J Hum Evol 2:473-485.
51. Jacob T. 1976. Early populations in the Indonesian region. In: Kirk RL, Thorne AG, editors. The origin of the Australians. Canberra: Australian Institute of Aboriginal Studies, p 81-93.
52. Jacob T. 1980. The Pithecanthropus of Indonesia: phenotype, genetics and ecology. In: Konigsson L-K, editor. Current argument on early man. Oxford: Pergamon Press, p 170-179.
53. Kaifu Y. 1999. Changes in alveolar prognathism and anterior teeth protrusion. Anthropol Sci 107:3-24.
54. Kaifu Y. 2000a. Tooth wear and compensatory modification of the anterior dentoalveolar complex in humans. Am J Phys Anthropol 111:369-392.
55. Kaifu Y. 2000b. Was extensive tooth wear normal in our ancestors? A preliminary examination in the genus Homo. Anthropol Sci 108:371-385.
56. Kaifu Y, Baba H, Aziz F, Schrenk F, Jacob T, Indriati E, Arif J. 2003. Primitive dentognathic morphology of Javanese Homo crectus. Am J Phys Anthropol [Suppl] 36:125.
57. Kaifu Y, Aziz F, Baba H. 2005. Hominid mandibular remains from Sangiran: 1952-1986 collection. Am J Phys Anthropol. In press.
58. Kennedy GE. 1991. On the autapomorphic traits of Homo erectus. J Hum Evol 20:375-412.
59. Kimbel WH. 1995. Hominid speciation and Pliocene climatic changes. In: Vrba ES, Denton GH, Partridge TC, Burckle LH, editors. Paleoclimate and evolution with emphasis on human origins. New Haven: Yale University Press, p 425-437.
60. Klein RG. 1999. The human career. 2nd ed. Chicago: University of Chicago Press.
61. Kramer A. 1989. The evolutionary and taxonomic affinities of the Sangiran mandibles of central Java, Indonesia. Ph.D. dissertation, University of Michigan.
62. Kramer A. 1993. Human taxonomic diversity in the Pleistocene: does Homo erectus represent multiple hominid species? Am J Phys Anthropol 91:161-171.
63. Kramer A. 1994. A critical analysis of claims for the existence of Southeast Asian australopithecines. J Hum Evol 26:3-21.
64. Kramer A, Konigsberg LW. 1994. The phyletic position of Sangiran 6 as determined by multivariate analyses. Courier Forsch Senckenberg 171:105-114.
65. Krantz GS. 1975. An explanation for the diastema of Javan erectus Skull IV. In: Tuttle RH, editor. Paleoanthropology, morphology and paleoecplogy. The Hague: Mouton, p 361-372.
66. Langbroek M, Roebroeks W. 2000. Extraterrestrial evidence on the age of the hominids from Java. J Hum Evol 38:595-600.
67. Larick R, Ciochon RL. 1996. The African emergence and early Asian dispersals of the genus Homo. Am Sci 84:538-551.
68. Larick R, Ciochon RL, Zaim Y, Sudijono, Suminto, Rizal Y, Aziz F. 2000. Lithostratigraphic context for Kin-1993.05-SNJ, a fossil colobine maxilla from Jolotingkir, Sangiran Dome. Int J Primatol 21:731-759.
69. 39 Ar ages for Bapang Formation hominins. Central Jawa. Indonesia. Proc Natl Acad Sci USA 98:4866-4871.
70. Leakey REF, Walker A. 1985. Further hominids from the Plio-Pleistocene of Koobi Fora, Kenya. Am J Phys Anthropol 67:135-163.
71. Leakey REF, Wood BA. 1973. New evidence of the genus Homo from east Rudolf, Kenya. II. Am J Phys Anthropol 39:355-368.
72. Leakey REF, Wood BA. 1974. New evidence of the genus Homo from east Rudolf, Kenya (IV). Am J Phys Anthropol 41:37-244.
73. Le Gros Clark WE. 1964. The fossil evidence of human evolution: an introduction to the study of paleoanthropology, 2nd ed. Chicago: University of Chicago Press.
74. Leigh SR. 1992. Cranial capacity evolution in Homo erectus and early Homo sapiens. Am J Phys Anthropol 87:1-13.
75. Lovejoy CO. 1970. The taxonomic status of the "Meganthropus" mandibular fragments from the Djetis Beds of Java. Man 5:228-236.
76. Mann A. 1981. The significance of the Sinanthropus casts, and some paleodemographic notes. In: Sigmon BA, Cybulski JS, editors. Homo erectus papers in honor of Davidson Black. Toronto: University of Toronto Press, p 41-62.
77. Matsu'ura S. 1982. A chronological framing for the Sangiran hominids: fundamental study by the fluorine dating method. Bull Natl Sci Mus Tokyo Ser D 8:1-53.
78. Matsu'ura S, Kondo S, Aziz F, Watanabe N. 1995. Chronology of four hominid mandibular fossils newly found from Sangiran and their possible evolutionary implications. In: Baba H, editor. Report of research project supported by grant-in-aid from Japanese Ministry of Education, Science, Sports and Culture, project no. 04454034. p 37-50.
79. Miller JMA. 2000. Craniofacial variation in Homo habilis: an analysis of the evidence for multiple species. Am J Phys Anthropol 112:103-128.
80. Orban-Segebarth R, Procureur F. 1983. Tooth size of Meganthropus palaeojavanicus: an analysis of distances between some fossil hominids and a modern human population. J Hum Evol 12:711-720.
81. Pope GG. 1988. Recent advances in Far Eastern paleoanthropology. Annu Rev Anthropol 17:43-77.
82. Qian F, Li Q, Wu P, Yuan S, Xing R, Chen H, Zhang H. 1991. Lower Pleistocene, Yuanmou Formation. In: Quarternary geology and paleoanthropology of Yuanmou, Yunnan, China. Beijing: Science Press, p 17-50 [in Chinese with English abstract, p 194-195].
83. Rightmire GP. 1980. Middle Pleistocene hominids from Olduvai Gorge, northern Tanzania. Am J Phys Anthropol 53:225-241.
84. Rightmire GP. 1990. The evolution of Homo erectus: comparative anatomical studies of an extinct human species. Cambridge: Cambridge University Press.
85. Rightmire GP. 1998. Evidence from facial morphology for similarity of Asian and African representatives of Homo erectus. Am J Phys Anthropol 106:61-85.
86. Robinson JT. 1953. Meganthropus, australopithecines, and hominids. Am J Phys Anthropol 11:1-38.
87. Robinson JT. 1955. Further remarks on the relationship between "Meganthropus" and australopithecines. Am J Phys Anthropol 13:429-446.
88. Robinson JT. 1956. The dentition of the australopithecinae. Pretoria: Transv Mus Mem 9.
89. Rosas A, Bermúdez de Castro JM. 1998. On the taxonomic affinities of the Dmanisi mandible (Georgia). Am J Phys Anthropol 107:145-162.
90. Santa Luca AP. 1980. The Ngandong fossil hominids: a comparative study for a Far Eastern Homo erectus group. Yale Univ Publ Anthropol 78:1-175.
91. Sartono S. 1974. Observations on a newly discovered jaw of Pithecanthropus modjokertensis from the Lower Pleistocene of Sangiran, Central Java. Proc K Ned Akad Wet Amsterdam 77:26-31.
92. Sartono S. 1982. Sagittal cresting in Meganthropus palaeojavanicus von Koenigswald. Mod Quat Res SE Asia 7:201-210.
93. Sartono S, Grimaud-Hervé D. 1983. Les pariétaux de l'hominidé Sangiran 31. Anthropologie (Paris) 87:465-468.
94. Sawyer GJ, Tattersall I. 1995. A new reconstruction of the skull of Homo erectus from Zhoukoudian, China. In: Gibert J, Sánchez F, Gibert L, Ribot F, editors. The hominids and their environment during the Lower and Middle Pleistocene of Eurasia. Proceedings of the International Conference of Human Palaeontology. Orce: Museo de Prehistoria Paleontologiá. p 277-286.
95. Schwartz JH, Tattersall I. 1996. Whose teeth? Nature 381:201-202.
96. Schwartz JH, Tattersall I. 2000. What constitutes Homo erectus? Acta Anthropol Sin [Suppl] 19:18-22.
97. Smith BH. 1984. Patterns of molar wear in hunter-gatherers and agriculturists. Am J Phys Anthropol 63:39-56.
98. Soeradi T, Shibasaki T, Kadar D, Sudijono, Itihara M, Kumai H, Hayashi T, Furuyama K, Aziz F, Siagian H, Furutani M, Suminto, Yoshikawa S. 1985. Geology and Stratigraphy of the Trinil area. In: Watanabe N, Kadar D, editors. Quaternary geology of the hominid fossil bearing formations in Java. Geo Res Dev Centre, Spec Publ. p 45-48.
99. Sudijono. 1985. The Grenzbank zone, a key marker bed. In: Watanabe N, Kadar D, editors. Quaternary geology of the hominid fossil bearing formations in Java. Geo Res Dev Centre, Spec Publ. 4:135-151.
100. Suwa G. 1990. A comparative analysis of hominid dental remains from the Sungura and Usno Formations, Omo Valley, Ethiopia. Ph.D. dissertation, University of California at Berkeley.
101. Suwa G, White TD, Howell FC. 1996. Mandibular postcanine dentition from the Sungra Formation, Ethiopia: crown morphology, taxonomic allocations, and Pilo-Pleistocene hominid evolution. Am J Phys Anthropol 101:247-282.
102. Swisher CC III, Curtis GH, Jacob T, Getty AG, Suprijo A, Widiasmoro. 1994. Age of the earliest known hominids in Java, Indonesia. Science 263:1118-1121.
103. Tattersall I. 1986. Species recognition in human paleontology. J Hum Evol 15:165-176.
104. Tobias PV. 1991. Olduvai Gorge, 4: the skulls, endocasts and teeth of Homo habilis. Cambridge: Cambridge University Press.
105. Tobias PV, von Koenigswald GHR. 1964. A comparison between the Olduvai hominines and those of Java and some implications for hominid phylogeny. Nature 204:515-518.
106. Tyler DE. 1991. A taxonomy of Javan hominid mandibles. Hum Evol 6:401-420.
107. Tyler DE. 2001. Two new "Meganthropus" mandibles from Java. Hum Evol 16:151-158.
108. Tyler DE, Sartono S. 2001. A new Homo erectus cranium from Sangiran. Hum Evol 16:13-25.
109. Tyler DE, Krantz GS, Sartono S. 1995a. The taxonomic status of the "Meganthropus" cranial (Sangiran 31) and the "Meganthropus" occipital fragment III. In: Bower JRF, Sartono S, editors. Evolution and ecology of Homo erectus. Leiden: Pithecanthropus Centennial Foundation, Leiden University, p 189-201.
110. Tyler DE, Sartono S, Krantz GS. 1995b. "Meganthropus" cranial material from Java, "Meganthropus" I: an announcement. In: Bower JRF, Sartono S, editors. Evolution and ecology of Homo erectus. Leiden: Pithecanthropus Centennial Foundation, Leiden University, p 221-224.
111. Vekua A, Lordkipanidze D, Rightmire GP, Agusti J, Ferring R, Maisuradze G, Mouskhelishivili A, Nioradze M, Ponce de Leon M, Tappen M, Tvalchrelidze M, Zollikofer C. 2002. A new skull of early Homo from Dmanisi, Georgia. Science 297:85-89.
112. Von Koenigswald GHR. 1950. Fossil hominids from the Lower Pleistocene of Java. Rep 18th Internat Geol Congr 1948, London, Part 9:59-61.
113. Von Koenigswald GHR. 1968. Observations upon two Pithecanthropus mandibles from Sangiran, Central Java. K Ned Akad Wet Amsterdam Ser B 71:99-107.
114. Von Koenigswald GHR. 1973. Austraropithecus, Meganthropus and Ramapithecus. J Hum Evol 2:487-491.
115. Walker A, Leakey R. 1993. The skull. In: Walker A, Leakey R, editors. The Nariokotome Homo erectus skeleton. Cambridge, MA: Harvard University Press, p 63-94.
116. Wang Q. 1996. Assignment of Longgupo Cave hominid incisor from Wushan. Acta Anthropol Sin 15:320-323.
117. Watanabe N, Kadar D, editors. 1985. Quaternary geology of the hominid fossil bearing formations in Java. Geo Res Dev Centre, Spec Publ.
118. Weidenreich F. 1936. The mandibles of Sinanthropus pekinensis: a comparative study. Paleontol Sin Ser D 7:1-162.
119. Weidenreich F. 1937. The dentition of Sinanthropus pekinensis: a comparative odontography of the hominids. Paleontol Sin New Ser D 1:1-180.
120. Weidenreich F. 1943. The skull of Sinanthropus pekinensis: a comparative study on a primitive hominid skull. Paleontol Sin New Ser D 10:1-484.
121. Weidenreich F. 1945. Giant early man from Java and South China. Anthropol Pap Am Mus Nat Hist 40:1-134.
122. Weidenreich F. 1951. Morphology of Solo man. Anthropol Pap Am Mus Nat Hist 43:205-290.
123. White TD. 1977. The anterior mandibular corpus of early African Hominidae: functional significance of shape and size. Ph.D. dissertation, University of Michigan.
124. White TD, Johanson DC. 1982. Pliocene hominid mandibles from the Hadar Formation, Ethiopia: 1974-1977 collections. Am J Phys Anthropol 57:501-544.
125. White TD, Johanson DC, Kimbel WH. 1981. Australopithecus africanus: its phyletic position reconsidered. S Afr J Sci 77:445-470.
126. White TD, Suwa G, Asfaw B. 2000. Jaws and teeth of Australopithecus afarensis from Maka, Middle Awash, Ethiopia. Am J Phys Anthropol 111:45-68.
127. Widianto H. 1993. Unité et diversité des hominides fossiles de Java: presentation de restes humains fossiles inédits. Ph.D. dissertation, Museum National d'Histoire Naturelle à l'Institut de Paleontologie Humaine.
128. Widianto H, Sémah A-M, Djubiantono T, Sémah F. 1994. A tentative reconstruction of the cranial human remains of Hanoman 1 from Bukuran, Sangiran (Central Java). Courier Forsch Senckenberg 171:47-59.
129. Wolpoff MH. 1984. Evolution in Homo erectus: the question of stasis. Paleobiology 10:389-406.
130. Wolpoff MH. 1999. Paleoanthropology, 2nd ed. Boston: McGraw-Hill.
131. Woo J. 1964. Mandible of the Sinanthropus-type discovered at Lantian, Shensi-Sinanthropus lantianensis. Sci Sin 5:801-811.
132. Wood BA. 1984. The origin of Homo erectus. Courier Forsch Senckenberg 69:99-111.
133. Wood BA. 1991. Koobi Fora research project, volume 4. Hominid cranial remains from Koobi Fora. Oxford: Clarendon Press.
134. Wood BA. 1994. Taxonomy and evolutionary relationship of Homo erectus. Courier Forsch Senckenberg 171:159-165.
135. Wood BA, van Noten FL. 1986. Preliminary observation on the BK 8518 mandible from Baringo, Kenya. Am J Phys Anthropol 69:117-127.
136. Wood BA, Abbott SA, Uytterschaut H. 1988. Analysis of the dental morphology of Plio-Pleistocene hominids IV. Mandibular postcanine root morphology. J Anat 156:107-139.
137. Wu M. 1983. Homo erectus from Hexian, Anhui found in 1981. Acta Anthropol Sin 2:109-115.
138. Wu R, Dong X. 1982. Preliminary study of Homo erectus remains from Hexian, Anhui. Acta Anthropol Sin 1:2-13.
139. Wu X. 2000. Longgupo hominoid mandible belongs to ape. Acta Anthropol Sin 19:1-10 [in Chinese with English abstract].
140. Wu X, Poirier FE. 1995. Human evolution in China: a metric description of the fossils and a review of the sites. Oxford: Oxford University Press.
141. Zhang Y. 1984. The "Australopithecus" of West Hubei and some Early Pleistocene hominids of Indonesia. Acta Anthropol Sin 3:85-92.
142. Zhu RX, Hoffuman KA, Potts R, Deng CL, Pan YX, Guo B, Shi CD, Guo ZT, Yuan BY, Hou YM, Huang WW. 2001. Earliest presence of humans in northeast Asia. Nature 413:413-417.