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This project was funded by two grants to M.H.T. and M.R.S. by the German Research Foundation (DFG) and a University College London Graduate School grant to MAM. We thank the government of Kenya and the Kenya Wildlife Service for research permits and support; L. Aiello, A. Bergner, P. Blisniuk, M. Collard, P. deMenocal, M. Dühnforth, F. Gasse, F. Grine, and G. Muchemi for inspiring discussions; and S. Higgins, S. Kabingu, T. Schlüter, and M. Ibs-von Seht for logistical support.
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