메뉴 건너뛰기




Volumn 17, Issue 3, 2005, Pages 691-704

Corona, a member of the class III homeodomain leucine zipper gene family in Arabidopsis, regulates stem cell specification and organogenesis

Author keywords

[No Author keywords available]

Indexed keywords

CELLS; CLONING; MATHEMATICAL MODELS; MUTAGENESIS; PLANTS (BOTANY); PROTEINS;

EID: 22144498191     PISSN: 10404651     EISSN: None     Source Type: Journal    
DOI: 10.1105/tpc.104.026179     Document Type: Article
Times cited : (137)

References (53)
  • 1
    • 0034649566 scopus 로고    scopus 로고
    • Analysis of the genome sequence of the flowering plant Arabidopsis thaliana
    • Arabidopsis Genome Initiative (2000). Analysis of the genome sequence of the flowering plant Arabidopsis thaliana. Nature 408, 796-815.
    • (2000) Nature , vol.408 , pp. 796-815
  • 2
    • 0029562071 scopus 로고
    • The expression of the Athb-8 homeobox gene is restricted to provascular cells in Arabidopsis thaliana
    • Baima, S., Nobili, F., Sessa, G., Lucchetti, S., Ruberti, I., and Morelli, G. (1995). The expression of the Athb-8 homeobox gene is restricted to provascular cells in Arabidopsis thaliana. Development 121, 4171-4182.
    • (1995) Development , vol.121 , pp. 4171-4182
    • Baima, S.1    Nobili, F.2    Sessa, G.3    Lucchetti, S.4    Ruberti, I.5    Morelli, G.6
  • 3
    • 0034984476 scopus 로고    scopus 로고
    • The Arabidopsis ATHB-8 HD-ZIP protein acts as a differentiation-promoting transcription factor of the vascular meristems
    • Baima, S., Possenti, M., Matteucci, A., Wisman, E., Altamura, M.M., Ruberti, I., and Morelli, G. (2001). The Arabidopsis ATHB-8 HD-ZIP protein acts as a differentiation-promoting transcription factor of the vascular meristems. Plant Physiol. 126, 643-655.
    • (2001) Plant Physiol. , vol.126 , pp. 643-655
    • Baima, S.1    Possenti, M.2    Matteucci, A.3    Wisman, E.4    Altamura, M.M.5    Ruberti, I.6    Morelli, G.7
  • 4
    • 0027379437 scopus 로고
    • Formation of the shoot apical meristem in Arabidopsis thaliana: An analysis of development in the wild type and shoot meristemless mutant
    • Barton, M.K., and Poethig, R.S. (1993). Formation of the shoot apical meristem in Arabidopsis thaliana: An analysis of development in the wild type and shoot meristemless mutant. Development 119, 823-831.
    • (1993) Development , vol.119 , pp. 823-831
    • Barton, M.K.1    Poethig, R.S.2
  • 5
    • 0031624102 scopus 로고    scopus 로고
    • In planta Agrobacterium-mediated transformation of adult Arabidopsis thaliana plants by vacuum infiltration
    • Bechtold, N., and Pelletier, G. (1998). In planta Agrobacterium-mediated transformation of adult Arabidopsis thaliana plants by vacuum infiltration. Methods. Mol. Biol. 82, 259-266.
    • (1998) Methods. Mol. Biol. , vol.82 , pp. 259-266
    • Bechtold, N.1    Pelletier, G.2
  • 6
    • 0028055069 scopus 로고
    • Assignment of 30 microsatellite loci to the linkage map of Arabidopsis
    • Bell, C.J., and Ecker, J.R. (1994). Assignment of 30 microsatellite loci to the linkage map of Arabidopsis. Genomics 19, 137-144.
    • (1994) Genomics , vol.19 , pp. 137-144
    • Bell, C.J.1    Ecker, J.R.2
  • 8
    • 0034725925 scopus 로고    scopus 로고
    • Dependence of stem cell fate in Arabidopsis on a feedback loop regulated by CLV3 activity
    • Brand, U., Fletcher, J.C., Hobe, M., Meyerowitz, E.M., and Simon, R. (2000). Dependence of stem cell fate in Arabidopsis on a feedback loop regulated by CLV3 activity. Science 289, 617-619.
    • (2000) Science , vol.289 , pp. 617-619
    • Brand, U.1    Fletcher, J.C.2    Hobe, M.3    Meyerowitz, E.M.4    Simon, R.5
  • 9
    • 0035983621 scopus 로고    scopus 로고
    • Regulation of CLV3 expression by two homeobox genes in Arabidopsis
    • Brand, U., Grunewald, M., Hobe, M., and Simon, R. (2002). Regulation of CLV3 expression by two homeobox genes in Arabidopsis. Plant Physiol. 129, 565-575.
    • (2002) Plant Physiol. , vol.129 , pp. 565-575
    • Brand, U.1    Grunewald, M.2    Hobe, M.3    Simon, R.4
  • 11
    • 0036337173 scopus 로고    scopus 로고
    • ASYMMETRIC LEAVES1 reveals knox gene redundancy in Arabidopsis
    • Byrne, M.E., Simorowski, J., and Martienssen, R.A. (2002). ASYMMETRIC LEAVES1 reveals knox gene redundancy in Arabidopsis. Development 129, 1957-1965.
    • (2002) Development , vol.129 , pp. 1957-1965
    • Byrne, M.E.1    Simorowski, J.2    Martienssen, R.A.3
  • 12
    • 0029942983 scopus 로고    scopus 로고
    • The CLAVATA and SHOOTMERISTEMLESS loci competitively regulate meristem activity in Arabidopsis
    • Clark, S.E., Jacobsen, S.E., Levin, J., and Meyerowitz, E.M. (1996). The CLAVATA and SHOOTMERISTEMLESS loci competitively regulate meristem activity in Arabidopsis. Development 122, 1567-1575.
    • (1996) Development , vol.122 , pp. 1567-1575
    • Clark, S.E.1    Jacobsen, S.E.2    Levin, J.3    Meyerowitz, E.M.4
  • 13
    • 0027426931 scopus 로고
    • CLAVATA1, a regulator of meristem and flower development in Arabidopsis
    • Clark, S.E., Running, M.P., and Meyerowitz, E.M. (1993). CLAVATA1, a regulator of meristem and flower development in Arabidopsis. Development 119, 397-418.
    • (1993) Development , vol.119 , pp. 397-418
    • Clark, S.E.1    Running, M.P.2    Meyerowitz, E.M.3
  • 14
    • 0028887377 scopus 로고
    • CLAVATA3 is a specific regulator of shoot and floral meristem development affecting the same process as CLAVATA1
    • Clark, S.E., Running, M.P., and Meyerowitz, E.M. (1995). CLAVATA3 is a specific regulator of shoot and floral meristem development affecting the same process as CLAVATA1. Development 121, 2057-2067.
    • (1995) Development , vol.121 , pp. 2057-2067
    • Clark, S.E.1    Running, M.P.2    Meyerowitz, E.M.3
  • 15
    • 0030729445 scopus 로고    scopus 로고
    • The CLAVATA1 gene encodes a putative receptor kinase that controls shoot and floral meristem size in Arabidopsis
    • Clark, S.E., Williams, R.E., and Meyerowitz, E.M. (1997). The CLAVATA1 gene encodes a putative receptor kinase that controls shoot and floral meristem size in Arabidopsis. Cell 8, 575-585.
    • (1997) Cell , vol.8 , pp. 575-585
    • Clark, S.E.1    Williams, R.E.2    Meyerowitz, E.M.3
  • 16
    • 0032447801 scopus 로고    scopus 로고
    • Floral dip: A simplified method for Agrobacterium-mediated transformation of Arabidopsis thaliana
    • Clough, S.J., and Bent, A.F. (1998). Floral dip: A simplified method for Agrobacterium-mediated transformation of Arabidopsis thaliana. Plant J. 16, 735-743.
    • (1998) Plant J. , vol.16 , pp. 735-743
    • Clough, S.J.1    Bent, A.F.2
  • 18
    • 0038366834 scopus 로고    scopus 로고
    • CLAVATA1 dominant-negative alleles reveal functional overlap between multiple receptor kinases that regulate meristem and organ development
    • Dievart, A., Dalal, M., Tax, F.E., Lacey, A.D., Huttly, A., Li, J., and Clark, S.E. (2003). CLAVATA1 dominant-negative alleles reveal functional overlap between multiple receptor kinases that regulate meristem and organ development. Plant Cell 15, 1198-1211.
    • (2003) Plant Cell , vol.15 , pp. 1198-1211
    • Dievart, A.1    Dalal, M.2    Tax, F.E.3    Lacey, A.D.4    Huttly, A.5    Li, J.6    Clark, S.E.7
  • 20
    • 0030454898 scopus 로고    scopus 로고
    • The SHOOT MERISTEMLESS gene is required for maintenance of undifferentiated cells in Arabidopsis shoot and floral meristems and acts at a different regulatory level than the meristem genes WUSCHEL and ZWILLE
    • Endrizzi, K., Moussain, B., Haecker, A., Levin, J.Z., and Laux, T. (1996). The SHOOT MERISTEMLESS gene is required for maintenance of undifferentiated cells in Arabidopsis shoot and floral meristems and acts at a different regulatory level than the meristem genes WUSCHEL and ZWILLE. Plant J. 10, 967-979.
    • (1996) Plant J. , vol.10 , pp. 967-979
    • Endrizzi, K.1    Moussain, B.2    Haecker, A.3    Levin, J.Z.4    Laux, T.5
  • 21
    • 0033583173 scopus 로고    scopus 로고
    • Signaling of cell fate decisions by CLAVATA3 in Arabidopsis shoot meristems
    • Fletcher, J.C., Brand, U., Running, M.P., Simon, R., and Meyerowitz, E.M. (1999). Signaling of cell fate decisions by CLAVATA3 in Arabidopsis shoot meristems. Science 283, 1911-1914.
    • (1999) Science , vol.283 , pp. 1911-1914
    • Fletcher, J.C.1    Brand, U.2    Running, M.P.3    Simon, R.4    Meyerowitz, E.M.5
  • 22
    • 4243678192 scopus 로고    scopus 로고
    • WUSCHEL signaling functions in interregional communication during Arabidopsis ovule development
    • Groß-Hardt, R., Lenhard, M., and Laux, T. (2002). WUSCHEL signaling functions in interregional communication during Arabidopsis ovule development. Genes Dev. 16, 1129-1138.
    • (2002) Genes Dev. , vol.16 , pp. 1129-1138
    • Groß-Hardt, R.1    Lenhard, M.2    Laux, T.3
  • 23
    • 1242352556 scopus 로고    scopus 로고
    • Expression dynamics of WOX genes mark cell fate decisions during early embryonic patterning in Arabidopsis thaliana
    • Haecker, A., Groß-Hardt, R., Geiges, B., Sarkar, A., Breuninger, H., Herrmann, M., and Laux, T. (2004). Expression dynamics of WOX genes mark cell fate decisions during early embryonic patterning in Arabidopsis thaliana. Development 131, 657-668.
    • (2004) Development , vol.131 , pp. 657-668
    • Haecker, A.1    Groß-Hardt, R.2    Geiges, B.3    Sarkar, A.4    Breuninger, H.5    Herrmann, M.6    Laux, T.7
  • 24
    • 0001282586 scopus 로고
    • In-situ hybridisation in plants
    • D.J. Bowles, S.J. Gurr, and M. McPherson, eds (Oxford: Oxford University Press)
    • Jackson, D. (1991). In-situ hybridisation in plants. Molecular Plant Pathology: A Practical Approach, D.J. Bowles, S.J. Gurr, and M. McPherson, eds (Oxford: Oxford University Press), pp. 163-174.
    • (1991) Molecular Plant Pathology: A Practical Approach , pp. 163-174
    • Jackson, D.1
  • 26
    • 0033213721 scopus 로고    scopus 로고
    • The Arabidopsis CLAVATA2 gene encodes a receptor-like protein required for the stability of the CLAVATA1 receptor-like-kinase
    • Jeong, S., Trotochaud, A.E., and Clark, S.E. (1999). The Arabidopsis CLAVATA2 gene encodes a receptor-like protein required for the stability of the CLAVATA1 receptor-like-kinase. Plant Cell 11, 1925-1933.
    • (1999) Plant Cell , vol.11 , pp. 1925-1933
    • Jeong, S.1    Trotochaud, A.E.2    Clark, S.E.3
  • 27
    • 0030087979 scopus 로고    scopus 로고
    • The AINTEGUMENTA gene of Arabidopsis required for ovule and female gametophyte development is related to the floral homeotic gene APETALA2
    • Klucher, M.K., Chow, H., Reiser, L., and Fischer, R.L. (1996). The AINTEGUMENTA gene of Arabidopsis required for ovule and female gametophyte development is related to the floral homeotic gene APETALA2. Plant Cell 8, 137-153.
    • (1996) Plant Cell , vol.8 , pp. 137-153
    • Klucher, M.K.1    Chow, H.2    Reiser, L.3    Fischer, R.L.4
  • 28
    • 0027650566 scopus 로고
    • A procedure for mapping Arabidopsis mutations using co-dominant ecotype-specific PCR-based markers
    • Konieczny, A., and Ausubel, F.M. (1993). A procedure for mapping Arabidopsis mutations using co-dominant ecotype-specific PCR-based markers. Plant J. 4, 403-410.
    • (1993) Plant J. , vol.4 , pp. 403-410
    • Konieczny, A.1    Ausubel, F.M.2
  • 29
    • 0030030901 scopus 로고    scopus 로고
    • The WUSCHEL gene is required for shoot and floral meristem integrity in Arabidopsis
    • Laux, T., Mayer, K.F.X., Berger, J., and Jürgens, G. (1996). The WUSCHEL gene is required for shoot and floral meristem integrity in Arabidopsis. Development 122, 87-96.
    • (1996) Development , vol.122 , pp. 87-96
    • Laux, T.1    Mayer, K.F.X.2    Berger, J.3    Jürgens, G.4
  • 30
    • 0036336801 scopus 로고    scopus 로고
    • The WUSCHEL and SHOOTMERISTEMLESS genes fulfill complementary roles in Arabidopsis shoot meristem regulation
    • Lenhard, M., Jürgens, G., and Laux, T. (2002). The WUSCHEL and SHOOTMERISTEMLESS genes fulfill complementary roles in Arabidopsis shoot meristem regulation. Development 129, 3195-3206.
    • (2002) Development , vol.129 , pp. 3195-3206
    • Lenhard, M.1    Jürgens, G.2    Laux, T.3
  • 31
    • 0042093777 scopus 로고    scopus 로고
    • Stem cell homeostasis in the Arabidopsis shoot meristem is regulated by intercellular movement of CLAVATA3 and its sequestration by CLAVATA1
    • Lenhard, M., and Laux, T. (2003). Stem cell homeostasis in the Arabidopsis shoot meristem is regulated by intercellular movement of CLAVATA3 and its sequestration by CLAVATA1. Development 130, 3163-3173.
    • (2003) Development , vol.130 , pp. 3163-3173
    • Lenhard, M.1    Laux, T.2
  • 32
    • 0030070190 scopus 로고    scopus 로고
    • A member of the KNOTTED class of homeodomain proteins encoded by the STM gene in Arabidopsis
    • Long, J.A., Moan, E.I., Medford, J.I., and Barton, M.K. (1996). A member of the KNOTTED class of homeodomain proteins encoded by the STM gene in Arabidopsis. Nature 379, 66-69.
    • (1996) Nature , vol.379 , pp. 66-69
    • Long, J.A.1    Moan, E.I.2    Medford, J.I.3    Barton, M.K.4
  • 33
    • 0033926906 scopus 로고    scopus 로고
    • Positional cloning in Arabidopsis. Why it feels good to have a genome initiative working for you
    • Lukowitz, W., Gillmore, C.S., and Scheible, W.R. (2000). Positional cloning in Arabidopsis. Why it feels good to have a genome initiative working for you. Plant Physiol. 123, 796-805.
    • (2000) Plant Physiol. , vol.123 , pp. 796-805
    • Lukowitz, W.1    Gillmore, C.S.2    Scheible, W.R.3
  • 34
    • 0032444227 scopus 로고    scopus 로고
    • Role of WUSCHEL in regulating stem cell fate in the Arabidopsis shoot meristem
    • Mayer, K.F.X., Schoof, H., Haecker, A., Lenhard, M., Jürgens, G., and Laux, T. (1998). Role of WUSCHEL in regulating stem cell fate in the Arabidopsis shoot meristem. Cell 95, 805-815.
    • (1998) Cell , vol.95 , pp. 805-815
    • Mayer, K.F.X.1    Schoof, H.2    Haecker, A.3    Lenhard, M.4    Jürgens, G.5    Laux, T.6
  • 35
    • 0031661098 scopus 로고    scopus 로고
    • Leaf polarity and meristem formation in Arabidopsis
    • McConnell, J.R., and Barton, M.K. (1998). Leaf polarity and meristem formation in Arabidopsis. Development 125, 2935-2942.
    • (1998) Development , vol.125 , pp. 2935-2942
    • McConnell, J.R.1    Barton, M.K.2
  • 36
    • 0035822096 scopus 로고    scopus 로고
    • Role of PHABULOSA and PHAVOLUTA in determining radial patterning in shoots
    • McConnell, J.R., Emery, J., Eshed, Y., Bao, N., Bowman, J., and Barton, M.K. (2001). Role of PHABULOSA and PHAVOLUTA in determining radial patterning in shoots. Nature 411, 709-713.
    • (2001) Nature , vol.411 , pp. 709-713
    • McConnell, J.R.1    Emery, J.2    Eshed, Y.3    Bao, N.4    Bowman, J.5    Barton, M.K.6
  • 37
    • 0742305480 scopus 로고    scopus 로고
    • HD-Zip III homeobox genes that include a novel member, ZeHB-13 (Zinnia)/ATHB-15 (Arabidopsis), are involved in procambium and xylem cell differentiation
    • Ohashi-lto, K., and Fukuda, H. (2003). HD-Zip III homeobox genes that include a novel member, ZeHB-13 (Zinnia)/ATHB-15 (Arabidopsis), are involved in procambium and xylem cell differentiation. Plant Cell Physiol. 44, 1350-1358.
    • (2003) Plant Cell Physiol. , vol.44 , pp. 1350-1358
    • Ohashi-lto, K.1    Fukuda, H.2
  • 38
    • 0035143177 scopus 로고    scopus 로고
    • REVOLUTA regulates meristem initiation at lateral positions
    • Otsuga, D., DeGuzman, B., Prigge, M.J., Drews, G.N., and Clark, S.E. (2001). REVOLUTA regulates meristem initiation at lateral positions. Plant J. 25, 223-236.
    • (2001) Plant J. , vol.25 , pp. 223-236
    • Otsuga, D.1    Deguzman, B.2    Prigge, M.J.3    Drews, G.N.4    Clark, S.E.5
  • 40
    • 22144438066 scopus 로고    scopus 로고
    • Class III homeodomain-leucine zipper gene family members have overlapping, antagonistic, and distinct roles in Arabidopsis development
    • Prigge, M.J., Otsuga, D., Alonso, J.M., Ecker, J.R., Drews, G.N., and Clark, S.E. (2005). Class III homeodomain-leucine zipper gene family members have overlapping, antagonistic, and distinct roles in Arabidopsis development. Plant Cell 17, 61-76.
    • (2005) Plant Cell , vol.17 , pp. 61-76
    • Prigge, M.J.1    Otsuga, D.2    Alonso, J.M.3    Ecker, J.R.4    Drews, G.N.5    Clark, S.E.6
  • 41
    • 0034020523 scopus 로고    scopus 로고
    • Auxin regulates the initiation and radial position of plant lateral organs
    • Reinhardt, D., Mandel, T., and Kuhlemeier, C. (2000). Auxin regulates the initiation and radial position of plant lateral organs. Plant Cell 12, 507-518.
    • (2000) Plant Cell , vol.12 , pp. 507-518
    • Reinhardt, D.1    Mandel, T.2    Kuhlemeier, C.3
  • 44
    • 0026605283 scopus 로고
    • HD-Zip proteins: Members of an Arabidopsis homeodomain protein superfamily
    • USA
    • Schena, M., and Davis, R.W. (1992). HD-Zip proteins: Members of an Arabidopsis homeodomain protein superfamily. Proc. Natl. Acad. Sci. USA 89, 3894-3898.
    • (1992) Proc. Natl. Acad. Sci. , vol.89 , pp. 3894-3898
    • Schena, M.1    Davis, R.W.2
  • 45
    • 0034677650 scopus 로고    scopus 로고
    • The stem cell population of Arabidopsis shoot meristems is maintained by a regulatory loop between the CLAVATA and WUSCHEL genes
    • Schoof, H., Lenhard, M., Haecker, A., Mayer, K.F.X., Jürgens, G., and Laux, T. (2000). The stem cell population of Arabidopsis shoot meristems is maintained by a regulatory loop between the CLAVATA and WUSCHEL genes. Cell 100, 635-644.
    • (2000) Cell , vol.100 , pp. 635-644
    • Schoof, H.1    Lenhard, M.2    Haecker, A.3    Mayer, K.F.X.4    Jürgens, G.5    Laux, T.6
  • 46
    • 0002839251 scopus 로고
    • Experiments on phyllotaxis. I. The effect of isolating a primordium
    • Snow, M., and Snow, R. (1931). Experiments on phyllotaxis. I. The effect of isolating a primordium. Philos. Trans. R. Soc. Lond. B Biol. Sci. 221, 1-43.
    • (1931) Philos. Trans. R. Soc. Lond. B Biol. Sci. , vol.221 , pp. 1-43
    • Snow, M.1    Snow, R.2
  • 48
    • 0029088028 scopus 로고
    • The REVOLUTA gene is necessary for apical meristem development and for limiting cell divisions in the leaves and stems of Arabidopsis thaliana
    • Talbert, P.B., Alder, H.T., Parks, D.W., and Comai, L. (1995). The REVOLUTA gene is necessary for apical meristem development and for limiting cell divisions in the leaves and stems of Arabidopsis thaliana. Development 121, 2723-2735.
    • (1995) Development , vol.121 , pp. 2723-2735
    • Talbert, P.B.1    Alder, H.T.2    Parks, D.W.3    Comai, L.4
  • 49
    • 0000215040 scopus 로고
    • Further experimental observations on the shoot apex of Dryopteris aristata Druce
    • Wardlaw, C.W. (1949). Further experimental observations on the shoot apex of Dryopteris aristata Druce. Phil. Trans. R. Soc. Lond. Ser. B 233, 415-451.
    • (1949) Phil. Trans. R. Soc. Lond. Ser. B , vol.233 , pp. 415-451
    • Wardlaw, C.W.1
  • 50
    • 0037314247 scopus 로고    scopus 로고
    • POLTERGEIST encodes a phosphatase 2C that regulates CLAVATA pathways controlling stem cell identity at Arabidopsis shoot and flower meristems
    • Yu, L.P., Miller, A.K., and Clark, S.E. (2003). POLTERGEIST encodes a phosphatase 2C that regulates CLAVATA pathways controlling stem cell identity at Arabidopsis shoot and flower meristems. Curr. Biol. 13, 170-188.
    • (2003) Curr. Biol. , vol.13 , pp. 170-188
    • Yu, L.P.1    Miller, A.K.2    Clark, S.E.3
  • 51
    • 0034107779 scopus 로고    scopus 로고
    • POLTERGEIST functions to regulate meristem development downstream of the CLAVATA loci
    • Yu, L.P., Simon, E.J., Trotochaud, A.E., and Clark, S.E. (2000). POLTERGEIST functions to regulate meristem development downstream of the CLAVATA loci. Development 137, 1661-1670.
    • (2000) Development , vol.137 , pp. 1661-1670
    • Yu, L.P.1    Simon, E.J.2    Trotochaud, A.E.3    Clark, S.E.4
  • 52
    • 0033230547 scopus 로고    scopus 로고
    • IFL1, a gene regulating interfascicular fiber differentiation in Arabidopsis encodes a homeodomain-leucine zipper protein
    • Zhong, R., and Ye, Z.-H. (1999). IFL1, a gene regulating interfascicular fiber differentiation in Arabidopsis encodes a homeodomain-leucine zipper protein. Plant Cell 11, 2139-2152.
    • (1999) Plant Cell , vol.11 , pp. 2139-2152
    • Zhong, R.1    Ye, Z.-H.2
  • 53
    • 0034983328 scopus 로고    scopus 로고
    • Alteration of auxin polar transport in the Arabidopsis ifl1 mutants
    • Zhong, R., and Ye, Z.-H. (2001). Alteration of auxin polar transport in the Arabidopsis ifl1 mutants. Plant Physiol. 126, 549-563.
    • (2001) Plant Physiol. , vol.126 , pp. 549-563
    • Zhong, R.1    Ye, Z.-H.2


* 이 정보는 Elsevier사의 SCOPUS DB에서 KISTI가 분석하여 추출한 것입니다.