On the structure of snake hemipenes with comments on their proper preparation for analysis: A reply to cadle, myers, prudente, and zaher

Herpetological Review

Volumn 35, Issue 4, 2004, Pages 320-328

  Dowling, Herndon G ^a  

^a Rendalia Biologists   (United States)

Author keywords

[No Author keywords available]

Indexed keywords

SERPENTES;

EID: 12244272489     PISSN: 0018084X     EISSN: None     Source Type: Journal    
DOI: None     Document Type: Review

References (40)

1. AUTH, D. L., J. MARIAUX, J. CLARY, D. CHISZAR, F. VAN BREUKLEN, AND H. M. SMITH. 1998. The report of the snake genus Conophis in South America is erroneous. Bull. Maryland Herpetol. Soc. 34(4):107-112.
2. BOGERT, C. M. 1940. Herpetological results of the Vernay Angola expedition, with notes on African reptiles in other collections. I. Snakes, including an arrangement of African Colubridae. Bull. Am. Mus. Nat. Hist. 77:1-107, 1 pl.
3. _. 1947. The status of the genus Leptodrymus Amaral, with comments on the modifications of colubrid premaxillae. Am. Mus. Novitates (1352):1-14.
4. BOURGEOIS, M. "1968" [1969]. Contribution a la Morphologie comparee du crane des ophidians de l'Afrique Centrale. Pub. Offic. Congo a Lubumbashi 18:1-293.
5. BRAIN, C. K. 1959. Mating in the South African Mole Snake, Pseudaspis cana (Linnaeus). Copeia 1959(1):71-72.
6. BROADLEY, D.G. 1983. Fitzsimons' Snakes of Southern Africa. Delta Books, Johannesburg.
7. CADLE, J. E. 1984. Molecular systematics of Neotropical xenodontine snakes. I. South American xenodontines. Herpetologica 40(1):8-20.
8. COLE, C. J., AND H. G. DOWLING. 1970. Chromosomes of the Sunbeam Snake, Xenopeltis unicolor Reinwardt (Reptilia: Xenopeltidae). Herpetol. Rev. 2(2):35-36.
9. _, AND L. M. HARDY. 1981. Systematics of North American colubrid snakes related to Tantilla planiceps (Blainville). Bull. Am. Mus. Nat. Hist. 171.
10. COPE, E. D. 1893. Prodromus of a new system of the non-venomous snakes. Am. Nat. 27:477-483.
11. _. 1895. The classification of the Ophidia. Trans. Am. Phil. Soc. (n.s.) 28:186-219, pl. 14-33.
12. DIXON, J. R. 1980. The Neotropical colubrid snake genus Liophis. The generic concept. Contrib. Biol. Geol., Milwaukee Publ. Mus. (31):1-40.
13. DOMERGUE, C. A. 1962. Observations sur les penis des Ophidiens (deuxieme note). Bull. Soc. Sci. Nat. Phys, Maroc 42(1-2):87-105.
14. DOWLING, H. G. 1958a. A taxonomic study of the ratsnakes. VI. Validation of the genera Gonyosoma Wagler and Elaphe Fitzinger. Copeia 1958(1):29-40.
15. _. 1958b. Pleistocene snakes of the Ozarkian Plateau. Am. Mus. Novitates (1882):1-9.
16. _. 1988. Families and higher taxa of the Serpentes: A new classification. In H. G. Dowling and J. V. Jenner, Snakes of Burma: checklist of reported species and bibliography. Smithsonian Herpetol. Inform. Serv. (76): 17-19.
17. _. 2002. "Intraspecific variation of the hemipenis," a correction, with comments on other erroneous descriptions. Herpetol. Rev. 33(1):12-14.
18. _, C. A. HASS, S. B. HEDGES, AND R. HIGHTON. 1996. Snake relationships revealed by slow-evolving proteins: a preliminary survey. J. Zool. (London) 240:1-28.
19. _, R. HIGHTON, G. C. MAHA, AND L. R. MAXSON. 1983. Biochemical evaluation of colubrid snake phylogeny. J. Zool. (London) 201:309-329.
20. _, AND T. PINOU. 2003. Xenodermatid snakes in America. Herpetol. Rev. 34(1):20-23. [Note that this paper, as published, was incomplete: the table with the list of species identified as xenodermatids was inadvertently omitted by the publisher. See: 〈http://pantheon.yale.edu/~tp36/correctedarticle. pdf〉 for a complete copy.]
21. _, AND R. M. PRICE. 1988. A proposed new genus for Elaphe subocularis and Elaphe rosaliae. The Snake [Japan] 20:52-63.
22. _, AND J. M. SAVAGE. 1960. A guide to the snake hemipenis: a survey of basic structure and systematic characteristics. Zoologica (N.Y.) 45:17-27, 3 pl.
23. DUNN, E. R. 1928. A tentative key and arrangement of the American genera of Colubridae. Bull. Antivenin Inst. Am. 2:18-24.
24. JENNER, J. V. 1981. A Zoogeographic Study and the Taxonomy of the Xenodontine Colubrid snakes. Unpubl. Ph. D. Dissertation, New York University, New York.
25. LEVITON, A. E. 1964. Contributions to a review of Philippine snakes, VI. The snakes of the genus Oxyrhabdium. Philippine J. Sci.: 407-422.
26. _, R. H. GIBBS, JR., E. HEAL, AND C. E. DAWSON. 1985. Standards in herpetology and ichthyology: Part I. Standard symbolic codes for institutional resource collections in herpetology and ichthyology. Copeia 1985(3):802-832.
27. MIRANDA, M. E., G. A. COUTURIER, AND J. D. WILLIAMS. 1982. Guia de los ofidios Bonaerenses. Asoc. Coop. Jardin Zool. de la Plata.
28. MYERS, C. W., AND J. E. CADLE. 1994. A new genus for South American snakes related to Rhadinaea obtusa Cope (Colubridae) and resurrection of Taeniophallus Cope for the "Rhadinaea" brevirostris group. Am. Mus. Novitates (3102):1-33.
29. _, AND _. 2003. On the snake hemipenis, with notes on Psomophis and techniques of eversion: A response to Dowling. Herpetol. Rev. 34(4):295-302.
30. ORTENBERGER, A. I. 1923. A method of preparing reptile penes. Copeia (119):71-73.
31. _. 1928. The whip snakes and racers: Genera Masticophis and Coluber. Mem. Univ. Michigan Mus. 1:1-247.
32. PESANTES, O. S. 1994. A method for preparing the hemipenis of preserved snakes. J. Herpetol. 28(1):93-95.
33. POPE, C. H. 1935. The Reptiles of China. Vol. 10. In C. A. Reeds (ed.), Natural History of Central Asia. Publ. Am. Mus. Nat. Hist., New York.
34. SMITH, M. A. 1943. Vol. III. Serpentes. In The Fauna of British India, Ceylon and Burma. Publ. British Mus. (Nat. Hist.), London.
35. WELLMAN, J. 1963. A revision of the genus Conophis (Family Colubridae, from Middle America). Univ. Kansas Publ. Mus. Nat. Hist. 15(6):251-295.
36. WITTE, G. F., DE. 1962. Genera des Serpents du Congo et du Ruanda-Urundi. Ann. Mus. Roy. Afr. Cent. - Sci. Zool. (104):1-203, xv pl.
37. YUKI, R. N. 1997. An observation technique for snake and lizard hemipenes with comments on preparation. Herpetol. Rev. 28(4):196.
38. ZAHER, H. 1999. Hemipenial morphology of the South American xenodontine snakes, with a proposal for a monophyletic Xenodontinae and a reappraisal of colubroid hemipenes. Bull. Am. Mus. Nat. Hist. (240).
39. _, AND A. L. C. PRUDENTE. 1999. Infraspecific variation of the hemipenis in Siphlophis and Tripanurgos. J. Herpetol. 33(4):698-702.
40. _, AND _. 2003. Hemipenes of Siphlophis (Serpentes, Xenodontinae) and techniques of hemipenial preparation in snakes: A response to Dowling. Herpetol. Rev. 34(4):302-307.