On the Snake Hemipenis, with Notes on Psomophis and Techniques of Eversion: A Response to Dowling

Herpetological Review

Volumn 34, Issue 4, 2003, Pages 295-302

  Myers, Charles W ^a   Cadle, John E ^b  

^a DIVISION OF INVERTEBRATE ZOOLOGY   (United States)

^b BROOKFIELD ZOO   (United States)

Author keywords

[No Author keywords available]

Indexed keywords

SERPENTES;

EID: 0346758168     PISSN: 0018084X     EISSN: None     Source Type: Journal    
DOI: None     Document Type: Review

References (36)

1. ARNOLD, E. N. 1984. Variation in the cloacal and hemipenial muscles of lizards and its bearing on their relationships. Symp. Zool. Soc. London 52:47-85.
2. BÖHME, W. 1988. Zur Genitalmorphologie der Sauria: funktionelle und stammesgeschichtliche Aspekte. Bonner Zool. Monogr. 27:176 pp.
3. _. 1995. Hemiclitoris discovered: a fully differentiated erectile structure in female monitor lizards (Varanus spp.) (Reptilia: Varanidae). J. Zool. Syst. Evol. Research 33:129-132.
4. BRANCH, W. R. 1986. Hemipenial morphology of African snakes: a taxonomic review. Part 1. Scolecophidia and Boidae. J. Herpetol. 20:285-299.
5. _, AND E. O. Z. WADE. "1976" [1977]. Hemipenial morphology of British snakes. British J. Herpetol. 5(7):548-553.
6. BROADLEY, D. G. 1980. A revision of the African snake genus Prosymna Gray (Colubridae). Occas. Papers Natl. Mus. and Monuments, series B, Nat. Sci. 6(7):481-556.
7. CADLE, J. E. 1996a. Snakes of the genus Liopholidophis (Colubridae) from eastern Madagascar: new species, revisionary notes, and an estimate of phylogeny. Bull. Mus. Comp. Zool. 154(5):369-464.
8. _. 1996b. Systematics of snakes of the genus Geodipsas (Colubridae) from Madagascar, with descriptions of new species and observations on natural history. Bull. Mus. Comp. Zool. 155(2):33-87.
9. _, AND C. W. MYERS. 2003. Systematics of snakes referred to Dipsas variegata in Panama and western South America, with revalidation of two species and notes on defensive behaviors in the Dipsadini (Colubridae). Amer. Mus. Novitates (3409):1-47.
10. COPE, E. D. 1895. The classification of the Ophidia. Trans. Amer. Phil. Soc., n.s., 18(pt. 2, art. 3):186-219, pls. 14-33 (reprint issued April 15, 1895, with identical pagination).
11. DOWLING, H. G. 2002. "Intraspecific variation of the hemipenis," a correction, with comments on other erroneous descriptions. Herpetol. Rev. 33:12-14.
12. _, AND W. E. DUELLMAN. "1974-1978" [1978]. Systematic herpetology: a synopsis of families and higher categories. New York, privately printed by senior author (as "Hiss Publications, Publ. Herpetology, no. 7, with minor revisions, 1975-1978"), issued loose-leaf, [302 pp.].
13. _, AND J. M. SAVAGE. 1960. A guide to the snake hemipenis: a survey of basic structure and systematic characteristics. Zoologica (New York) 45(1): 17-28 + 3 pls.
14. GIBSON, F. W[AITE]. 1966. Head muscles of Boa constrictor. Zoologica (New York) 51(1):29-48
15. _. 1970. The "quadrifurcate" hemipenis of Tropidophis. Herpetol. Rev. 2(4):29.
16. _. [MS. circa 1970-1974]. Procedure for everting hemipenes of preserved snakes. Unpublished manuscript; copy filed in AMNH Herpetology Archives (Myers collection, Waite Gibson folder).
17. HARDY, L. M. 1975. A systematic revision of the colubrid snake genus Gyalopion. J. Herpetol. 9:107-132.
18. HOGE, A. R., H. E. BELLUOMINI, G. SCHREIBER, AND A. M. PENHA. 1959. Sexual abnormalities in Bothrops insularis. Mem. Inst. Butantan 29:17-88.
19. MANZANI, P. R., AND A. S. ABE. 1988. Sobre dois novos métodos de prepara do hemipênis de serpentes. Mem. Inst. Butantan 50(1):15-20.
20. MACLEAN, W. P. 1968. A case of intersexuality in Bothrops moojeni Hoge. Copeia 1968(1):170.
21. MYERS, C. W. 1974. The systematics of Rhadinaea (Colubridae), a genus of New World snakes. Bull. Amer. Mus. Nat. Hist. 153(1):1-262.
22. _. 2003. Rare snakes - five new species from eastern Panama: reviews of northern Atractus and southern Geophis (Colubridae: Dipsadinae). Amer. Mus. Novitates (3391):1-47.
23. _, AND J. E. CADLE. 1994. A new genus for South American snakes related to Rhadinaea obtusa Cope (Colubridae) and resurrection of Taeniophallus Cope for the "Rhadinaea" brevirostris group. Amer. Mus. Novitates (3102):1-33.
24. _, AND M. A. DONNELLY. 2001. Herpetofauna of the Yutajé-Corocoro massif, Venezuela: second report from the Robert G. Goelet American Museum-TERRAMAR Expedition to the northwestern tepuis. Bull. Amer. Mus. Nat. Hist. (261):1-85.
25. ORTENBURGER, A. I. 1923. A method of preparing reptile penes. Copeia 119:71-73.
26. PESANTES, O. S. 1994. A method for preparing the hemipenis of preserved snakes. J. Herpetol. 28:93-95.
27. SAVAGE, J. M. 1997. On terminology for the description of the hemipenis of squamate reptiles. Herpetol. J. 7:23-25.
28. SCHULZ, K.-D., 1996. A monograph of the colubrid snakes of the genus Elaphe Fitzinger. Havlickuv Brod, Czech Republic: Koeltz Scientific Books, iii, 439 pp.+ 58 color pls.
29. VAN CLEAVE, H. J., AND J. A. Ross. 1947a. A method for reclaiming dried zoological specimens. Science 105:318.
30. _, AND J. A. Ross. 1947b. Use of trisodium phosphate in microscopical technic. Science 106:194.
31. YUKI, R. N. 1997. An observation technique for snake and lizard hemipenes with comments on preparation. Herpetol. Rev. 28:196.
32. ZAHER, H. 1999. Hemipenial morphology of the South American xenodontine snakes, with a proposal for a monophyletic Xenodontinae and a reappraisal of colubroid hemipenes. Bull. Amer. Mus. Nat. Hist. (240):1-168.
33. _, AND A. L. C. PRUDENTE. 1999. Intraspecific variation of the hemipenis in Siphlophis and Tripanurgos. J. Herpetol. 33:698-702.
34. _, AND _. 2003. Hemipenes of Siphlophis (Serpentes, Xenodontinae) and techniques of hemipenial preparation in snakes: a response to Dowling. Herpetol. Rev. 34:302-307.
35. ZIEGLER, T., AND W. B̈OHME. 1996. Zur Hemiclitoris der squamaten Reptilien: Auswirkungen auf einige Methoden der Geschlechtsunterscheidung. Herpetofauna 18(101):11-19.
36. _, AND _. 1997. Genitalstrukturen und Paarungsbiologie bei squamaten Reptilien, speziell den Platynota, mit Bemerkungen zur Systematik. Mertensiella 8:1-210.