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The present study is part of the PhD of Pierre Franck which he performed under the joint scientific authority of Jean-Marie Comuet (director of research at the French National Institute of Agronomical Research) and Michel Solignac (Professor at Paris XI University). Lionel Garnery, a former PhD student of J.M.C. and M.S. is Maître de Conférence at the University of Saint Quentin en Yveline (near Paris) and Giovanna Celebrano is researcher in Ecology at the University of Torino.
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