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We thank B. Aaby, R. Below, and P. J. C. Kuiper for stimulating discussions and for constructive comments on the topic. Supported by the Deutsche Forschungsgemeinschaft (DFG) and the Council for Earth and Life Sciences (ALW) of the Netherlands Organization for Scientific Research (NWO). This is contribution 990501 from the Netherlands Research School of Sedimentary Geology and contribution 510 from the University of Florida Contributions to Paleobiology.
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