Feral asses (Equus africanus): Intraspecific variation in social organization in arid and mesic habitats

Applied Animal Behaviour Science

Volumn 60, Issue 2-3, 1998, Pages 171-195

  Moehlman, Patricia D ^a  

^a Box 2031 ^*  (Tanzania)

Author keywords

Equids; Feral asses; Intraspecific variation; Mating systems; Social organization

Indexed keywords

ENVIRONMENTAL CONDITIONS; FERAL ORGANISM; INTRASPECIFIC VARIATION; REPRODUCTIVE STRATEGY; SOCIAL STRUCTURE; UNGULATE;

UNITED STATES;

EQUUS AFRICANUS; EQUUS ASINUS;

EID: 0032533639     PISSN: 01681591     EISSN: None     Source Type: Journal    
DOI: 10.1016/S0168-1591(98)00163-4     Document Type: Article

References (67)

1. Alexander R.D. The evolution of social behavior. Annu. Rev. Ecol. Syst. 5:1974;325-383.
2. Altmann J. Observational study of behaviour: sampling methods. Behaviour. 49:1974;227-267.
3. Becker C.D., Ginsberg J.R. Mother-infant behaviour of wild Grevy's zebra: adaptations for survival in semidesert East Africa. Anim. Behav. 40:1990;111-1118.
4. Berger, J., 1986. Wild Horses of the Great Basin: Social Competition and Population Size. University of Chicago Press, 326 pp.
5. Carothers, S.W., Stitt, M.E., Johnson, R.R., 1976. Feral asses on public lands: an analysis of biotic impact, legal consideration, and management alternatives. Trans. 41st North Amer. Wildl. Nat. Res. Conf., pp. 396-406.
6. Crowell-Davis S.L. Nursing behaviour and maternal aggression among Welsh ponies (Equus caballus). Appl. Anim. Behav. Sci. 14:1985;11-25.
7. Duncan P. Foal killing by stallions. Appl. Anim. Ethol. 8:1982;567-570.
8. Duncan P. Determinants of the use of habitat by horses in a Mediterranean wetland. J. Anim. Ecol. 52:1983;93-109.
9. Duncan, P., 1992. Zebras, Asses, and Horses: an Action Plan for the Conservation of Wild Equids, IUCN Specialist Group, IUCN, Gland, Switzerland, 36 pp.
10. Emlen S.T., Oring S.W. Ecology, sexual selection, and the evolution of mating systems. Science. 197(4300):1977;215-223.
11. Evans R.M. The relationship between parental input and parental investment. Anim. Behav. 39:1990;797-798.
12. Feh C., Boldsukh T., Tourenq C. Are family groups in Equids a response to cooperative hunting by predators? The case of Mongolian Kulans (Equus hemionus luteus Matschie). Rev. Ecol. (Terre Vie). 49:1994;11-20.
13. Feist, J.D., 1971. Behavior of feral horses in the Pryor Mountain wild horse range. MS thesis, University of Michigan, Ann Arbor, 129 pp.
14. Gentry A., Clutton-Brock J., Groves C.P. Proposed conservation of usage of 15 mammal specific names based on wild species which are antedated by or contemporary with those based on domestic animals. Bull. Zoological Nomenclature. 53(1):1996;28-37.
15. George M., Ryder O.A. Mitochondrial DNA evolution in the genus Equus. Mol. Biol. Evol. 3(6):1986;535-546.
16. Ginsberg, J.R., 1988. Social organization and mating strategies of an arid adapted equid: the Grevy's zebra. PhD dissertation, Princeton University, Princeton, NJ, 268 pp.
17. Ginsberg, J.R., 1989. The ecology of female behavior and male mating success in the Grevy's zebra, Equus grevyi. Symp. Zool. (Lond.) Soc., 6, pp. 89-110.
18. Ginsberg J.R., Rubenstein D.I. Sperm competition and variation in zebra mating behavior. Behav. Ecol. Sociobiol. 26:1990;427-434.
19. Green N.F., Green H.D. The wild horse population of Stone Cabin Valley, Nevada: a preliminary report. Proc. Nat. Wild Horse Forum. 1:1977;59-65.
20. Groves, C.P., 1974. Horses, Asses, and Zebras in the Wild. David & Charles, Newton Abbot, London.
21. Groves, C.P., 1986. The taxonomy, distribution and adaptations of recent equids. In: Meadow, R.H., Uerpman, H.P. (Eds.), Equids in the Ancient World, Dr. Ludwig Reichert Verlag, Wiesbaden, pp. 11-65.
22. Hanley, T.A., 1976. Carrying capacity relationships of feral burro (Equus asinus) habitat in a Sonoran desert ecosystem. MS thesis, Arizona State University, Tempe, 156 pp.
23. Hanley T.A., Brady W.W. Feral burro impact on a Sonoran desert range. J. Range Manage. 30(5):1977;374-377.
24. Hughes J.P., Stabenfeldt G.H., Evans J.W. Estrus cycle and ovulation in the mare. J. Am. Vet. Med. Assoc. 161(11):1972;1367-1374.
25. Imanishi K. Social life of semi-wild horse in Toimisaki: III. Summary for three surveys undertaken in 1948-1949. Physiol. Ecol. (Japan). 4:1950;29-42.
26. Jarman P.J. The social organization of antelope in relation to their ecology. Behaviour. 48:1974;215-267.
27. Johnson, A.S., Hillestad, H.O., Shanholtzer, S.F., Shanholtzer, G.F., 1974. An ecological survey of the coastal region of Georgia. Nat. Park Serv. Sci. Monogr. Ser., No. 3.
28. Joubert E. The social organization and associated behavior in Hartmann zebra, Equus zebra hartmannae. Madoqua. I(6):1972;17-56.
29. Keiper R.R. Social organization of feral ponies. Proc. Penn. Acad. Sci. 50:1976;69-70.
30. Kenneth, J.H., 1953. Gestation periods. Commonwealth Agricultural Bureau. Farntham Royal, Slough, Bucks, England, 39 pp.
31. Klingel H. Soziale organisation und verhalten freilebender Steppenzebras (Equus quagga). Z. Tierpsychol. 24:1967;580-624.
32. Klingel H. Soziale organisation und verhaltenweisen von Hartmann- und Bergzebras (Equus zebra hartmannae and E. zebra zebra). Z. Tierpsychol. 25:1968;76-88.
33. Klingel H. Reproduction in the plains zebra Equus burchelli boehmi: behaviour and ecological factors. J. Reprod. Fert. 6:1969;339-345. Suppl.
34. Klingel H. Zur Soziologie der Grevy-Zebras. Zool. Anz. 33:1969;311-316. Suppl.
35. Klingel H. Social behaviour of African Equidae. Zool. Afr. 7:1972;175-186.
36. Klingel H. Social organization and behaviour of the Grevy's zebra. Z. Tierpsych. 35:1974;37-70.
37. Klingel H. Social organization and reproduction in equids. J. Reprod. Fert. 23:1975;7-11. Suppl.
38. Klingel H. Observations on social organization and behaviour of African and Asiatic asses (Equus africanus and E. hemionus). Z. Tierpsychol. 44:1977;323-331.
39. Lloyd P.H., Rasa O.A.E. Status, reproductive success and fitness in Cape mountain zebra (Equus zebra zebra). Behav. Ecol. Sociobiol. 25:1989;411-420.
40. Maloiy G.M.O. Water economy of the Somali donkey. Am. J. Physiol. 219(5):1970;1522-1527.
41. Maloiy G.M.O., Boarer D.H. Response of the Somali donkey to dehydration hematological changes. Am. J. Physiol. 221:1971;37-41.
42. McCort, W.D., 1980. The feral asses (Equus asinus) of Ossabaw Island, Georgia. PhD dissertation, Pennsylvania State University, University Park, 219 pp.
43. Monard A., Duncan P., Boy V. The proximate mechanisms of natal dispersal in female horses. Behaviour. 133:1996;1095-1124.
44. Morgart, J.R., 1978. Burro behavior and population dynamics in Bandelier National Monument, New Mexico. MS thesis, Arizona State University, Tempe, 94 pp.
45. Miller R., Denniston R.H. Interband dominance in feral horses. Z. Tierpsychol. 51:1979;41-47.
46. Moehlman P.D. Social organization and reproductive behavior of feral asses (Equus asinus). Am. Zool. 13:1973;1261.
47. Moehlman, P.D., 1974. Behavior and ecology of feral asses (Equus asinus). PhD dissertation, University of Wisconsin, Madison, 251 pp.
48. Moehlman, P.D., 1979. Behavior and ecology of feral asses (Equus asinus). Nat. Geogr. Soc. Res. Reports, 1970, pp. 405-411.
49. National Research Council, 1978. Nutritional requirements of horses. National Academy Press, Washington, DC.
50. Nelson, K.J., 1979. On the question of male-limited population growth in feral horses (Equus caballus). MS thesis, New Mexico State University, Las Cruces, 29 pp.
51. Norment, C., Douglas, C.C., 1977. Ecological studies of feral burros in Death Valley. University of Nevada Cooperative National Park Research Studies Unit, Contribution 17.
52. O'Farrell, M.J., 1978. An assessment of impact of feral burros on natural ecosystems of the Lake Mead National Recreation Area, Arizona-Nevada. LAME Tech. Report No. 4, 37 pp.
53. Penzhorn B.L. A long-term study of social organization and behaviour of Cape mountain zebra Equus zebra zebra. Z. Tierpsychol. 64:1984;97-146.
54. Rowen, M., 1992. Mother-Infant Behavior and Ecology of Grevy's Zebra, Equus grevyi. PhD dissertation, Yale University, New Haven, CT, 159 pp.
55. Rowen, M., Ginsberg, J.R., 1992. Grevy's zebra. In: Duncan, P. (Ed.), Zebras, Asses, and Horses: an Action Plan for the Conservation of Wild Equids, IUCN Specialist Group, IUCN, Gland, Switzerland, 36 pp.
56. Rubenstein D.I. Behavioral ecology of island feral horses. Equine Vet. J. 13:1981;27-34.
57. Rubenstein, D.I., 1986. Ecology and sociality in horses and zebras. In: Rubenstein, D.I., Wrangham, R.W. (Eds.), Ecological Aspects of Social Evolution. Princeton University Press, Princeton, NJ, pp. 282-302.
58. Rubenstein, D.I. 1994. The ecology of female social behaviour. In: Jarman, P.J., Rossiter, A. (Eds.), Animal Societies, Individuals, Interactions, and Organization. Horses, Zebras, and Asses. Kyoto University Press, Kyoto, Japan, pp. 13-28.
59. Rudman, R., 1990. The behavior and ecology of feral burros on St. John, U.S. Virgin Islands, PhD dissertation, Cornell University, Ithaca, NY, 184 pp.
60. Seegmiller R.F., Ohmart R.D. Ecological relationships of feral burros and desert bighorn sheep. Wild. Monogr. 45(78):1981;1-58.
61. Shah, N.V., 1993. Ecology of Wild Ass (Equus hemionus khur) in Little Rann of Kutch. PhD dissertation, Maharaja Sayajirao University of Baroda, India, 182 pp.
62. Tomkiewicz, S.M., Jr., 1979. Heterothermy and Water Turnover in Feral Burros (Equus asinus) of the Desert Southwest. MS thesis, Arizona State University, Tempe, 89 pp.
63. Trivers R.L. Parent-offspring conflict. Am. Zool. 14:1974;249-264.
64. Tyler S. The behavior and social organization of the New Forest ponies. Anim. Behav. Monogr. 5(2):1972;1-196.
65. United States Geological Survey, 1972. Surface water records of California. Denver, CO.
66. Walker, M.T., 1978. Ecological similarities between feral burros and desert bighorn sheep, Black Mountains, northwestern Arizona. MS thesis, Arizona State University, Tempe, 127 pp.
67. Woodward S.L. The social system of feral asses (Equus asinus). Z. Tierpsychol. 49:1979;304-316.