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note
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We thank Carol Bilbrough, Jill Baron, Diane McKnight, Russ Monson, Tim Seastedt, and two anonymous reviewers for comments on this manuscript, and Steve Schmidt for advice with field measurements. Financial support for the work was provided by the NSF sponsored Niwot Ridge Long-Term Ecological Research Program, the U.S EPA, the Andrew W. Mellon Foundation, and an NSF predoctoral fellowship to H. Steltzer
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note
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First submitted 28 April 1997. Accepted for publication after revision 29 August 1997.
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