Crinoid biofacies in Upper Carboniferous cyclothems, midcontinent North America: Faunal tracking and the role of regional processes in biofacies recurrence

Palaeogeography, Palaeoclimatology, Palaeoecology

Volumn 127, Issue 1-4, 1996, Pages 47-81

  Holterhoff, Peter F ^a  

^a UNIVERSITY OF ARIZONA   (United States)

Author keywords

crinoid; North America; palaeocommunity; palaeoecology; Pennsylvanian

Indexed keywords

BIOFACIES; CARBONIFEROUS; CRINOID; CYCLOTHEM; FAUNAL TRACKING;

EID: 0030595824     PISSN: 00310182     EISSN: None     Source Type: Journal    
DOI: 10.1016/S0031-0182(96)00088-0     Document Type: Article

References (127)

1. Albrecht, G.H., 1980. Multivariate analysis and the study of form, with special reference to canonical variate analysis. Am. Zool., 20: 679-693.
2. Arvidson, R.S., 1990. Stratigraphy, carbonate petrology, diagenesis, and trace element cement Geochemistry of the Wyandotte Limestone (Upper Pennsylvanian), Miami County, Kansas. Thesis. Univ. Iowa, 205 pp.
3. Bambach, R.K., 1994. The null hypothesis for community stability and recurrence-environmental selection of adapted organisms. Geol. Soc. Am. Abstr. Programs, 26(7): A-519.
4. Barkai, A. and McQuaid, C., 1988. Predator-prey role reversal in a marine benthic ecosystem. Science, 242: 62-64.
5. Baumiller, T.K., 1990. Crinoid functional morphology and the energetics of passive suspension feeding: implications to the evolutionary history of Paleozoic Crinoidea. Theis. Univ. Chicago, 150 pp. (unpublished).
6. Baumiller, T.K., 1993. Survivorship analysis of Paleozoic Crinoidea: effect of filter morphology on evolutionary rates. Paleobiology, 19(3): 304-321.
7. Bennington, J.B. and Bambach, R.K., 1996. Statistical testing for paleocommunity recurrence: are similar fossil assemblages ever the same? In: L.C. Ivany and K.M. Schopf (Editors), New Perspectives on Faunal Stability in the Fossil Record. Palaeogeogr. Palaeoclimatol. Palaeoecol., xx:xx-xx.
8. Bennison, A.P., 1984. Shelf to trough correlations of Late Desmoinesian and Early Missourian carbonate banks and related strata, Northeast Oklahoma. In: N.J. Hyne (Editor), Limestones of the Mid-Continent. Tulsa Geol. Soc. Spec. Publ., 2: 93-126.
9. Bennison, A.P., 1985. Trough-to-shelf sequence of the early Missourian Skiatook Group, Oklahoma and Kansas. In: W.L. Watney et al. (Editors), Recent Interpretations of Late Paleozoic Cyclothems. SEPM Mid-Cont. Sect. Proc. 3rd Annu. Meet. Field Conf., pp. 219-245.
10. Bjerstedt, T.W., 1988. Multivariate analyses of trace fossil distribution from an Early Mississippian oxygen-deficient basin, central Appalachians. Palaios, 3: 53-68.
11. Boardman, D.R. and Heckel, P.H., 1989. Glacial-eustatic sealevel curve for early Late Pennsylvanian sequence in north-central Texas and biostratigraphic correlation with curve for midcontinent North America. Geology, 17: 802-805.
12. Boardman, D.R., Mapes, R.H., Yancey, T.E. and Malinky, J.M., 1984. A new model for the depth-related allogenic community succession within North American Pennsylvanian cyclothems and implications on the black shale problem. In: N.J. Hyne (Editor), Limestones of the Midcontinent. Tulsa Geol. Soc. Spec. Publ., 2: 141-182.
13. Boucot, A.J., 1983. Does evolution take place in an ecological vacuum? II. J. Paleontol., 57 (1): 1-30.
14. Boucot, A.J., 1986. Ecostratigraphic criteria for evaluating the magnitude, character and duration of bioevents. In: O.H. Walliser (Editor), Global Bio-Events. Lecture Notes Earth Sci., 8: 25-45.
15. Boucot, A.J., 1990. Community evolution: its evolutionary and biostratigraphic significance. In: W. Miller (Editor), Paleocommunity Temporal Dynamics: The Long-Term Development of Multispecies Assemblages. Paleontol. Soc. Spec. Publ., 5: 48-70.
16. Brett, C.E. and Baird, G.C., 1985. Carbonate-shale cycles in the Middle Devonian of New York: an evaluation of models for the origin of limestones in terrigenous shelf sequences. Geology, 13: 324-327.
17. Brett, C.E. and Baird, G.C., 1986. Symmetrical and upward shallowing cycles in the Middle Devonian of New York State and their implications for the punctuated aggradational cycle hypothesis. Paleoceanography, 1: 431-445.
18. Brett, C.E. and Baird, G.C., 1995. Coordinated stasis and evolutionary ecology of Silurian to middle Devonian faunas in the Appalachian Basin. In: D.H. Erwin and R.L. Anstey (Editors), New Approaches to Speciation in the Fossil Record. Columbia Univ. Press, New York, pp. 285-315.
19. Brett, C.E., Miller, K.B. and Baird, G.C., 1990. A temporal hierarchy of paleoecological processes within a Middle Devonian epeiric sea. In: W. Miller (Editor), Paleocommunity Temporal Dynamics: The Long-Term Development of Multispecies Assemblages. Paleontol. Soc. Spec. Publ., 5: 178-209.
20. Brown, J.H., 1995. Macroecology. Univ. Chicago Press, Chicago, 269 pp.
21. Burchett, R.R., 1971. Guidebook to the Geology Along Portions of the Lower Platte River Valley and Weeping Water Valley of Eastern Nebraska. Conserv. Surv. Div. Nebraska Geol. Surv., Univ. Nebraska, 39 pp.
22. Burchett, R.R. and Reed, E.C., 1967. Centennial Guidebook to the Geology of Southeastern Nebraska. Conservation and Survey Division, Nebraska Geological Survey, University of Nebraska, Lincoln, 83 pp.
23. Buzas, M.A. and Culver, S.J., 1994. Species pool and dynamics of marine paleocommunities. Science, 264: 1439-1441.
24. Cocke, J.M. and Strimple, H.L., 1974. Distribution of Algae and Corals in Upper Pennsylvanian Missourian Rocks in Northeastern Oklahoma. In: Field Trip Guidebook, South-Central Sect. Geol. Soc. Am., 42 pp.
25. Cohen, A.S., 1994. Extinction in ancient lakes: biodiversity crises and conservation 40 years after J.L. Brooks. In: K. Martens et al. (Editors), Speciation in Ancient Lakes. Arch. Hydrobiol. Beih. Ergebn. Limnol., 44: 451-479.
26. Coope, G.R., 1979. Late Cenozoic fossil Coleoptera: Evolution, Biogeography, and Ecology. Annu. Rev. Ecol. Syst., 10: 247-267.
27. Crame, J.A., 1986. Late Pleistocene molluscan assemblages from the coral reefs of the Kenya coast. Coral Reefs, 4: 183-196.
28. DiMichele, W.A., 1994. Ecological patterns in time and space. Paleobiology, 20(2): 89-92.
29. Drake, J.A., 1990. Communities as assembled structures: do rules govern pattern? TREE, 5: 159-164.
30. Elias, M.K., 1937. Depth of deposition of the Big Blue (Late Paleozoic) sediments in Kansas. Geol. Soc. Am. Bull., 48: 403-432.
31. Fasham, M.J.R., 1977. A comparison of nonmetric multidimensional scaling, principal components and reciprocal averaging for the ordination of simulated coenoclines and coenoplanes. Ecology, 58: 551-561.
32. Fursich, F.T., Oschmann, W., Jaitly, A.K. and Singh, I.B., 1991. Faunal response to transgressive-regressive cycles: example from the Jurassic of Western India. Palaeogeogr. Palaeoclimatol. Palaeoecol., 85: 149-159.
33. Galloway, W.E., Yancey, M.S. and Whipple, A.P., 1977. Seismic stratigraphic model of depositional platform margin, eastern Anadarko Basin, Oklahoma. AAPG Bull., 61: 1437-1447.
34. Gay, S.P., 1989. Gravitational compaction, a neglected mechanism in structural and stratigraphic studies: new evidence from mid-continent, USA. AAPG Bull., 73: 641-657.
35. Gnanadesikan, R., 1977. Methods for Statistical Data Analysis of Multivariate Observations. Wiley, New York, 311 pp.
36. Gnanadesikan, R. and 11 others, 1989. Discriminant analysis and clustering. Statist. Sci., 4: 34-69.
37. Greenstein, B.I., 1991. An integrated study of echinoid taphonomy: predictions for the fossil record of four echinoid families. Palaios, 6: 519-540.
38. Hallock, P. and Schlanger, W., 1986. Nutrient excess and the demise of coral reefs and carbonate platforms. Palaios, 1: 389-398.
39. Handford, C.R. and Dutton, S.P., 1980. Pennsylvanian-Early Permian depositional systems and shelf-margin evolution, Palo Duro Basin, Texas. AAPG Bull., 64: 88-106.
40. Hanski, I., 1989. Metapopulation dynamics: does it help to have more of the same? TREE, 4: 113-114.
41. Hanski, I., 1991. Single-species metapopulation dynamics: concepts, models and observations. Biol. J. Linnean Soc., 42: 17-38.
42. Hanski, I. and Gilpin, M., 1991. Metapopulation dynamics: brief history and conceptual domain. Biol. J. Linnean Soc., 42: 3-16.
43. Harbaugh, J.W., Merriam, D.F., Wray, J.L. and Jacques, T.E., 1965. Pennsylvanian Marine Banks in Southeastern Kansas. Geol. Soc. Am. Field Conf. Guide Book Annu. Meet., 66 pp.
44. Heckel, P.H., 1975a. Stratigraphy and Depositional Framework of the Stanton Formation in Southeastern Kansas. Kans. Geol. Surv. Bull., 210, 45 pp.
45. Heckel, P.H., 1975b. Upper Pennsylvanian Limestone Facies in Southeastern Kansas. In: Kans. Geol. Soc. 31st Annu. Field Conf. Guidebook, 66 pp.
46. Heckel, P.H., 1977. Origin of phosphatic black shale facies in Pennsylvanian cyclothems of midcontinent North America. AAPG Bull., 61: 1045-1068.
47. Heckel, P.H., 1978. Upper Pennsylvanian cyclothemic limestone facies in Eastern Kansas. Kans. Geol. Surv. Guidebook Sen., 2, pp. 79.
48. Heckel, P.H., 1980. Paleogeography of eustatic model for deposition of midcontinent Upper Pennsylvanian cyclothems. In: T.D. Fouch and E.R. Magathan (Editors), Paleozoic Paleogeography of West Central United States. Rocky Mount. Sect. SEPM Spec. Publ., Denver, pp. 197-215.
49. Heckel, P.H., 1986. Sea-level curve for Pennsylvanian eustatic marine transgressive-regressive depositional cycles along midcontinent outcrop belt, North America. Geology, 14: 330-334.
50. Heckel, P.H. and Pope, J.P., 1992. Stratigraphy and cyclic sedimentation of Middle and Upper Pennsylvanian Strata around Winterset, Iowa. Iowa Geol. Surv. Guidebook Ser., 14, 53 pp.
51. Heckel, P.H., Brady, L.L., Ebanks, W.J. and Pabian, R.K., 1979. Pennsylvanian Cyclic Platform Deposits of Kansas and Nebraska (Guidebook Ser., 4). Kans. Geol. Surv., 79 pp.
52. Hoffman, A., 1981. The ecostratigraphic paradigm. Lethaia, 14: 1-7.
53. Holterhoff, P.F., 1993. The application of filtration theory to the paleoecology of crinoids from the Lansing Group (Missourian, Upper Pennsylvanian), Midcontinent, North America, Thesis. Univ. Cincinnati, 307 pp.
54. Holterhoff, P.F., 1995. Diversity structure of Permo-Pennsylvanian crinoid faunas from midcontinent North America: relationships between alpha and gamma. Geol. Soc. Am. Abstr. Programs, 27(3), p. 60.
55. Holterhoff, P.F., submitted. Filtration models, guilds, and biofacies: crinoid paleoecology of the Stanton Formation (Upper Pennsylvanian), midcontinent, North America, Palaeogeogr. Palaeoclimatol. Palaeoecol.
56. Huntley, B. and Webb, T., 1989. Migration: species' response to climatic variations caused by changes in the earth's orbit. J. Biogeogr., 16: 5-19.
57. Huston, M.A., 1994. Biological Diversity. Cambridge Univ. Press, Cambridge, 681 pp.
58. Imbrie, W.S., 1985. A theoretical framework for the Pleistocene ice ages. J. Geol. Soc. London, 142: 417-432.
59. Ireland, H.A., 1955. Pre-cambrian surface in northeastern Oklahoma and parts of adjacent states. AAPG Bull., 39: 468-483.
60. Joeckel, R.M., 1989. Geomorphology of a Pennsylvanian land surface: pedogenesis in the Rock Lake Shale Member, southeastern Nebraska. J. Sediment. Petrol., 59: 469-481.
61. Kammer, T.W., 1985. Aerosol filtration theory applied to Mississippian deltaic crinoids. J. Paleontol., 59: 551-560.
62. Kammer, T.W. and Ausich, W.I., 1987. Aerosol suspension feeding and current velocities: distributional controls for Late Osagean crinoids. Paleobiology, 13: 379-395.
63. Kidweil, S.M., 1991a. The stratigraphy of shell concentrations. In: P.A. Allison and D.E.G. Briggs (Editors), Taphonomy: Releasing the Data Locked in the Fossil Record. Plenum, New York, pp. 212-290.
64. Kidwell, S.M., 1991b. Condensed deposits in siliciclastic sequences: expected and observed features. In: G. Einsele et al. (Editors), Cycles and Events in Stratigraphy. Springer, Berlin, pp. 682-695.
65. Kidwell, S.M. and Flessa, K., 1995. The Quality of the Fossil Record: Populations, Species, and Communities. Annu. Rev. Ecol. Syst., 26: 269-299.
66. Kovach, W.L., 1988. Multivariate methods of analyzing paleoecological data. In: W.A. DiMichele and S.L. Wing (Editors), Methods and Applications of Plant Paleoecology, Paleontol. Soc. Spec. Publ., 3: 72-104.
67. Kumar, N. and Slatt, R.M., 1984. Submarine-fan and slope facies of Tonkawa (Missourian-Virgilian) Sandstone in deep Anadarko Basin. AAPG Bull., 68: 1839-1856.
68. Lane, N.G. and Webster, G.D., 1980. Crinoidea. In: T.W. Broadhead and J.A. Waters (Editors), Echinoderms. Univ. Tenn. Stud. Geol., 3: 144-157.
69. Lewis, R., 1980. Taphonomy. In: T.W. Broadhead and J.A. Waters (Editors), Echinoderms. Univ. Tenn. Stud. Geol., 3: 27-39.
70. Macurda, D.B. and Meyer, D.L., 1983. Sea lillies and feather stars. Am. Sci., 71: 354-365.
71. Meyer, D.L., 1973. Feeding behavior and ecology of shallowwater unstalked crinoids (Echinodermata) in the Caribbean Sea. Mar. Biol., 22: 105-129.
72. Meyer, D.L., 1979. Length and spacing of the tube feet in crinoids (Echinodermata) and their role in suspension-feeding. Mar. Biol., 51: 361-369.
73. Meyer, D.L., 1982. Food and feeding mechanisms: Crinozoa. In: M. Jangoux and J.M. Lawrence (Editors), Echinoderm Nutrition. Balkema, Rotterdam, pp. 25-42.
74. Meyer, D.L., 1985. Evolutionary implications of predation on Recent comatulid crinoids from the Great Barrier Reef. Paleobiology, 11: 154-164.
75. Meyer, D.L. and Macurda, D.B., 1977. Adaptive radiation of the comatulid crinoids. Paleobiology, 3: 74-82.
76. Meyer, D.L. and Ausich, W.I., 1983. Biotic interactions among Recent and among fossil crinoids. In: M.J.S. Tevesz and P.L. McCall (Editors), Biotic Interactions in Recent and Fossil Benthic Communities. Plenum, New York, pp. 377-427.
77. Meyer, D.L. and Meyer, K., 1986. Biostratinomy of Recent crinoids (Echinodermata) at Lizard Island, Great Barrier Reef, Australia. Palaios, 1: 294-302.
78. Meyer, D.L., Ausich, W.I. and Terry, R.E., 1989. Comparative taphonomy of echinoderms in carbonate facies: Fort Payne Formation (Lower Mississippian) of Kentucky and Tennessee. Palaios, 4: 533-552.
79. Miller, A.I., 1988. Spatial resolution in subfossil molluscan remains: implications for paleobiological analysis. Paleobiology, 14: 91-103.
80. Miller, K.B., Brett, C.E. and Parsons, K.M., 1988. The paleoecologic significance of storm-generated disturbance within a Middle Devonian muddy epeiric sea. Palaios, 3: 35-52.
81. Miller, W., 1990. Hierarchy, individuality, and paleoecosystems. In: W. Miller (Editor), Paleocommunity Temporal Dynamics: The Long-Term Development of Multispecies Assemblages. Paleontol. Soc. Spec. Publ., 5: 31-47.
82. Miller, W., 1993a. Models of recurrent fossil assemblages. Lethaia, 26: 182-183.
83. Miller, W., 1993b. Benthic community replacement and population response. Neues Jahrb. Geol. Paläontol. Abh., 188:133-146.
84. Miller, W., 1994. Ecology of coordinated stasis. Geol. Soc. Am. Abstr. with Programs, 26(7): A-520.
85. Moore, R.C. and 8 others, 1978. Systematic descriptions. In: R.C. Moore and C. Teichert (Editors). Treatise on Invertebrate Paleontology, Part T, Echinodermata 2, Vol. 2. Geol. Soc. Am., Boulder, Univ. Kansas Press, Lawrence, pp. 405-812.
86. Morris, P.J., Ivany, L.C. and Schopf, K.M., 1992. Paleoecological stasis in evolutionary theory. Geol. Soc. Am. Abstr. with Programs, 24(7): A313.
87. Morris, P.J., Ivany, L.C., Schopf, K.M. and Brett, C.E., 1995. The challenge of paleoecological stasis: reassessing sources of evolutionary stability. Proc. Natl. Acad. Sci. USA, 92: 11,269-11,273.
88. Neumann, A.C. and McIntyre, I., 1985. Reef response to sea-level rise: keep-up, catch-up, give-up. Proc. 5th Int. Coral Reef Congr., 3, pp. 105-110.
89. O'Neill, R.V., DeAngelis, D.L., Waide, J.B. and Allen, T.F.H., 1986. A Hierarchical Concept of Ecosystems (Monogr. Popul. Biol., 23). Princeton Univ. Press, 253 pp.
90. Pabian, R.K. and Strimple, H.L., 1993. Taxonomy, Paleoecology and Biostratigraphy of the Crinoids of the South Bend Limestone (Late Pennsylvanian-Missourian, ?Virgilian) in Southeastern Nebraska and Southeastern Kansas. Nebraska Geol. Surv. Prof. Publ., 1, 55 pp.
91. Pauly, G., 1990. Effects of late Cenozoic sea-level fluctuations on the bivalve faunas of tropical oceanic islands. Paleobiology, 16: 415-434.
92. Pauly, G., 1991. Late Cenozoic sea level fluctuations and the diversity and species composition of insular shallow water marine faunas. In: E.C. Dudley (Editor), The Unity of Evolutionary Biology. Dioscorides Press, Portland, pp. 184-193.
93. Petuch, E.J., 1976. An unusual molluscan assemblage from Venezuela. Veliger, 18: 322-325.
94. Petuch, E.J., 1981. A relict Neogene caenogastropod fauna from northern South America. Malacologia, 20: 307-347.
95. Petuch, E.J., 1982. Geographical heterochrony: contemporaneous coexistence of Neogene and Recent molluscan faunas in the Americas. Palaeogeogr. Palaeoclimatol, Palaeoecol., 37: 277-312.
96. Plotnick, R.E., McKinney, M. and Gardner, R., 1994. Multiscale ecosystem stasis and multiscale disturbances: the thousand natural shocks that flesh is heir to. Geol. Soc. Am. Abstr. with Programs, 26(7): A-520.
97. Pulliam, H.R., 1988. Sources, sinks, and population regulation. Am. Nat., 132: 652-661.
98. Rascoe, B., 1962. Regional stratigraphic analysis of Pennsylva-nian and Permian rocks in western mid-continent, Colorado, Kansas, Oklahoma, Texas. AAPG Bull., 46: 1345-1370.
99. Rascoe, B., 1975. Tectonic origin of preconsolidation deformation in Upper Pennsylvanian rocks near Bartlesville, Oklahoma. AAPG Bull., 59: 1626-1638.
100. Rohlf, F.J., 1989. NTSYS-pc, version 1.50, Exeter Publ., Setauket.
101. Roughgarden, J., Pennington, T., and Alexander, S., 1994. Dynamics of the rocky intertidal zone with remarks on generalization in ecology. Philos. Trans. R. Soc. Lond. B, 343: 79-85.
102. SAS Institute Inc., 1989. SAS/STAT User's Guide, Version 6, 4th Ed., Vol. 1. SAS Institute Inc., Cary, NC, 943 pp.
103. Savarese, M., Gray, L.M. and Brett, C.E., 1986. Faunal and lithologic cyclicity in the Centerfield Member ( Middle Devonian, Hamilton Group) of Western New York: a reinterpretation of depositional history. In: C.E. Brett (Editor), Dynamic Stratigraphy and Depositional Environments of the Hamilton Group (Middle Devonian) in New York State, Part I. N.Y. State Mus. Bull., 457: 32-56.
104. Schneider, J.A., 1988. Frequency of arm regeneration of comatulid crinoids in relation to life habit. In: R.D. Burke et al. (Editors), Echinoderm Biology. Balkema, Rotterdam, pp. 531-538.
105. Schopf, K.M., 1996. Coordinated stasis: biofacies revisited and the conceptual modeling of whole-fauna dynamics. In: L.C. Ivany and K.M. Schopf (Editors), New Perspectives on Faunal Stability in the Fossil Record. Palaeogeogr. Palaeoclimatol. Palaeoecol., 127: 157-175.
106. Schopf, K.M., Ivany, L.C. and Morris, P.J., 1992. Onshore-offshore trends in light of ecological locking. Geol. Soc. Am. Abstr. with Programs, 24 (7):A313.
107. Schubert, J.K. and Bottjer, D.J., 1995. Aftermath of the Permian-Triassic mass extinction event: paleoecology of Lower Triassic carbonates in the western USA. Palaeogeogr. Palaeoclimatol. Palaeoecol., 116: 1-39.
108. Sheehan, P.M., 1991. Patterns of synecology during the Phanerozoic. In: E.C. Dudley (Editor), The Unity of Evolutionary Biology. Dioscorides Press, Portland, pp. 103-118.
109. Sheldon, P.R., 1993. Making sense of microevolutionary patterns. In: D.R. Lees and D. Edwards (Editors), Evolutionary Patterns and Processes. Academic Press, London, pp. 19-31.
110. Smith, T.M. and Urban, D.L., 1988. Scale and resolution of forest structural pattern. Vegetatio, 74: 143-150.
111. Sneath, P.H.A. and Sokal, R.R., 1973. Numerical Taxonomy. Freeman. San Francisco, 573 pp.
112. Sokal, R.R. and Rohlf, F.J., 1981. Biometry. Freeman, San Francisco, 859 pp.
113. Sorensen, J.H., Johnsgard, S.J. and Wozencraft, C., 1989. Bibliography of Kansas Geology, 1823-1984. Kans. Geol. Surv. Bull., 221, 416 pp.
114. Springer, D.A. and Bambach, R.K., 1985. Gradient versus cluster analysis of fossil assemblages: a comparison from the Ordovician of southwestern Virginia. Lethaia, 18: 181-198.
115. Sutherland, J.P., 1974. Multiple stable points in natural communities. Am. Nat., 108: 859-873.
116. SYSTAT, 1986. Version 3, Evanston, IL, 417 pp.
117. Taylor, J.D., 1978. Faunal response to the instability of reef habitats: Pleistocene molluscan assemblages of Aldabra Atoll. Palaeontology, 21: 1-30.
118. Thompson, J.N., 1994. The Coevolutionary Process. Univ, Chicago Press, 376 pp.
119. Underwood, A.J. and Fairweather, P.G., 1989. Supply-side ecology and benthic marine assemblages. TREE, 4: 16-20.
120. Underwood, M.R., 1984. Stratigraphy and depositional history of the Bourbon Flags (Upper Pennsylvanian), an enigmatic unit in Linn and Bourbon Counties in east-Central Kansas. Thesis. Univ. Iowa, Iowa City, 111 pp. (unpublished).
121. Valentine, J.W. and Jablonski, D., 1993. Fossil communities: compositional variation at many time scales. In: R.E. Ricklefs and D. Schluter (Editors), Species Diversity in Ecological Communities. Univ. Chicago Press, pp. 341-349.
122. Vermeij, G.J., 1987. Evolution and Escalation: An Ecological History of Life. Princeton Univ. Press, 527 pp.
123. Warshauer, S.M. and Smosna, R., 1981. On minimum spanning trees and the intergradation of clusters. Math. Geol., 13: 225-235.
124. Watney, W.L., Kaesler, R.L. and Newell, K.D., 1985. Recent interpretations of Late Paleozoic cyclothems. In: Mid-continent Sect. SEPM 3rd Annu. Field Conf. Guidebook, 271 pp.
125. Watney, W.L., French, J.A. and Franseen, E.K., 1989. Sequence stratigraphic interpretations and modeling of cyclothems. In: Kans. Geol. Soc. 41st Annu. Field Conf. Guidebook, 211 pp.
126. Watt, A.S., 1947. Pattern and process in the plant community. J. Ecol., 35: 1-22.
127. Webster, G.D., 1981. New crinoids from the Naco Formation (Middle Pennsylvanian) of Arizona and a revision of the Family Cromyocrinidae. J. Paleontol., 55: 1176-1199.