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Volumn 119, Issue 5, 2013, Pages 436-447

Assessing the Effects of Rearing Environment, Natural Selection, and Developmental Stage on the Emergence of a Behavioral Syndrome

Author keywords

[No Author keywords available]

Indexed keywords

AGGRESSION; DEVELOPMENTAL STAGE; EMERGENCE; HERITABILITY; MARK-RECAPTURE METHOD; NATURAL SELECTION; PHENOTYPIC PLASTICITY; REARING; SPIDER; WILD POPULATION;

EID: 84876121125     PISSN: 01791613     EISSN: 14390310     Source Type: Journal    
DOI: 10.1111/eth.12081     Document Type: Article
Times cited : (49)

References (58)
  • 1
    • 54249168349 scopus 로고    scopus 로고
    • Acoustic experience shapes female mate choice in field crickets
    • Bailey, N. W. & Zuk, M. 2008: Acoustic experience shapes female mate choice in field crickets. Proc. R. Soc. B Biol. Sci. 275, 2645-2650.
    • (2008) Proc. R. Soc. B Biol. Sci. , vol.275 , pp. 2645-2650
    • Bailey, N.W.1    Zuk, M.2
  • 2
    • 14844356173 scopus 로고    scopus 로고
    • Behavioural differences between individuals and two populations of stickleback (Gasterosteus aculeatus)
    • Bell, A. M. 2005: Behavioural differences between individuals and two populations of stickleback (Gasterosteus aculeatus). J. Evol. Biol. 18, 464-473.
    • (2005) J. Evol. Biol. , vol.18 , pp. 464-473
    • Bell, A.M.1
  • 3
    • 78650606994 scopus 로고    scopus 로고
    • What can whole genome expression data tell us about the ecology and evolution of personality?
    • Bell, A. M. & Aubin-Horth, N. 2010: What can whole genome expression data tell us about the ecology and evolution of personality? Philos. Trans. R. Soc. Lond. B Biol. Sci. 365, 4001-4012.
    • (2010) Philos. Trans. R. Soc. Lond. B Biol. Sci. , vol.365 , pp. 4001-4012
    • Bell, A.M.1    Aubin-Horth, N.2
  • 4
    • 79957969264 scopus 로고    scopus 로고
    • Behavior and the dynamic genome
    • Bell, A. M. & Robinson, G. E. 2011: Behavior and the dynamic genome. Science 332, 1161-1162.
    • (2011) Science , vol.332 , pp. 1161-1162
    • Bell, A.M.1    Robinson, G.E.2
  • 5
    • 34547422785 scopus 로고    scopus 로고
    • Exposure to predation generates personality in three spined sticklebacks (Gasterosteus aculeatus)
    • Bell, A. M. & Sih, A. 2007: Exposure to predation generates personality in three spined sticklebacks (Gasterosteus aculeatus). Ecol. Lett. 10, 828-834.
    • (2007) Ecol. Lett. , vol.10 , pp. 828-834
    • Bell, A.M.1    Sih, A.2
  • 6
    • 84865369783 scopus 로고    scopus 로고
    • Sexual cannibalism is associated with female behavioural type, hunger state and increased hatching success
    • Berning, A. W., Gadd, R. D. H., Sweeney, K., MacDonald, L., Eng, R. Y. Y., Hess, Z. L. & Pruitt, J. N. 2012: Sexual cannibalism is associated with female behavioural type, hunger state and increased hatching success. Anim. Behav. 84, 715-721.
    • (2012) Anim. Behav. , vol.84 , pp. 715-721
    • Berning, A.W.1    Gadd, R.D.H.2    Sweeney, K.3    MacDonald, L.4    Eng, R.Y.Y.5    Hess, Z.L.6    Pruitt, J.N.7
  • 7
    • 77957907224 scopus 로고    scopus 로고
    • Do consistent individual differences in metabolic rate promote consistent individual differences in behavior?
    • Biro, P. A. & Stamps, J. A. 2010: Do consistent individual differences in metabolic rate promote consistent individual differences in behavior? Trends Ecol. Evol. 25, 653-659.
    • (2010) Trends Ecol. Evol. , vol.25 , pp. 653-659
    • Biro, P.A.1    Stamps, J.A.2
  • 8
    • 0024877695 scopus 로고
    • Repeatability - its role in evolutionary studies of mating-behavior
    • Boake, C. R. B. 1989: Repeatability - its role in evolutionary studies of mating-behavior. Evol. Ecol. 3, 173-182.
    • (1989) Evol. Ecol. , vol.3 , pp. 173-182
    • Boake, C.R.B.1
  • 10
    • 35748944103 scopus 로고    scopus 로고
    • The quick and the dead: correlational selection on morphology, performance, and habitat use in island lizards
    • Calsbeek, R. & Irschick, D. J. 2007: The quick and the dead: correlational selection on morphology, performance, and habitat use in island lizards. Evolution 61, 2493-2503.
    • (2007) Evolution , vol.61 , pp. 2493-2503
    • Calsbeek, R.1    Irschick, D.J.2
  • 11
    • 77952054900 scopus 로고    scopus 로고
    • The pace of life under artificial selection: personality, energy expenditure, and longevity are correlated in domestic dogs
    • Careau, V., Reale, D., Humphries, M. M. & Thomas, D. W. 2010: The pace of life under artificial selection: personality, energy expenditure, and longevity are correlated in domestic dogs. Am. Nat. 175, 753-758.
    • (2010) Am. Nat. , vol.175 , pp. 753-758
    • Careau, V.1    Reale, D.2    Humphries, M.M.3    Thomas, D.W.4
  • 12
    • 0027883920 scopus 로고
    • Dynamic-models of behavior - an extension of life-history theory
    • Clark, C. W. 1993: Dynamic-models of behavior - an extension of life-history theory. Trends Ecol. Evol. 8, 205-209.
    • (1993) Trends Ecol. Evol. , vol.8 , pp. 205-209
    • Clark, C.W.1
  • 13
    • 0028178681 scopus 로고
    • Antipredator behavior and the asset-protection principle
    • Clark, C. W. 1994: Antipredator behavior and the asset-protection principle. Behav. Ecol. 5, 159-170.
    • (1994) Behav. Ecol. , vol.5 , pp. 159-170
    • Clark, C.W.1
  • 14
    • 33747326883 scopus 로고    scopus 로고
    • Natural selection and animal personality
    • Dingemanse, N. J. & Reale, D. 2005: Natural selection and animal personality. Behaviour 142, 1159-1184.
    • (2005) Behaviour , vol.142 , pp. 1159-1184
    • Dingemanse, N.J.1    Reale, D.2
  • 15
    • 35148875040 scopus 로고    scopus 로고
    • Behavioural syndromes differ predictably between 12 populations of three-spined stickleback
    • Dingemanse, N. J., Wright, J., Kazem, A. J. N., Thomas, D. K., Hickling, R. & Dawnay, N. 2007: Behavioural syndromes differ predictably between 12 populations of three-spined stickleback. J. Anim. Ecol. 76, 1128-1138.
    • (2007) J. Anim. Ecol. , vol.76 , pp. 1128-1138
    • Dingemanse, N.J.1    Wright, J.2    Kazem, A.J.N.3    Thomas, D.K.4    Hickling, R.5    Dawnay, N.6
  • 16
    • 84866904961 scopus 로고    scopus 로고
    • Juvenile exposure to acoustic sexual signals from conspecifics alters growth trajectory and an adult personality trait
    • DiRienzo, N., Pruitt, J. N. & Hedrick, A. V. 2012: Juvenile exposure to acoustic sexual signals from conspecifics alters growth trajectory and an adult personality trait. Anim. Behav. 84, 861-868.
    • (2012) Anim. Behav. , vol.84 , pp. 861-868
    • DiRienzo, N.1    Pruitt, J.N.2    Hedrick, A.V.3
  • 17
    • 33749530852 scopus 로고    scopus 로고
    • Behavioral correlations across breeding contexts provide a mechanism for a cost of aggression
    • Duckworth, R. A. 2006: Behavioral correlations across breeding contexts provide a mechanism for a cost of aggression. Behav. Ecol. 17, 1011-1019.
    • (2006) Behav. Ecol. , vol.17 , pp. 1011-1019
    • Duckworth, R.A.1
  • 19
    • 0024875653 scopus 로고
    • Genetically-based variation between 2 spider populations in foraging behavior
    • Hedrick, A. V. & Riechert, S. E. 1989: Genetically-based variation between 2 spider populations in foraging behavior. Oecologia 80, 533-539.
    • (1989) Oecologia , vol.80 , pp. 533-539
    • Hedrick, A.V.1    Riechert, S.E.2
  • 20
    • 84855220133 scopus 로고    scopus 로고
    • Individual deviation from behavioural correlations: a simple approach to study the evolution of behavioural syndromes
    • Herczeg, G. & Garamszegi, L. Z. 2012: Individual deviation from behavioural correlations: a simple approach to study the evolution of behavioural syndromes. Behav. Ecol. Sociobiol. 66, 161-169.
    • (2012) Behav. Ecol. Sociobiol. , vol.66 , pp. 161-169
    • Herczeg, G.1    Garamszegi, L.Z.2
  • 21
    • 60649106034 scopus 로고    scopus 로고
    • Predation mediated population divergence in complex behaviour of nine-spined stickleback (Pungitius pungitius)
    • Herczeg, G., Gonda, A. & Merila, J. 2009: Predation mediated population divergence in complex behaviour of nine-spined stickleback (Pungitius pungitius). J. Evol. Biol. 22, 544-552.
    • (2009) J. Evol. Biol. , vol.22 , pp. 544-552
    • Herczeg, G.1    Gonda, A.2    Merila, J.3
  • 22
    • 0032443351 scopus 로고    scopus 로고
    • Logistic regression for empirical studies of multivariate selection
    • Janzen, F. J. & Stern, H. S. 1998: Logistic regression for empirical studies of multivariate selection. Evolution 52, 1564-1571.
    • (1998) Evolution , vol.52 , pp. 1564-1571
    • Janzen, F.J.1    Stern, H.S.2
  • 23
    • 22844446041 scopus 로고    scopus 로고
    • Precopulatory sexual cannibalism in fishing spiders (Dolomedes triton): a role for behavioral syndromes
    • Johnson, J. C. & Sih, A. 2005: Precopulatory sexual cannibalism in fishing spiders (Dolomedes triton): a role for behavioral syndromes. Behav. Ecol. Sociobiol. 58, 390-396.
    • (2005) Behav. Ecol. Sociobiol. , vol.58 , pp. 390-396
    • Johnson, J.C.1    Sih, A.2
  • 24
    • 35748932031 scopus 로고    scopus 로고
    • Fear, food, sex and parental care: A syndrome of boldness in the fishing spider, Dolomedes triton
    • Johnson, J. C. & Sih, A. 2007: Fear, food, sex and parental care: A syndrome of boldness in the fishing spider, Dolomedes triton. Anim. Behav. 74, 1131-1138.
    • (2007) Anim. Behav. , vol.74 , pp. 1131-1138
    • Johnson, J.C.1    Sih, A.2
  • 25
    • 78650726498 scopus 로고    scopus 로고
    • Does personality in small rodents vary depending on population density?
    • Korpela, K., Sundell, J. & Ylonen, H. 2011: Does personality in small rodents vary depending on population density? Oecologia 165, 67-77.
    • (2011) Oecologia , vol.165 , pp. 67-77
    • Korpela, K.1    Sundell, J.2    Ylonen, H.3
  • 26
    • 78649554566 scopus 로고    scopus 로고
    • Parasitism, predation and the evolution of animal personalities
    • Kortet, R., Hedrick, A. V. & Vainikka, A. 2010: Parasitism, predation and the evolution of animal personalities. Ecol. Lett. 13, 1449-1458.
    • (2010) Ecol. Lett. , vol.13 , pp. 1449-1458
    • Kortet, R.1    Hedrick, A.V.2    Vainikka, A.3
  • 27
    • 5344227529 scopus 로고    scopus 로고
    • Apprehension and time allocation in gerbils: the effects of predatory risk and energetic state
    • Kotler, B. P., Brown, J. S. & Bouskila, A. 2004: Apprehension and time allocation in gerbils: the effects of predatory risk and energetic state. Ecology 85, 917-922.
    • (2004) Ecology , vol.85 , pp. 917-922
    • Kotler, B.P.1    Brown, J.S.2    Bouskila, A.3
  • 28
    • 84862638369 scopus 로고    scopus 로고
    • Individual behavioural consistency and plasticity in an urban spider
    • Kralj-Fiser, S. & Schneider, J. M. 2012: Individual behavioural consistency and plasticity in an urban spider. Anim. Behav. 84, 197-204.
    • (2012) Anim. Behav. , vol.84 , pp. 197-204
    • Kralj-Fiser, S.1    Schneider, J.M.2
  • 30
    • 0001214721 scopus 로고
    • The measurement of selection on correlated characters
    • Lande, R. & Arnold, S. J. 1983: The measurement of selection on correlated characters. Evolution 37, 1210-1226.
    • (1983) Evolution , vol.37 , pp. 1210-1226
    • Lande, R.1    Arnold, S.J.2
  • 31
    • 78650597371 scopus 로고    scopus 로고
    • Risk, resources and state- dependent adaptive behavioural syndromes
    • Luttbeg, B. & Sih, A. 2010: Risk, resources and state- dependent adaptive behavioural syndromes. Philos. Trans. R. Soc. Lond. B Biol. Sci. 365, 3977-3990.
    • (2010) Philos. Trans. R. Soc. Lond. B Biol. Sci. , vol.365 , pp. 3977-3990
    • Luttbeg, B.1    Sih, A.2
  • 32
    • 33750296011 scopus 로고    scopus 로고
    • How noisy information and individual asymmetries can make 'personality' an adaptation: a simple model
    • McElreath, R. & Strimling, P. 2006: How noisy information and individual asymmetries can make 'personality' an adaptation: a simple model. Anim. Behav. 72, 1135-1139.
    • (2006) Anim. Behav. , vol.72 , pp. 1135-1139
    • McElreath, R.1    Strimling, P.2
  • 33
    • 84862678743 scopus 로고    scopus 로고
    • Predator-induced changes in the boldness of naive field crickets, Gryllus integer, depends on behavioural type
    • Niemela, P. T., DiRienzo, N. & Hedrick, A. V. 2012: Predator-induced changes in the boldness of naive field crickets, Gryllus integer, depends on behavioural type. Anim. Behav. 84, 129-135.
    • (2012) Anim. Behav. , vol.84 , pp. 129-135
    • Niemela, P.T.1    DiRienzo, N.2    Hedrick, A.V.3
  • 34
    • 79953313786 scopus 로고    scopus 로고
    • Correlated response to selection of testosterone levels and immunocompetence in lines selected for avian personality
    • van Oers, K., Buchanan, K. L., Thomas, T. E. & Drent, P. J. 2011: Correlated response to selection of testosterone levels and immunocompetence in lines selected for avian personality. Anim. Behav. 81, 1055-1061.
    • (2011) Anim. Behav. , vol.81 , pp. 1055-1061
    • van Oers, K.1    Buchanan, K.L.2    Thomas, T.E.3    Drent, P.J.4
  • 35
    • 0032128294 scopus 로고    scopus 로고
    • Management of fat reserves in tufted titmice Baelophus bicolor in relation to risk of predation
    • Pravosudov, V. V. & Grubb, T. C. 1998: Management of fat reserves in tufted titmice Baelophus bicolor in relation to risk of predation. Anim. Behav. 56, 49-54.
    • (1998) Anim. Behav. , vol.56 , pp. 49-54
    • Pravosudov, V.V.1    Grubb, T.C.2
  • 36
    • 73949135023 scopus 로고    scopus 로고
    • Context-dependent running speed in funnel-web spiders from divergent populations
    • Pruitt, J. N. & Husak, J. F. 2010: Context-dependent running speed in funnel-web spiders from divergent populations. Funct. Ecol. 24, 165-171.
    • (2010) Funct. Ecol. , vol.24 , pp. 165-171
    • Pruitt, J.N.1    Husak, J.F.2
  • 37
    • 77956593300 scopus 로고    scopus 로고
    • The adaptive value of gluttony: predators mediate the life history trade-offs of satiation threshold
    • Pruitt, J. N. & Krauel, J. J. 2010: The adaptive value of gluttony: predators mediate the life history trade-offs of satiation threshold. J. Evol. Biol. 23, 2104-2111.
    • (2010) J. Evol. Biol. , vol.23 , pp. 2104-2111
    • Pruitt, J.N.1    Krauel, J.J.2
  • 38
    • 67349147912 scopus 로고    scopus 로고
    • Sex matters: sexually dimorphic fitness consequences of a behavioural syndrome
    • Pruitt, J. N. & Riechert, S. E. 2009: Sex matters: sexually dimorphic fitness consequences of a behavioural syndrome. Anim. Behav. 78, 175-181.
    • (2009) Anim. Behav. , vol.78 , pp. 175-181
    • Pruitt, J.N.1    Riechert, S.E.2
  • 39
    • 84863910998 scopus 로고    scopus 로고
    • The ecological consequences of temperament in spiders
    • Pruitt, J. N. & Riechert, S. E. 2012: The ecological consequences of temperament in spiders. Curr Zool 58, 589-596.
    • (2012) Curr Zool , vol.58 , pp. 589-596
    • Pruitt, J.N.1    Riechert, S.E.2
  • 40
    • 49149087502 scopus 로고    scopus 로고
    • Behavioural syndromes and their fitness consequences in a socially polymorphic spider, Anelosimus studiosus
    • Pruitt, J. N., Riechert, S. E. & Jones, T. C. 2008: Behavioural syndromes and their fitness consequences in a socially polymorphic spider, Anelosimus studiosus. Anim. Behav. 76, 871-879.
    • (2008) Anim. Behav. , vol.76 , pp. 871-879
    • Pruitt, J.N.1    Riechert, S.E.2    Jones, T.C.3
  • 41
    • 77952540403 scopus 로고    scopus 로고
    • Population differences in behaviour are explained by shared within-population trait correlations
    • Pruitt, J. N., Riechert, S. E., Iturralde, G., Vega, M., Fitzpatrick, B. M. & Aviles, L. 2010: Population differences in behaviour are explained by shared within-population trait correlations. J. Evol. Biol. 23, 748-756.
    • (2010) J. Evol. Biol. , vol.23 , pp. 748-756
    • Pruitt, J.N.1    Riechert, S.E.2    Iturralde, G.3    Vega, M.4    Fitzpatrick, B.M.5    Aviles, L.6
  • 42
    • 82355175219 scopus 로고    scopus 로고
    • Amazonian social spiders share similar within-colony behavioural variation and behavioural syndromes
    • Pruitt, J. N., Iturralde, G., Aviles, L. & Riechert, S. E. 2011: Amazonian social spiders share similar within-colony behavioural variation and behavioural syndromes. Anim. Behav. 82, 1449-1455.
    • (2011) Anim. Behav. , vol.82 , pp. 1449-1455
    • Pruitt, J.N.1    Iturralde, G.2    Aviles, L.3    Riechert, S.E.4
  • 45
    • 0000089322 scopus 로고
    • Web-site selection in desert spider Agelenopsis aperta
    • Riechert, S. E. 1976: Web-site selection in desert spider Agelenopsis aperta. Oikos 27, 311-315.
    • (1976) Oikos , vol.27 , pp. 311-315
    • Riechert, S.E.1
  • 46
    • 0002255059 scopus 로고
    • The evolution of behavioral phenotypes - lessons learned from divergent spider populations
    • Riechert, S. E. 1993: The evolution of behavioral phenotypes - lessons learned from divergent spider populations. In: Advances in the Study of Behavior, vol 22. pp. 103-134.
    • (1993) Advances in the Study of Behavior , vol.22 , pp. 103-134
    • Riechert, S.E.1
  • 47
    • 0027843856 scopus 로고
    • A test for correlations among fitness-linked behavioral traits in the spider Agelenopsis aperta (Araneae, Agelenidae)
    • Riechert, S. E. & Hedrick, A. V. 1993: A test for correlations among fitness-linked behavioral traits in the spider Agelenopsis aperta (Araneae, Agelenidae). Anim. Behav. 46, 669-675.
    • (1993) Anim. Behav. , vol.46 , pp. 669-675
    • Riechert, S.E.1    Hedrick, A.V.2
  • 48
    • 0037626707 scopus 로고    scopus 로고
    • Do female spiders select heavier males for the genes for behavioral aggressiveness they offer their offspring?
    • Riechert, S. E. & Johns, P. M. 2003: Do female spiders select heavier males for the genes for behavioral aggressiveness they offer their offspring? Evolution 57, 1367-1373.
    • (2003) Evolution , vol.57 , pp. 1367-1373
    • Riechert, S.E.1    Johns, P.M.2
  • 49
    • 0024484474 scopus 로고
    • Genetic analyses of 2 behavioral traits linked to individual fitness in the desert spider Agelenopsis aperta
    • Riechert, S. E. & Smith, J. M. 1989: Genetic analyses of 2 behavioral traits linked to individual fitness in the desert spider Agelenopsis aperta. Anim. Behav. 37, 624-637.
    • (1989) Anim. Behav. , vol.37 , pp. 624-637
    • Riechert, S.E.1    Smith, J.M.2
  • 50
    • 0035708875 scopus 로고    scopus 로고
    • High gene flow levels lead to gamete wastage in a desert spider system
    • Riechert, S. E., Singer, F. D. & Jones, T. C. 2001: High gene flow levels lead to gamete wastage in a desert spider system. Genetica 112, 297-319.
    • (2001) Genetica , vol.112 , pp. 297-319
    • Riechert, S.E.1    Singer, F.D.2    Jones, T.C.3
  • 52
    • 78650329144 scopus 로고    scopus 로고
    • Response to environmental change in rainbow trout selected for divergent stress coping styles
    • Ruiz-Gomez, M. D., Huntingford, F. A., Overli, O., Thornqvist, P. O. & Hoglund, E. 2011: Response to environmental change in rainbow trout selected for divergent stress coping styles. Physiol. Behav. 102, 317-322.
    • (2011) Physiol. Behav. , vol.102 , pp. 317-322
    • Ruiz-Gomez, M.D.1    Huntingford, F.A.2    Overli, O.3    Thornqvist, P.O.4    Hoglund, E.5
  • 53
    • 2942742936 scopus 로고    scopus 로고
    • Behavioral syndromes: an ecological and evolutionary overview
    • Sih, A., Bell, A. & Johnson, J. C. 2004a: Behavioral syndromes: an ecological and evolutionary overview. Trends Ecol. Evol. 19, 372-378.
    • (2004) Trends Ecol. Evol. , vol.19 , pp. 372-378
    • Sih, A.1    Bell, A.2    Johnson, J.C.3
  • 54
    • 2942757889 scopus 로고    scopus 로고
    • Behavioral syndromes: an integrative overview
    • Sih, A., Bell, A. M., Johnson, J. C. & Ziemba, R. E. 2004b: Behavioral syndromes: an integrative overview. Q. Rev. Biol. 79, 241-277.
    • (2004) Q. Rev. Biol. , vol.79 , pp. 241-277
    • Sih, A.1    Bell, A.M.2    Johnson, J.C.3    Ziemba, R.E.4
  • 55
  • 56
    • 77950817905 scopus 로고    scopus 로고
    • The development of animal personality: relevance, concepts and perspectives
    • Stamps, J. & Groothuis, T. G. G. 2010a: The development of animal personality: relevance, concepts and perspectives. Biol. Rev. 85, 301-325.
    • (2010) Biol. Rev. , vol.85 , pp. 301-325
    • Stamps, J.1    Groothuis, T.G.G.2
  • 57
    • 78650593550 scopus 로고    scopus 로고
    • Developmental perspectives on personality: implications for ecological and evolutionary studies of individual differences
    • Stamps, J. A. & Groothuis, T. G. G. 2010b: Developmental perspectives on personality: implications for ecological and evolutionary studies of individual differences. Philos. Trans. R. Soc. Lond. B Biol. Sci. 365, 4029-4041.
    • (2010) Philos. Trans. R. Soc. Lond. B Biol. Sci. , vol.365
    • Stamps, J.A.1    Groothuis, T.G.G.2
  • 58
    • 51249121392 scopus 로고    scopus 로고
    • Estimating nonlinear selection gradients using quadratic regression coefficients: double or nothing ?
    • Stinchcombe, J. R., Agrawal, A. F., Hohenlohe, P. A., Arnold, S. J. & Blows, M. W. 2008: Estimating nonlinear selection gradients using quadratic regression coefficients: double or nothing ? Evolution 62, 2435-2440.
    • (2008) Evolution , vol.62 , pp. 2435-2440
    • Stinchcombe, J.R.1    Agrawal, A.F.2    Hohenlohe, P.A.3    Arnold, S.J.4    Blows, M.W.5


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