메뉴 건너뛰기




Volumn 82, Issue 2, 2013, Pages 418-428

Pigments versus structure: Examining the mechanism of age-dependent change in a carotenoid-based colour

Author keywords

Age dichromatism; Aging; Carotenoid availability; Great tit; Within individual change

Indexed keywords

AZEMIOPS FEAE VENOM; CAROTENOID; PIGMENT; VIPER VENOM;

EID: 84874000659     PISSN: 00218790     EISSN: 13652656     Source Type: Journal    
DOI: 10.1111/1365-2656.12008     Document Type: Article
Times cited : (22)

References (66)
  • 1
    • 28044443946 scopus 로고    scopus 로고
    • Quantifying colors
    • eds G.E. Hill & K.J. McGraw) - Harvard University Press, Cambridge, Massachusetts.
    • Andersson, S. & Prager, M. (2006) Quantifying colors. Bird Coloration. Volume I: Mechanisms and Measurements (eds G.E. Hill & K.J. McGraw), pp. 41-89. Harvard University Press, Cambridge, Massachusetts.
    • (2006) Bird Coloration. Volume I: Mechanisms and Measurements , pp. 41-89
    • Andersson, S.1    Prager, M.2
  • 2
    • 0003203292 scopus 로고
    • The significance of variations in body weight and wing length in the great tit, Parus major
    • van Balen, J.H. (1967) The significance of variations in body weight and wing length in the great tit, Parus major. Ardea, 55, 1-59.
    • (1967) Ardea , vol.55 , pp. 1-59
    • van Balen, J.H.1
  • 5
    • 34948825378 scopus 로고    scopus 로고
    • Exploring the genetics of aging in a wild passerine bird
    • Brommer, J.E., Wilson, A.J. & Gustafsson, L. (2007) Exploring the genetics of aging in a wild passerine bird. American Naturalist, 170, 643-650.
    • (2007) American Naturalist , vol.170 , pp. 643-650
    • Brommer, J.E.1    Wilson, A.J.2    Gustafsson, L.3
  • 6
    • 78651518941 scopus 로고    scopus 로고
    • How many color metrics do we need? Evaluating how different color-scoring procedures explain carotenoid pigment content in avian bare-part and plumage ornaments
    • Butler, M.W., Toomey, M.B. & McGraw, K.J. (2011) How many color metrics do we need? Evaluating how different color-scoring procedures explain carotenoid pigment content in avian bare-part and plumage ornaments. Behavioral Ecology and Sociobiology, 65, 401-413.
    • (2011) Behavioral Ecology and Sociobiology , vol.65 , pp. 401-413
    • Butler, M.W.1    Toomey, M.B.2    McGraw, K.J.3
  • 7
    • 0036149163 scopus 로고    scopus 로고
    • Individual covariation in life-history traits: seeing the trees despite the forest
    • Cam, E., Link, W.A., Cooch, E., Monnat, J.-Y. & Danchin, E. (2002) Individual covariation in life-history traits: seeing the trees despite the forest. American Naturalist, 159, 96-105.
    • (2002) American Naturalist , vol.159 , pp. 96-105
    • Cam, E.1    Link, W.A.2    Cooch, E.3    Monnat, J.-Y.4    Danchin, E.5
  • 8
    • 0034619844 scopus 로고    scopus 로고
    • Changes in expression and honesty of sexual signalling over the reproductive lifetime of sticklebacks
    • Candolin, U. (2000) Changes in expression and honesty of sexual signalling over the reproductive lifetime of sticklebacks. Proceedings of the Royal Society of London, Series B, 267, 2425-2430.
    • (2000) Proceedings of the Royal Society of London, Series B , vol.267 , pp. 2425-2430
    • Candolin, U.1
  • 10
    • 75249105687 scopus 로고    scopus 로고
    • Age-dependent allocation of carotenoids to coloration versus antioxidant defences
    • Cote, J., Arnoux, E., Sorci, G., Gaillard, M. & Faivre, B. (2010) Age-dependent allocation of carotenoids to coloration versus antioxidant defences. Journal of Experimental Biology, 213, 271-277.
    • (2010) Journal of Experimental Biology , vol.213 , pp. 271-277
    • Cote, J.1    Arnoux, E.2    Sorci, G.3    Gaillard, M.4    Faivre, B.5
  • 12
    • 33847743337 scopus 로고    scopus 로고
    • Intraspecific variation in coloration
    • eds G.E. Hill & K.J. McGraw) - Harvard University Press, Cambridge, Massachusetts.
    • Dale, J. (2006) Intraspecific variation in coloration. Bird Coloration. Volume 2: Function and Evolution (eds G.E. Hill & K.J. McGraw), pp. 36-86. Harvard University Press, Cambridge, Massachusetts.
    • (2006) Bird Coloration. Volume 2: Function and Evolution , pp. 36-86
    • Dale, J.1
  • 14
    • 33747819803 scopus 로고    scopus 로고
    • Age differences in blue tit Parus caeruleus plumage colour: within-individual changes or colour-biased survival?
    • Delhey, K. & Kempenaers, B. (2006) Age differences in blue tit Parus caeruleus plumage colour: within-individual changes or colour-biased survival? Journal of Avian Biology, 37, 339-348.
    • (2006) Journal of Avian Biology , vol.37 , pp. 339-348
    • Delhey, K.1    Kempenaers, B.2
  • 15
    • 77953323840 scopus 로고    scopus 로고
    • Carotenoid composition of invertebrates consumed by two insectivorous bird species
    • Eeva, T., Helle, S., Salminen, J.-P. & Hakkarainen, H. (2010) Carotenoid composition of invertebrates consumed by two insectivorous bird species. Journal of Chemical Ecology, 36, 608-613.
    • (2010) Journal of Chemical Ecology , vol.36 , pp. 608-613
    • Eeva, T.1    Helle, S.2    Salminen, J.-P.3    Hakkarainen, H.4
  • 16
    • 0021084757 scopus 로고
    • Natural and sexual selection on color patterns in poeciliid fishes
    • Endler, J.A. (1983) Natural and sexual selection on color patterns in poeciliid fishes. Environmental Biology of Fishes, 9, 173-190.
    • (1983) Environmental Biology of Fishes , vol.9 , pp. 173-190
    • Endler, J.A.1
  • 17
    • 79958061869 scopus 로고    scopus 로고
    • Divergent patterns of age-dependence in ornamental and reproductive traits in the collared flycatcher
    • Evans, S.R., Gustafsson, L. & Sheldon, B.C. (2011) Divergent patterns of age-dependence in ornamental and reproductive traits in the collared flycatcher. Evolution, 65, 1623-1636.
    • (2011) Evolution , vol.65 , pp. 1623-1636
    • Evans, S.R.1    Gustafsson, L.2    Sheldon, B.C.3
  • 18
    • 83655164468 scopus 로고    scopus 로고
    • Quantitative genetics of a carotenoid-based colour: heritability and persistent natal environmental effects in the great tit
    • Evans, S.R. & Sheldon, B.C. (2012) Quantitative genetics of a carotenoid-based colour: heritability and persistent natal environmental effects in the great tit. American Naturalist, 179, 79-94.
    • (2012) American Naturalist , vol.179 , pp. 79-94
    • Evans, S.R.1    Sheldon, B.C.2
  • 19
    • 84862333057 scopus 로고    scopus 로고
    • Seasonality of carotenoid-based plumage coloration: modelling wavelength-specific change through spectral reconstruction
    • Evans, S.R., Summers, A.G.R. & Sheldon, B.C. (2012) Seasonality of carotenoid-based plumage coloration: modelling wavelength-specific change through spectral reconstruction. Journal of Avian Biology, 43, 234-243.
    • (2012) Journal of Avian Biology , vol.43 , pp. 234-243
    • Evans, S.R.1    Summers, A.G.R.2    Sheldon, B.C.3
  • 22
    • 2342655777 scopus 로고    scopus 로고
    • Species-specific differences in oak foliage affect preference and performance of gypsy moth caterpillars
    • Foss, L.K. & Rieske, L.K. (2003) Species-specific differences in oak foliage affect preference and performance of gypsy moth caterpillars. Entomologia Experimentalis et Applicata, 108, 87-93.
    • (2003) Entomologia Experimentalis et Applicata , vol.108 , pp. 87-93
    • Foss, L.K.1    Rieske, L.K.2
  • 24
    • 84873999065 scopus 로고    scopus 로고
    • The quantitative genetics of plumage colour in the blue tit (Parus caeruleus). PhD thesis, Imperial College.
    • Hadfield, J.D. (2005) The quantitative genetics of plumage colour in the blue tit (Parus caeruleus). PhD thesis, Imperial College.
    • (2005)
    • Hadfield, J.D.1
  • 25
    • 0034065597 scopus 로고    scopus 로고
    • Visual pigments, oil droplets, ocular media and cone photoreceptor distribution in two species of passerine bird: the blue tit (Parus caeruleus L.) and the blackbird (Turdus merula L.)
    • Hart, N.S., Partridge, J.C., Cuthill, I.C. & Bennett, A.T.D. (2000) Visual pigments, oil droplets, ocular media and cone photoreceptor distribution in two species of passerine bird: the blue tit (Parus caeruleus L.) and the blackbird (Turdus merula L.). Journal of Comparative Physiology A-Neuroethology, Sensory, Neural and Behavioral Physiology, 186, 375-387.
    • (2000) Journal of Comparative Physiology A-Neuroethology, Sensory, Neural and Behavioral Physiology , vol.186 , pp. 375-387
    • Hart, N.S.1    Partridge, J.C.2    Cuthill, I.C.3    Bennett, A.T.D.4
  • 27
    • 0025593946 scopus 로고
    • Female house finches prefer colourful males: sexual selection for a condition-dependent trait
    • Hill, G.E. (1990) Female house finches prefer colourful males: sexual selection for a condition-dependent trait. Animal Behavior, 40, 563-572.
    • (1990) Animal Behavior , vol.40 , pp. 563-572
    • Hill, G.E.1
  • 29
    • 13844250981 scopus 로고    scopus 로고
    • Carotenoid-based plumage coloration reflects hemoparasite infection and local survival in breeding great tits
    • Hõrak, P., Ots, I., Vellau, H., Spottiswoode, C. & Møller, A.P. (2001) Carotenoid-based plumage coloration reflects hemoparasite infection and local survival in breeding great tits. Oecologia, 126, 166-173.
    • (2001) Oecologia , vol.126 , pp. 166-173
    • Hõrak, P.1    Ots, I.2    Vellau, H.3    Spottiswoode, C.4    Møller, A.P.5
  • 30
    • 35448954032 scopus 로고    scopus 로고
    • Sexual, seasonal, and environmental variation in plasma carotenoids in great tits, Parus major
    • Isaksson, C., von Post, M. & Andersson, S. (2007) Sexual, seasonal, and environmental variation in plasma carotenoids in great tits, Parus major. Biological Journal of the Linnean Society, 92, 521-527.
    • (2007) Biological Journal of the Linnean Society , vol.92 , pp. 521-527
    • Isaksson, C.1    von Post, M.2    Andersson, S.3
  • 31
    • 56649106268 scopus 로고    scopus 로고
    • Sex and age differences in reflectance and biochemistry of carotenoid-based colour variation in the great tit Parus major
    • Isaksson, C., Ornborg, J., Prager, M. & Andersson, S. (2008) Sex and age differences in reflectance and biochemistry of carotenoid-based colour variation in the great tit Parus major. Biological Journal of the Linnean Society, 95, 758-765.
    • (2008) Biological Journal of the Linnean Society , vol.95 , pp. 758-765
    • Isaksson, C.1    Ornborg, J.2    Prager, M.3    Andersson, S.4
  • 32
    • 77953759254 scopus 로고    scopus 로고
    • Dissecting carotenoid from structural components of carotenoid-based coloration: a field experiment with great tits (Parus major)
    • Jacot, A., Romero-Diaz, C., Tschirren, B., Richner, H. & Fitze, P.S. (2010) Dissecting carotenoid from structural components of carotenoid-based coloration: a field experiment with great tits (Parus major). American Naturalist, 176, 55-62.
    • (2010) American Naturalist , vol.176 , pp. 55-62
    • Jacot, A.1    Romero-Diaz, C.2    Tschirren, B.3    Richner, H.4    Fitze, P.S.5
  • 34
    • 77953486354 scopus 로고    scopus 로고
    • From spectral information to animal colour vision: experiments and concepts
    • Kelber, A. & Osorio, D. (2010) From spectral information to animal colour vision: experiments and concepts. Proceedings of the Royal Society of London, Series B, 277, 1617-1625.
    • (2010) Proceedings of the Royal Society of London, Series B , vol.277 , pp. 1617-1625
    • Kelber, A.1    Osorio, D.2
  • 35
    • 0002860306 scopus 로고
    • Territory and breeding density in the great tit, Parus major L
    • Krebs, J.R. (1971) Territory and breeding density in the great tit, Parus major L. Ecology, 52, 2-22.
    • (1971) Ecology , vol.52 , pp. 2-22
    • Krebs, J.R.1
  • 36
    • 33750171851 scopus 로고    scopus 로고
    • Mechanics of carotenoid-based coloration
    • eds G.E. Hill & K.J. McGraw) - Harvard University Press, Cambridge, Massachusetts.
    • McGraw, K.J. (2006) Mechanics of carotenoid-based coloration. Bird Coloration. Volume I: Mechanisms and Measurements (eds G.E. Hill & K.J. McGraw), pp. 177-242. Harvard University Press, Cambridge, Massachusetts.
    • (2006) Bird Coloration. Volume I: Mechanisms and Measurements , pp. 177-242
    • McGraw, K.J.1
  • 37
    • 33847754189 scopus 로고    scopus 로고
    • Analyzing colors
    • eds G.E. Hill & K.J. McGraw) - Harvard University Press, Cambridge, Massachusetts.
    • Montgomerie, R. (2006) Analyzing colors. Bird Coloration. Volume I: Mechanisms and Measurements (eds G.E. Hill & K.J. McGraw), pp. 90-147. Harvard University Press, Cambridge, Massachusetts.
    • (2006) Bird Coloration. Volume I: Mechanisms and Measurements , pp. 90-147
    • Montgomerie, R.1
  • 38
    • 56749176789 scopus 로고    scopus 로고
    • Post-fledging range use of Great Tit Parus major families in relation to chick body condition
    • Naef-Daenzer, B. & Grüebler, M.U. (2008) Post-fledging range use of Great Tit Parus major families in relation to chick body condition. Ardea, 96, 181-190.
    • (2008) Ardea , vol.96 , pp. 181-190
    • Naef-Daenzer, B.1    Grüebler, M.U.2
  • 39
    • 0028975713 scopus 로고
    • Selection for the timing of great tit breeding in relation to caterpillar growth and temperature
    • van Noordwijk, A.J., McCleery, R.H. & Perrins, C.M. (1995) Selection for the timing of great tit breeding in relation to caterpillar growth and temperature. Journal of Animal Ecology, 64, 451-458.
    • (1995) Journal of Animal Ecology , vol.64 , pp. 451-458
    • van Noordwijk, A.J.1    McCleery, R.H.2    Perrins, C.M.3
  • 40
    • 43949140895 scopus 로고    scopus 로고
    • Measuring senescence in wild animal populations: towards a longitudinal approach
    • Nussey, D.H., Coulson, T., Festa-Bianchet, M. & Gaillard, M. (2008) Measuring senescence in wild animal populations: towards a longitudinal approach. Functional Ecology, 22, 393-406.
    • (2008) Functional Ecology , vol.22 , pp. 393-406
    • Nussey, D.H.1    Coulson, T.2    Festa-Bianchet, M.3    Gaillard, M.4
  • 41
    • 26244462984 scopus 로고    scopus 로고
    • Photoreceptor spectral sensitivities in terrestrial animals: adaptations for luminance and colour vision
    • Osorio, D. & Voroboyev, M. (2005) Photoreceptor spectral sensitivities in terrestrial animals: adaptations for luminance and colour vision. Proceedings of the Royal Society of London, Series B, 272, 1745-1752.
    • (2005) Proceedings of the Royal Society of London, Series B , vol.272 , pp. 1745-1752
    • Osorio, D.1    Voroboyev, M.2
  • 42
    • 79959371346 scopus 로고    scopus 로고
    • Fecundity selection on ornamental plumage colour differs between ages and sexes and varies over small spatial scales
    • Parker, T.H., Wilkin, T.A., Barr, I.R., Sheldon, B.C., Rowe, L. & Griffith, S.C. (2011) Fecundity selection on ornamental plumage colour differs between ages and sexes and varies over small spatial scales. Journal of Evolutionary Biology, 24, 1584-1597.
    • (2011) Journal of Evolutionary Biology , vol.24 , pp. 1584-1597
    • Parker, T.H.1    Wilkin, T.A.2    Barr, I.R.3    Sheldon, B.C.4    Rowe, L.5    Griffith, S.C.6
  • 44
    • 33646723279 scopus 로고    scopus 로고
    • Age-dependent traits: a new statistical model to separate within- and between individual effects
    • van de Pol, M. & Verhulst, S. (2006) Age-dependent traits: a new statistical model to separate within- and between individual effects. American Naturalist, 167, 765-773.
    • (2006) American Naturalist , vol.167 , pp. 765-773
    • van de Pol, M.1    Verhulst, S.2
  • 45
    • 59849095705 scopus 로고    scopus 로고
    • A simple method for distinguishing within- versus between-subject effects using mixed models
    • van de Pol, M. & Wright, J. (2009) A simple method for distinguishing within- versus between-subject effects using mixed models. Animal Behavior, 77, 753-758.
    • (2009) Animal Behavior , vol.77 , pp. 753-758
    • van de Pol, M.1    Wright, J.2
  • 48
    • 0027141310 scopus 로고
    • Aggregation of higher plant xanthophylls: differences in absorption spectra and in the dependency on solvent polarity
    • Ruban, A.V., Horton, P. & Young, A.J. (1993) Aggregation of higher plant xanthophylls: differences in absorption spectra and in the dependency on solvent polarity. Journal of Photochemistry and Photobiology: Biology, 21, 229-234.
    • (1993) Journal of Photochemistry and Photobiology: Biology , vol.21 , pp. 229-234
    • Ruban, A.V.1    Horton, P.2    Young, A.J.3
  • 49
    • 25444441540 scopus 로고    scopus 로고
    • Carotenoids need structural colours to shine
    • Shawkey, M.D. & Hill, G.E. (2005) Carotenoids need structural colours to shine. Biology Letters, 1, 121-124.
    • (2005) Biology Letters , vol.1 , pp. 121-124
    • Shawkey, M.D.1    Hill, G.E.2
  • 51
    • 0000659293 scopus 로고
    • Variation in plumage colour of the Great Tit Parus major in relation to habitat, season and food
    • Slagsvold, T. & Lifjeld, J.T. (1985) Variation in plumage colour of the Great Tit Parus major in relation to habitat, season and food. Journal of Zoology, 206, 321-328.
    • (1985) Journal of Zoology , vol.206 , pp. 321-328
    • Slagsvold, T.1    Lifjeld, J.T.2
  • 52
    • 43949090490 scopus 로고    scopus 로고
    • Evolution of avian plumage color in a tetrahedral color space: a phylogenetic analysis of New World buntings
    • Stoddard, M.C. & Prum, R.O. (2008) Evolution of avian plumage color in a tetrahedral color space: a phylogenetic analysis of New World buntings. American Naturalist, 171, 755-776.
    • (2008) American Naturalist , vol.171 , pp. 755-776
    • Stoddard, M.C.1    Prum, R.O.2
  • 53
    • 0348196716 scopus 로고    scopus 로고
    • Community structure of arboreal caterpillars within and among four tree species of the eastern deciduous forest
    • Summerville, K.S., Crist, T.O., Kahn, J.K. & Gering, J.C. (2003) Community structure of arboreal caterpillars within and among four tree species of the eastern deciduous forest. Ecological Entomology, 28, 747-757.
    • (2003) Ecological Entomology , vol.28 , pp. 747-757
    • Summerville, K.S.1    Crist, T.O.2    Kahn, J.K.3    Gering, J.C.4
  • 55
    • 0035970824 scopus 로고    scopus 로고
    • Energetic and fitness costs of mismatching resource supply and demand in seasonally breeding birds
    • Thomas, D.W., Blondel, J., Perret, P., Lambrechts, M.M. & Speakman, J.R. (2001) Energetic and fitness costs of mismatching resource supply and demand in seasonally breeding birds. Science, 291, 2598-2600.
    • (2001) Science , vol.291 , pp. 2598-2600
    • Thomas, D.W.1    Blondel, J.2    Perret, P.3    Lambrechts, M.M.4    Speakman, J.R.5
  • 56
    • 35148828946 scopus 로고    scopus 로고
    • Male reproductive senescence: the price of immune-induced oxidative damage on sexual attractiveness in the blue-footed booby
    • Torres, R. & Velando, A. (2007) Male reproductive senescence: the price of immune-induced oxidative damage on sexual attractiveness in the blue-footed booby. Journal of Animal Ecology, 76, 1161-1168.
    • (2007) Journal of Animal Ecology , vol.76 , pp. 1161-1168
    • Torres, R.1    Velando, A.2
  • 57
    • 78649256435 scopus 로고    scopus 로고
    • Age-related differences in a carotenoid-based coloration trait are due to within-individual changes in great tits Parus major
    • del Val, E., Quesada, J. & Senar, J.C. (2010) Age-related differences in a carotenoid-based coloration trait are due to within-individual changes in great tits Parus major. Ardea, 98, 179-184.
    • (2010) Ardea , vol.98 , pp. 179-184
    • del Val, E.1    Quesada, J.2    Senar, J.C.3
  • 58
    • 77951472070 scopus 로고    scopus 로고
    • Senescing sexual ornaments recover after a sabbatical
    • Velando, A., Drummond, H. & Torres, R. (2010) Senescing sexual ornaments recover after a sabbatical. Biology Letters, 6, 194-196.
    • (2010) Biology Letters , vol.6 , pp. 194-196
    • Velando, A.1    Drummond, H.2    Torres, R.3
  • 61
    • 63449133232 scopus 로고    scopus 로고
    • Habitat quality, nestling diet, and provisioning behaviour in great tits Parus major
    • Wilkin, T.A., King, L.E. & Sheldon, B.C. (2009) Habitat quality, nestling diet, and provisioning behaviour in great tits Parus major. Journal of Avian Biology, 40, 135-145.
    • (2009) Journal of Avian Biology , vol.40 , pp. 135-145
    • Wilkin, T.A.1    King, L.E.2    Sheldon, B.C.3
  • 62
    • 36448968250 scopus 로고    scopus 로고
    • The use of GIS in estimating spatial variation in habitat quality: a case study of lay-date in the Great Tit Parus Major
    • Wilkin, T.A., Perrins, C.M. & Sheldon, B.C. (2007) The use of GIS in estimating spatial variation in habitat quality: a case study of lay-date in the Great Tit Parus Major. Ibis, 149, 110-118.
    • (2007) Ibis , vol.149 , pp. 110-118
    • Wilkin, T.A.1    Perrins, C.M.2    Sheldon, B.C.3
  • 63
    • 0021057568 scopus 로고
    • The role of alternative host-plant species in the life of a polyphagous moth, Operophtera brumata (Lepidoptera: Geometridae)
    • Wint, W. (1983) The role of alternative host-plant species in the life of a polyphagous moth, Operophtera brumata (Lepidoptera: Geometridae). Journal of Animal Ecology, 52, 439-450.
    • (1983) Journal of Animal Ecology , vol.52 , pp. 439-450
    • Wint, W.1
  • 64
    • 0032891498 scopus 로고    scopus 로고
    • Interacting phenotypes and the evolutionary process. II. Selection resulting from social interactions
    • Wolf, J.B., Brodie, E.D. & Moore, A.J. (1999) Interacting phenotypes and the evolutionary process. II. Selection resulting from social interactions. American Naturalist, 153, 254-266.
    • (1999) American Naturalist , vol.153 , pp. 254-266
    • Wolf, J.B.1    Brodie, E.D.2    Moore, A.J.3
  • 65
    • 0028160075 scopus 로고
    • A comparison of three sampling techniques to estimate the population size of caterpillars in trees
    • Zandt, H.S. (1994) A comparison of three sampling techniques to estimate the population size of caterpillars in trees. Oecologia, 97, 399-406.
    • (1994) Oecologia , vol.97 , pp. 399-406
    • Zandt, H.S.1
  • 66
    • 0005876450 scopus 로고
    • The preparation and absorption spectra of five pure carotenoid pigments
    • Zsceile, F.P., White, J.W. & Beadle, B.W. (1942) The preparation and absorption spectra of five pure carotenoid pigments. Plant Physiology, 17, 331-346.
    • (1942) Plant Physiology , vol.17 , pp. 331-346
    • Zsceile, F.P.1    White, J.W.2    Beadle, B.W.3


* 이 정보는 Elsevier사의 SCOPUS DB에서 KISTI가 분석하여 추출한 것입니다.