Landscape moderation of biodiversity patterns and processes - eight hypotheses

Biological Reviews

Volumn 87, Issue 3, 2012, Pages 661-685

  Tscharntke, Teja ^a   Tylianakis, Jason M ^b   Rand, Tatyana A ^c   Didham, Raphael K ^b,d,e   Fahrig, Lenore ^f   Batáry, Péter ^a,g   Bengtsson, Janne ^h   Clough, Yann ^a   Crist, Thomas O ⁱ   Dormann, Carsten F ^j   Ewers, Robert M ^k   Fründ, Jochen ^a   Holt, Robert D ^l   Holzschuh, Andrea ^m   Klein, Alexandra M ⁿ   Kleijn, David ^o   Kremen, Claire ^p   Landis, Doug A ^q   Laurance, William ^r   Lindenmayer, David ^s   Scherber, Christoph ^a   Sodhi, Navjot ^t   Steffan Dewenter, Ingolf ^m   Thies, Carsten ^a   van der Putten, Wim H ^o,u   Westphal, Catrin ^a   more..

^a UNIVERSITY OF GÖTTINGEN   (Germany)

^b UNIVERSITY OF CANTERBURY   (New Zealand)

^c USDA ARS Northern Plains Agricultural Research Laboratory   (United States)

^d UNIVERSITY OF WESTERN AUSTRALIA   (Australia)

^e CSIRO   (Australia)

^f CARLETON UNIVERSITY   (Canada)

^g EÖTVÖS LORÁND UNIVERSITY   (Hungary)

^h SWEDISH UNIVERSITY OF AGRICULTURAL SCIENCES   (Sweden)

ⁱ MIAMI UNIVERSITY   (United States)

^j HELMHOLTZ CENTRE FOR ENVIRONMENTAL RESEARCH UFZ   (Germany)

^k IMPERIAL COLLEGE LONDON   (United Kingdom)

^l UNIVERSITY OF FLORIDA   (United States)

^m UNIVERSITY OF WÜRZBURG   (Germany)

ⁿ LEUPHANA UNIVERSITY OF LÜNEBURG   (Germany)

^o WAGENINGEN UNIVERSITY   (Netherlands)

^p UNIVERSITY OF CALIFORNIA   (United States)

^q MICHIGAN STATE UNIVERSITY   (United States)

^r JAMES COOK UNIVERSITY   (Australia)

^s AUSTRALIAN NATIONAL UNIVERSITY   (Australia)

^t NATIONAL UNIVERSITY OF SINGAPORE   (Singapore)

^u NETHERLANDS INSTITUTE OF ECOLOGY NIOO KNAW   (Netherlands)

more..

Author keywords

Belowground aboveground patterns; Beta diversity; Conservation management; Ecosystem functioning and services; Functional traits; Insurance hypothesis; Landscape composition and configuration; Multitrophic interactions; Resilience and stability; Spatial heterogeneity

Indexed keywords

ANIMAL; BIODIVERSITY; ECOSYSTEM; ENVIRONMENTAL MONITORING; ENVIRONMENTAL PROTECTION; METHODOLOGY; PLANT; REVIEW;

ANIMALS; BIODIVERSITY; CONSERVATION OF NATURAL RESOURCES; ECOSYSTEM; ENVIRONMENTAL MONITORING; PLANTS;

EID: 84861670005     PISSN: 14647931     EISSN: 1469185X     Source Type: Journal    
DOI: 10.1111/j.1469-185X.2011.00216.x     Document Type: Review

References (329)

1. Aarssen, L. W. & Schamp, B. S. (2002). Predicting distributions of species richness and species size in regional floras: applying the species pool hypothesis to the habitat templet model. Perspectives in Plant Ecology, Evolution and Systematics 5, 3-12.
2. Andrén, H. (1994). Effects of habitat fragmentation on birds and mammals in landscapes with different proportions of suitable habitat: a review. Oikos 71, 355-366.
3. Báldi, A. & Batáry, P. (2011). The past and future of farmland birds in Hungary. Bird Study 58, 365-377.
4. Bardgett, R. D. (2005). The Biology of Soils. A Community and Ecosystem Approach. Oxford University Press, Oxford.
5. Barlow, J., Gardner, T. A., Araujo, I. S., Avila-Pires, T. C., Bonaldo, A. B., Costa, J. E., Esposito, M. C., Ferreira, L. V., Hawes, J., Hernandez, M. I. M., Hoogmoed, M. S., Leite, R. N., Lo-Man-Hung, N. F., Malcolm, J. R., Martins, M. B., Mestre, L. A. M., Miranda-Santos, R., Nunes-Gutjahr, A. L., Overal, W. L., Parry, L., Peters, S. L., Ribeiro-Junior, M. A., da Silva, M. N. F., da Silva Motta, C. & Peres, C. A. (2007). Quantifying the biodiversity value of trcondary, and plantation forests. Proceedings of the National Academy of Sciences of the United States of America 104, 18555-18560.
6. Batáry, P. & Báldi, A. (2004). Evidence of an edge effect on avian nest success: a meta-analysis. Conservation Biology 18, 389-400.
7. Batáry, P., Báldi, A., Kleijn, D. & Tscharntke, T. (2011). Landscape-moderated biodiversity effects of agri-environment management-a meta-analysis. Proceedings of the Royal Society B 278, 1894-1902.
8. Batáry, P., Báldi, A., Sárospataki, M., Kohler, F., Verhulst, J., Knop, E., Herzog, F. & Kleijn, D. (2010a). Effect of conservation management on bees and insect-pollinated grassland plant communities in three European countries. Agriculture Ecosystems and Environment 136, 35-39.
9. Batáry, P., Matthiesen, T. & Tscharntke, T. (2010b). Landscape-moderated importance of hedges in conserving farmland bird diversity of organic vs. conventional croplands and grasslands. Biological Conservation 143, 2020-2027.
10. Batáry, P., Báldi, A., Szél, G., Podlussány, A., Rozner, I. & Erdos, S. (2007). Responses of grassland specialist and generalist beetles to management and landscape complexity. Diversity and Distributions 13, 196-202.
11. Bengtsson, J. (2010). Applied (meta)community ecology: diversity and ecosystem services at the intersection of local and regional processes. In Community Ecology (eds H. A. Verhoef and P. J. Morin), pp. 115-130. Oxford University Press, Oxford.
12. Bengtsson, J., Angelstam, P., Elmquvist, T., Emanuelsson, U., Forbes, C., Ihse, M., Moberg, F. & Nyström, M. (2003). Reserves, resilience and dynamic landscapes. Ambio 32, 389-396.
13. Bengtsson, J. & Milbrink, G. (1995). Predicting extinctions: interspecific competition, predation, and population variability in experimental Daphnia populations. Oecologia 101, 397-406.
14. Bennett, A. F. & Saunders, D. A. (2010). Habitat fragmentation and landscape change. In Conservation Biology for All (eds N. S. Sodhi and P. R. Ehrlich), pp. 88-106. Oxford University Press, Oxford.
15. Bezemer, T. M., Fountain, M. T., Barea, J. M., Christensen, S., Dekker, S. C., Duyts, H., van Hal, R., Harvey, J. A., Hedlund, K., Maraun, M., Mikola, J., Mladenov, A. G., Robin, C., De Ruiter, P. C., Scheu, S., Setälä, H., Šmilauer, P. & van der Putten, W. H. (2010). Divergent composition but similar function of soil food webs of individual plants: plant species and community effects. Ecology 91, 3027-3036.
16. Bianchi, F. J. J. A., Booij, C. J. H. & Tscharntke, T. (2006). Sustainable pest regulation in agricultural landscapes: a review on landscape composition, biodiversity and natural pest control. Proceedings of the Royal Society B 273, 1715-1727.
17. Bierregaard, R. O., Lovejoy, T. E., Kapos, V., Dossantos, A. A. & Hutchings, R. W. (1992). The biological dynamics of tropical rainforest fragments. BioScience 42, 859-866.
18. Birkhofer, K., Bezemer, T. M., Bloem, J., Bonkowski, M., Christensen, S., Ekelund, F., Fließbach, A., Hedlund, K., Mikola, J., Robin, C., Mäder, P., Setälä, H., Tatin-Froux, F., Van der Putten, W. H. & Scheu, S. (2008). Improved internal nutrient cycling and conservation biological control through long-term organic farming. Soil Biology and Biochemistry 40, 2297-2308.
19. Blitzer, E. J., Dormann, C., Holzschuh, A., Klein, A.-M., Rand, T. & Tscharntke, T. (2012). Functionally important spillover of organisms across the managed system-natural habitat interface-a review. Agriculture Ecosystems and Environment 146, 34-43.
20. Blondel, J., Thomas, D. W., Charmantier, A., Perret, P., Bourgault, P. & Lambrechts, M. M. (2006). A thirty-year study of phenotypic and genetic variation of blue tits in Mediterranean habitat mosaics. Bioscience 56, 661-673.
21. Blüthgen, N. & Klein, A. M. (2011). Functional complementarity and specialisation: the role of biodiversity in plant-pollinator interactions. Basic and Applied Ecology 12, 282-291.
22. Brose, U., Pavao-Zuckerman, M., Eklöf, A., Bengtsson, J., Berg, M. P., Cousins, S. H., Mulder, C., Verhoef, H. A. & Wolters, V. (2005). Spatial aspects of food webs. In: Dynamic Food webs (eds P.C. de Ruiter, J.C. Moore and V. Wolters), pp. 463-470. Academic Press, San Diego.
23. Brown, J. H. (1995). Macroecology. University Chicago Press, Chicago.
24. Brückmann, S. V., Krauss, J. & Steffan-Dewenter, I. (2010). Butterfly and plant specialists suffer from reduced connectivity in fragmented landscapes. Journal of Applied Ecology 47, 799-809.
25. Burel, F. (1996). Hedgerows and their role in agricultural landscapes. Critical Reviews in Plant Sciences 15, 169-190.
26. Chapron, G., Andrén, H. & Liberg, O. (2008). Conserving top predators in ecosystems. Science 320, 47.
27. Chase, J. M. & Bengtsson, J. (2010). Increasing spatio-temporal scales: metacommunity ecology. In Community Ecology (eds H. A. Verhoef and P. J. Morin), pp. 57-68. Oxford University Press, Oxford.
28. Chesson, P., Donahue, M. J., Melbourne, B. A. & Sears, A. L. W. (2005). Scale transition theory for understanding mechanisms in metacommunities. In Metacommunities: Spatial Dynamics and Ecological Communities (eds M. Holyoak, M. A. Leibold and R. D. Holt), pp. 279-306. University of Chicago Press, Chicago.
29. Clough, Y., Barkmann, J., Juhrbandt, J., Kessler, M., Wanger, T. C., Anshary, A., Buchori, D., Cicuzza, D., Darras, D., Dwi Putra, D., Erasmi, S., Pitopang, R., Schmidt, C., Schulze, C. H., Seidel, D., Steffan-Dewenter, I., Stenchly, K., Vidal, S., Weist, M., Wielgoss, A. C. & Tscharntke, T. (2011). Combining high biodiversity with high yields in tropical agroforests. Proceedings of the National Academy of Sciences of the United States of America 108, 8311-8316.
30. Clough, Y., Holzschuh, A., Gabriel, D., Purtauf, T., Kruess, A., Steffan-Dewenter, I., Kleijn, D. & Tscharntke, T. (2007). Alpha and beta diversity of arthropods and plants in organically and conventionally managed wheat fields. Journal of Applied Ecology 44, 804-812.
31. Clough, Y., Kruess, A., Kleijn, D. & Tscharntke, T. (2005). Spider diversity in cereal fields: comparing factors at local, landscape and regional scales. Journal of Biogeography 32, 2007-2014.
32. Cochrane, M. A. & Laurance, W. F. (2002). Fire as a large-scale edge effect in Amazonian forests. Journal of Tropical Ecology 18, 311-325.
33. Coleman, B. D., Mares, M. A., Willig, M. R. & Hsieh, Y. H. (1982). Randomness, area, and species richness. Ecology 63, 1121-1133.
34. Collinge, S. K. (2009). Ecology of Fragmented Landscapes. The John Hopkins University Press, Baltimore.
35. Collinge, S. K. & Forman, R. T. T. (1998). A conceptual model of land conversion processes: predictions and evidence from a microlandscape experiment with grassland insects. Oikos 82, 66-84.
36. Concepción, E. D. & Díaz, M. (2011). Field, landscape and regional effects of farmland management on specialist open-land birds: does body size matter? Agriculture Ecosystems and Environment 142, 303-310.
37. Concepción, E. D., Díaz, M. & Baquero, R. A. (2008). Effects of landscape complexity on the ecological effectiveness of agri-environment schemes. Landscape Ecology 23, 135-148.
38. Cook, W. M., Anderson, R. M. & Schweiger, E. W. (2004). Is the matrix really inhospitable? Vole runway distribution in a fragmented landscape. Oikos 104, 5-14.
39. Cook, W. M., Yao, J., Foster, B. L., Holt, R. D. & Patrick, L. B. (2005). Secondary succession in an experimentally fragmented landscape: community patterns across space and time. Ecology 86, 1267-1279.
40. Cornell, H. V. & Lawton, J. H. (1992). Species interactions, local and regional processes, and limits to the richness of ecological communities: a theoretical perspective. Journal of Animal Ecology 61, 1-12.
41. Crawley, M. J. (1997). Biodiversity. In Plant Ecology. Second Edition (ed. M. J. Crawley), pp. 595-632. Blackwell, Oxford.
42. Cronin, J. T. (2007). From population sources to sieves: the matrix alters host-parasitoid source-sink structure. Ecology 88, 2966-2976.
43. Crowder, D. W., Northfield, T. D., Strand, M. R. & Snyder, W. E. (2010). Organic agriculture promotes evenness and natural pest control. Nature 466, 109-112.
44. Daily, G. C., Ehrlich, P. R. & Sanchez-Azofeifa, G. A. (2001). Countryside biogeography: use of human-dominated habitats by the avifauna of southern Costa Rica. Ecological Applications 11, 1-13.
45. Dallimer, M., Gaston, K. J., Skinner, A. M. J., Hanley, N., Acs, A. & Armsworth, P. R. (2010). Field-level bird abundances are enhanced by landscape-scale agri-environment scheme uptake. Biology Letters 6, 643-646.
46. Damschen, E. I., Haddad, N. M., Orrock, J. L., Tewksbury, J. J. & Levey, D. J. (2006). Corridors increase plant species richness at large scales. Science 313, 1284-1286.
47. Debinski, D. M. & Holt, R. D. (2000). A survey and overview of habitat fragmentation experiments. Conservation Biology 14, 342-355.
48. De Deyn, G. B. & Van der Putten, W. H. (2005). Linking above- and belowground biodiversity. Trends in Ecology and Evolution 20, 625-633.
49. Didham, R. K. (2011). Life after logging: strategic withdrawal from the garden of Eden or tactical error for wilderness conservation? Biotropica 43, 393-395.
50. Didham, R. K., Ghazoul, J., Stork, N. E. & Davis, A. J. (1996). Insects in fragmented forests: a functional approach. Trends in Ecology and Evolution 11, 255-260.
51. Didham, R. K., Hammond, P. M., Lawton, J. H., Eggleton, P. & Stork, N. E. (1998). Beetle species responses to tropical forest fragmentation. Ecological Monographs 68, 295-323.
52. Dormann, C. F., Schweiger, O., Augenstein, I., Bailey, D., Billeter, R., De Blust, G., Bugter, R., Frenzel, M., Hendrickx, F., Herzog, F., Klotz, S., Liira, J., Maelfait, J.-P., Schmidt, T., Speelmans, M., van Wingerden, W. K. R. E. & Zobel, M. (2007). Effects of land use on similarity of plant and animal communities. Global Ecology and Biogeography 16, 774-787.
53. Duelli, P. & Obrist, M. K. (2003). Regional biodiversity in an agricultural landscape: the contribution of seminatural habitat islands. Basic and Applied Ecology 4, 129-138.
54. Dunning, J. B., Danielson, B. J. & Pulliam, H. R. (1992). Ecological processes that affect populations in complex landscapes. Oikos 65, 169-175.
55. Ebeling, A., Klein, A.-M., Schumacher, J., Weisser, W. W. & Tscharntke, T. (2008). How does plant richness affect pollinator richness and temporal stability of flower visits? Oikos 117, 1808-1815.
56. Ebeling, A., Klein, A.-M. & Tscharntke, T. (2011). Plant-flower visitor interaction webs: temporal stability and pollinator specialization increases along an experimental plant diversity gradient. Basic and Applied Ecology 12, 300-309.
57. Eckert, C. G., Samis, K. E. & Lougheed, S. C. (2008). Genetic variation across species' geographical ranges: the central-marginal hypothesis and beyond. Molecular Ecology 17, 1170-1188.
58. Eilers, E. J. & Klein, A.-M. (2009). Landscape context and management effects on an important insect pest and its natural enemies in almond. Biological Control 51, 388-394.
59. Elmqvist, T., Folke, C., Nyström, M., Peterson, G., Bengtsson, J., Walker, B. & Norberg, J. (2003). Response diversity, ecosystem change, and resilience. Frontiers in Ecology and the Environment 1, 488-494.
60. Ethier, K. & Fahrig, L. (2011). Positive effects of forest fragmentation, independent of forest amount, on bat abundance in eastern Ontario, Canada. Landscape Ecology 26, 865-876.
61. Ettema, E. H. & Wardle, D. A. (2002). Spatial soil ecology. Trends in Ecology & Evolution 17, 177-183.
62. Ewers, R. M. & Didham, R. K. (2006). Confounding factors in the detection of species responses to habitat fragmentation. Biological Reviews 81, 117-142.
63. Ewers, R. M. & Didham, R. K. (2007). Habitat fragmentation: panchreston or paradigm? Trends in Ecology and Evolution 22, 511.
64. Ewers, R. M. & Didham, R. K. (2008). Pervasive impact of large-scale edge effects on a beetle community. Proceedings of the National Academy of Sciences 105, 5426-5429.
65. Ewers, R. M., Kapos, V., Coomes, D. A., Lafortezza, R. & Didham, R. K. (2009). Mapping community change in modified landscapes. Biological Conservation 142, 2872-2880.
66. Fagan, W. E., Cantrell, R. S. & Cosner, C. (1999). How habitat edges change species interactions. The American Naturalist 153, 165-182.
67. Fahrig, L. (2003). Effects of habitat fragmentation on biodiversity. Annual Reviews of Ecology and Systematics 34, 487-515.
68. Fahrig, L. (2007). Non-optimal animal movement in human-altered landscapes. Functional Ecology 21, 1003-1015.
69. Fahrig, L., Baudry, J., Brotons, L., Burel, F. G., Crist, T. O., Fuller, R. J., Sirami, C., Siriwardena, G. M. & Martin, J. L. (2011). Functional landscape heterogeneity and animal biodiversity in agricultural landscapes. Ecology Letters 14, 101-112.
70. Fischer, J. & Lindenmayer, D. B. (2005). Perfectly nested or significantly nested-an important difference for conservation management. Oikos 109, 485-494.
71. Fischer, J., Stott, J. & Law, B. S. (2010). The disproportionate value of scattered trees. Biological Conservation 143, 1564-1567.
72. Flohre, A., Fischer, C., Aavik, T., Bengtsson, J., Berendse, F., Bommarco, R., Ceryngier, P., Clement, L. W., Dennis, C., Eggers, S., Emmerson, M., Geiger, F., Guerrero, I., Hawro, V., Inchausti, P., Liira, J., Morales, M. B., Onate, J. J., Pärt, T., Weisser, W. W., Winqvist, C., Thies, C. & Tscharntke, T. (2011). Agricultural intensification and biodiversity partitioning in European landscapes comparing plants, carabids, and birds. Ecological Applications 21, 1772-1781.
73. Flohre, A., Rudnick, M., Traser, G., Tscharntke, T. & Eggers, T. (2011). Does soil biota benefit from organic farming in complex vs. simple landscapes? Agriculture Ecosystems and Environment 141, 210-214.
74. Forman, R. T. T. (1995). Some general principles of landscape and regional ecology. Landscape Ecology 10, 133-142.
75. Fortin, M. J. & Dale, M. R. T. (2005). Spatial Analysis: A Guide for Ecologists. Cambridge University Press, Cambridge, UK.
76. Foster, W. A., Snaddon, J. L., Turner, E. C., Fayle, T. M., Cockerill, T. D., Ellwood, M. D. F., Broad, G. R., Chung, A. Y. C., Eggleton, P., Ken, C. V. & Yusah, K. M. (2011). Establishing the evidence base for maintaining biodiversity and ecosystem function in the oilpalm landscape of South East Asia. Philosophical Transactions of the Royal Society B 366, 3277-3291.
77. Gabriel, D., Roschewitz, I., Tscharntke, T. & Thies, C. (2006). Beta diversity at different spatial scales: plant communities in organic and conventional agriculture. Ecological Applications 16, 2011-2021.
78. Gabriel, D., Sait, S. M., Hodgson, J. A., Schmutz, U., Kunin, W. E. & Benton, T. G. (2010). Scale matters: the impact of organic farming on biodiversity at different spatial scales. Ecology Letters 13, 858-869.
79. Gabriel, D., Thies, C. & Tscharntke, T. (2005). Local diversity of arable weeds increases with landscape complexity. Perspectives in Plant Ecology, Evolution and Systematics 7, 85-93.
80. Gagic, V., Tscharntke, T., Dormann, C. F., Gruber, B., Wilstermann, A. & Thies, C. (2011). Food web structure and biocontrol in a four-trophic level system across a landscape complexity gradient. Proceedings of the Royal Society B 278, 2946-2953.
81. Gardiner, M. M., Landis, D. A., Gratton, C., DiFonzo, C. D., O'Neal, M., Chacon, J. M., Wayo, M. T., Schmidt, N. P., Mueller, E. E. & Heimpel, G. E. (2009a). Landscape diversity enhances the biological control of an introduced crop pest in the north-central U.S. Ecological Applications 19, 143-154.
82. Gardiner, M. M., Landis, D. A., Gratton, C., Schmidt, N., O'Neal, M., Mueller, E., Chacon, J., Heimpel, G. E. & Difonzo, C. D. (2009b). Landscape composition mediates exotic and native coccinellid community structure. Diversity and Distributions 15, 554-564.
83. Gascon, C., Lovejoy, T. E., Bierregaard, R. O. Jr, Malcolm, J. R., Stouffer, P. C., Vasconcelos, H. L., Laurance, W. F., Zimmerman, B., Tocher, M. & Borges, S. (1999). Matrix habitat and species richness in tropical forest remnants. Biological Conservation 91, 223-229.
84. Gaston, K. (2000). Global patterns in biodiversity. Nature 405, 220-227.
85. Gaston, K. J. & Fuller, R. A. (2008). Commonness, population depletion and conservation biology. Trends in Ecology and Evolution 23, 14-19.
86. Geiger, F., De Snoo, G. R., Berendse, F., Guerrero, I., Morales, M. B., Oñate, J. J., Eggers, S., Pärt, T., Bommarco, R., Bengtsson, J., Clement, L. W., Weisser, W. W., Olszewski, A., Ceryngier, P., Hawro, V., Inchausti, P., Fischer, C., Flohre, A., Thies, C. & Tscharntke, T. (2010). Landscape composition influences farm management effects on farmland birds in winter: a pan-European approach. Agriculture Ecosystems & Environment 139, 571-577.
87. Gering, J. C. & Crist, T. O. (2002). The alpha-beta-regional relationship: providing new insights into local-regional patterns of species richness and scale dependence of diversity components. Ecology Letters 5, 433-444.
88. Gering, J. C., Crist, T. O. & Veech, J. A. (2003). Additive partitioning of beetle species diversity across multiple spatial scales: implications for regional conservation of biodiversity. Conservation Biology 17, 488-499.
89. Gilroy, J. J. & Sutherland, W. J. (2007). Beyond ecological traps: perceptual errors and undervalued resources. Trends in Ecology and Evolution 22, 351-356.
90. Gladbach, D. J., Holzschuh, A., Scherber, C., Thies, C., Dormann, C. F. & Tscharntke, T. (2011). Crop-noncrop spillover: arable fields affect trophic interactions on wild plants in surrounding habitats. Oecologia 166, 433-441.
91. Goodwin, B. J. & Fahrig, L. (2002). How does landscape structure influence landscape connectivity? Oikos 99, 552-570.
92. Greenleaf, S. S. & Kremen, C. (2006). Wild bees enhance honey bees' pollination of hybrid sunflower. Proceedings of the National Academy of Sciences of the United States of America 103, 13890-13895.
93. Grez, A., Zaviezo, T., Tischendorf, L. & Fahrig, L. (2004). A transient, positive effect of habitat fragmentation on insect population densities. Oecologia 141, 444-451.
94. Griffin, J. N., O'Gorman, E. J., Emmerson, M. C., Jenkins, S. R., Klein, A.-M., Loreau, M. & Symstad, A. (2009). Biodiversity and the stability of ecosystem functioning. In Biodiversity, Ecosystem Functioning, and Human Wellbeing-An Ecological and Economic Perspective (eds S. Naeem, D. E. Bunker, A. Hector, M. Loreau and C. Perrings), pp. 78-93. Oxford University Press, Oxford.
95. Griffiths, B. S., Ritz, K., Wheatley, R., Kuan, H. L., Boag, B., Christensen, S., Ekelund, F., Sorensen, S. J., Muller, S. & Bloem, J. (2001). An examination of the biodiversity-ecosystem function relationship in arable soil microbial communities. Soil Biology and Biochemistry 33, 1713-1722.
96. Gurr, G. M., Wratten, S. D. & Altieri, M. A. (2004). Ecological Engineering for Pest Management. CSIRO Publishing, Collingwood.
97. Haenke, S., Scheid, B., Schaefer, M., Tscharntke, T. & Thies, C. (2009). Increasing syrphid fly diversity and density in sown flower strips within simple vs. complex landscapes. Journal of Applied Ecology 46, 1106-1114.
98. Haila, Y. (2002). A conceptual genealogy of fragmentation research: from island biogeography to landscape ecology. Ecological Applications 12, 321-334.
99. Hambäck, P. A., Summerville, K. S., Steffan-Dewenter, I., Krauss, J., Englund, G. & Crist, T. O. (2007). Habitat specialization, body size, and family identity explain lepidopteran density-area relationships in a cross-continental comparison. Proceedings of the National Academy of Sciences 104, 8368-8373.
100. Hanski, I. (2011). Habitat loss, the dynamics of biodiversity, and a perspective on conservation. Ambio 40, 248-255.
101. Harrison, S. & Bruna, E. (1999). Habitat fragmentation and large-scale conservation: what do we know for sure? Ecography 22, 225-232.
102. Harrison, S. & Cornell, H. (2008). Toward a better understanding of the regional causes of local community richness. Ecology Letters 11, 969-979.
103. Harrison, S., Ross, S. J. & Lawton, J. H. (1992). Beta diversity on geographic gradients in Britain. Journal of Animal Ecology 61, 151-158.
104. He, F. & Hubbell, S. P. (2011). Species-area relationships always overestimate extinction rates from habitat loss. Nature 473, 368-371.
105. Hedlund, K., Griffiths, B., Christensen, S., Scheu, S., Setälä, H., Tscharntke, T. & Verhoef, H. (2004). Trophic interactions in changing landscapes: responses of soil food webs. Basic and Applied Ecology 5, 495-503.
106. Helgason, T., Daniell, T. J., Husband, R., Fitter, A. H. & Young, J. P. W. (1998). Ploughing up the world-wide web? Nature 394, 431.
107. Hendrickx, F., Maelfait, J., van Wingerden, W., Schweiger, O., Speelmans, M., Aviron, S., Augenstein, I., Billeter, R., Bailey, D., Bukacek, R., Burel, F., Diekötter, T., Dirksen, J., Herzog, F., Liira, J., Roubalova, M., Vandomme, V. & Bugster, R. (2007). How landscape structure, land-use intensity and habitat diversity affect components of total arthropod diversity in agricultural landscapes. Journal of Applied Ecology 44, 340-351.
108. Henle, K., Davies, K. F., Kleyer, M., Margules, C. & Settele, J. (2004). Predictors of species sensitivity to fragmentation. Biodiversity and Conservation 13, 207-251.
109. Herrmann, F., Westphal, C., Moritz, R. F. A. & Steffan-Dewenter, I. (2007). Genetic diversity and mass resources promote colony size and forager densities of a social bee (Bombus pascuorum) in agricultural landscapes. Molecular Ecology 16, 1167-1178.
110. Holland, J. D., Fahrig, L. & Cappuccino, N. (2005). Fecundity determines the extinction threshold in a Canadian assemblage of longhorned beetles (Coleoptera: Cerambycidae). Journal of Insect Conservation 9, 109-119.
111. Holt, R. D. (1996). Food webs in space: an island biogeographic perspective. In Food Webs (eds G. A. Polis and K. O. Winemiller), pp. 313-323. Chapman & Hall, New York.
112. Holt, R. D. (2008). Theoretical perspectives on resource pulses. Ecology 89, 671-681.
113. Holt, R. D. (2010). Toward a trophic island biogeography: reflections on the interface of island biogeography and food web ecology. In The Theory of Island Biogeography (eds J. B. Losos and R. E. Ricklefs), pp. 143-185. Princeton University Press, Princeton, USA.
114. Holt, R. D., Barfield, M. & Gomulkiewicz, R. (2005). Theories of niche conservatism and evolution: could exotic species be potential tests? In Species Invasions: Insights into Ecology, Evolution, and Biogeography (eds D. Sax, J. Stachowicz and D. Gaines), pp. 259-290. Sinauer Associates, Sunderland.
115. Holt, R. D. & Hochberg, M. (2001). Evaluating indirect ecological effects of biological control. In Indirect Interactions, Community Modules, and Biological Control: A Theoretical Perspective (eds E. Wajnberg, J. K. Scott and P. C. Quimby), pp. 13-37. CABI Publishing, Wallingford.
116. Holt, R. D., Lawton, J. H., Polis, G. A. & Martinez, N. D. (1999). Trophic rank and the species-area relationship. Ecology 80, 1495-1504.
117. Holzschuh, A., Dormann, C. F., Tscharntke, T. & Steffan-Dewenter, I. (2011). Expansion of mass-flowering crops leads to transient pollinator dilution and reduced wild plant pollination. Proceedings of the Royal Society B 278, 3444-3451.
118. Holzschuh, A., Steffan-Dewenter, I., Kleijn, D. & Tscharntke, T. (2007). Diversity of flower-visiting bees in cereal fields: effects of farming system, landscape composition and regional context. Journal of Applied Ecology 44, 41-49.
119. Holzschuh, A., Steffan-Dewenter, I. & Tscharntke, T. (2008). Agricultural landscapes with organic crops support higher pollinator diversity. Oikos 117, 354-361.
120. Holzschuh, A., Steffan-Dewenter, I. & Tscharntke, T. (2009). Grass strip corridors in agricultural landscapes enhance nest-site colonization by solitary wasps. Ecological Applications 19, 123-132.
121. Holzschuh, A., Steffan-Dewenter, I. & Tscharntke, T. (2010). How do landscape composition and configuration, organic farming and fallow strips affect the diversity of bees, wasps and their parasitoids? Journal of Animal Ecology 79, 491-500.
122. Hooper, D. U., Chapin, F. S. III, Ewel, J. J., Hector, A., Inchausti, P., Lavorel, S., Lawton, J. H., Lodge, D., Loreau, M., Naeem, S., Schmid, B., Setälä, H., Symstad, A. J., Vandermeer, J. & Wardle, D. A. (2005). Effects of biodiversity on ecosystem functioning: a consensus of current knowledge. Ecological Monographs 75, 3-35.
123. Hubbell, S. P. (2001). The Unified Theory of Biodiversity and Biogeography. Princeton University Press, Princeton.
124. Hughes, A. R. & Stachowicz, J. J. (2004). Genetic diversity enhances the resistance of a seagrass ecosystem to disturbance. Proceedings of the National Academy of Sciences 101, 8998-9002.
125. Isaacs, R., Tuell, J., Fiedler, A., Gardiner, M. & Landis, D. (2009). Maximizing arthropod-mediated ecosystem services in agricultural landscapes: the role of native plants. Frontiers in Ecolology and the Environment 7, 196-203.
126. Janzen, D. H. (1983). No park is an island: increased interference from outside as park size decreases. Oikos 41, 402-410.
127. Janzen, D. H. (1986). The eternal external threat. In Conservation Biology: The Science of Scarcity and Diversity (ed. M. E. Soule), pp. 286-303. Sinauer, Sunderland.
128. Jha, S. & Dick, C. W. (2010). Native bees mediate long-distance pollen dispersal in a shade coffee landscape mosaic. Proceedings of the National Academy of Sciences 107, 13760-13764.
129. Kardol, P., Cregger, M. A., Campany, C. E. & Classen, A. T. (2010). Soil ecosystem functioning under climate change: plant species and community effects. Ecology 91, 767-981.
130. Kardol, P., Newton, J. S., Bezemer, T. M., Maraun, M. & van der Putten, W. H. (2009). Contrasting diversity patterns of soil mites and nematodes in secondary succession. Acta Oecologia 35, 603-609.
131. Kareiva, P. (1990). Population dynamics in spatially complex environments: theory and data. Philosophical Transactions of the Royal Society London B 330, 175-190.
132. Kareiva, P. & Wennergren, U. (1995). Connecting landscape patterns to ecosystem and population processes. Nature 373, 299-302.
133. Kawecki, T. J. (2008). Adaptation to marginal habitats. Annual Review of Ecology, Evolution and Systematics 39, 321-342.
134. Kessler, M., Abrahamczyk, S., Bos, M., Buchori, D., Putra, D. D., Gradstein, S. R., Höhn, P., Kluge, J., Orend, F., Pitopang, R., Shahabuddin, S., Schulze, C. H., Sporn, S. G., Steffan-Dewenter, I., Tjitrosoedirdjo, S. S. & Tscharntke, T. (2009). Alpha and beta diversity of plants and animals along a tropical land-use gradient. Ecological Applications 19, 2142-2156.
135. Kier, G., Kreft, H., Lee, T. M., Jetz, W., Ibisch, P. I., Nowicki, C., Mutke, J. & Barthlott, W. (2009). A global assessment of endemism and species richness across island and mainland regions. Proceedings of the National Academy of Sciences 106, 9322-9327.
136. Kleijn, D. (1996). The use of nutrient resources from arable fields by plants in field boundaries. Journal of Applied Ecology 33, 1433-1440.
137. Kleijn, D., Baquero, R. A., Clough, Y., Díaz, M., De Esteban, J., Fernández, F., Gabriel, D., Herzog, F., Holzschuh A., Jöhl, R., Knop, E., Kruess, A., Marshall, E. J. P., Steffan-Dewenter, I., Tscharntke, T., Verhulst, J, West, T. M. & Yela, J. L. (2006). Mixed biodiversity benefits of agri-environment schemes in five European countries. Ecology Letters 9, 243-254.
138. Kleijn, D., Berendse, F., Smit, R. & Gilissen, N. (2001). Agri-environment schemes do not effectively protect biodiversity in Dutch agricultural landscapes. Nature 413, 723-725.
139. Kleijn, D., Rundlöf, M., Scheper, J., Smith, H. G. & Tscharntke, T. (2011) Does conservation on farmland contribute to halt biodiversity decline? Trends in Ecology and Evolution 26, 474-481.
140. Klein, A.-M., Cunningham, S. A., Bos, M. & Steffan-Dewenter, I. (2008). Advances in pollination ecology from tropical plantation crops. Ecology 89, 935-943.
141. Klein, A.-M., Steffan-Dewenter, I. & Tscharntke, T. (2003a). Fruit set of highland coffee depends on the diversity of pollinating bees. Proceedings of the Royal Society B 270, 955-961.
142. Klein, A.-M., Steffan-Dewenter, I. & Tscharntke, T. (2003b). Flower visitation and fruit set of Coffea canephora in relation to local and regional agroforestry management. Journal of Applied Ecolology 40, 837-845.
143. Klein, A.-M., Vaissière, B. E., Cane, J. H., Steffan-Dewenter, I., Cunningham, S. A., Kremen, C. & Tscharntke, T. (2007). Importance of pollinators in changing landscapes for world crops. Proceedings of the Royal Society B 274, 303-313.
144. Knight, T. M., McCoy, M. W., Chase, J. M., McCoy, K. A. & Holt, R. D. (2005). Trophic cascades across ecosystems. Nature 437, 880-884.
145. Krauss, J., Klein, A. M., Steffan-Dewenter, I. & Tscharntke, T. (2004). Effects of habitat area, isolation, and landscape diversity on plant species richness of calcareous grasslands. Biodiversity and Conservation 13, 1427-1439.
146. Krauss, J., Steffan-Dewenter, I. & Tscharntke, T. (2003a). How does landscape context contribute to effects of habitat fragmentation on diversity and population density of butterflies? Journal of Biogeography 30, 889-900.
147. Krauss, J., Steffan-Dewenter, I. & Tscharntke, T. (2003b). Local species immigration, extinction, and turnover of butterflies in relation to habitat area and habitat isolation. Oecologia 137, 591-602.
148. Kremen, C. (2005). Managing ecosystem services: what do we need to know about their ecology? Ecology Letters 8, 468-479.
149. Kremen, C., Cameron, A., Moilanen, A., Phillips, S. J., Thomas, C. D., Beentje, H., Dransfield, J., Fisher, B. L., Glaw, F., Good, T. C., Harper, G. J., Hijmans, R. J., Lees, D. C., Louis, E., Nussbaum, R. A., Raxworthy, C. J., Razafimpahanana, A., Schatz, G. E., Vences, M., Vieites, D. R., Wright, P. C. & Zjhra, M. L. (2008). Aligning conservation priorities across taxa in Madagascar with high-resolution planning tools. Science 320, 222-226.
150. Kremen, C., Williams, N. M., Aizen, M. A., Gemmill-Herren, B., LeBuhn, G., Minckley, R., Packer, L., Potts, S. G., Roulston, T., Steffan-Dewenter, I., Vazquez, D. P., Winfree, R., Adams, L., Crone, E. E., Greenleaf, S. S., Keitt, T. H., Klein, A.-M., Regetz, J. & Ricketts, T. H. (2007). Pollination and other ecosystem services produced by mobile organisms: a conceptual framework for the effects of land-use change. Ecology Letters 10, 299-314.
151. Kremen, C., Williams, N. M., Bugg, R. L., Fay, J. P. & Thorp, R. W. (2004). The area requirements of an ecosystem service: crop pollination by native bee communities in California. Ecology Letters 7, 1109-1119.
152. Kremen, C., Williams, N. M. & Thorp, R. W. (2002). Crop pollination from native bees at risk from agricultural intensification. Proceedings of the National Academy of Sciences 99, 16812-16816.
153. Kruess, A. & Tscharntke, T. (1994). Habitat fragmentation, species loss, and biological control. Science 264, 1581-1584.
154. Kruess, A. & Tscharntke, T. (2000). Species richness and parasitism in a fragmented landscape: experiments and field studies with insects on Vicia sepium. Oecologia 122, 129-137.
155. Kupfer, J. A., Malanson, G. P. & Franklin, S. B. (2006). Not seeing the ocean for the islands: the mediating influence of matrix-based processes on forest fragmentation effects. Global Ecology and Biogeography 15, 8-20.
156. Kuussaari, M., Bommarco, R., Heikkinen, R. K., Helm, A., Krauss, J., Lindborg, R., Öckinger, E., Pärtel, M., Pino, J., Rodà, F., Stefanescu, C., Teder, T., Zobel, M. & Steffan-Dewenter, I. (2009). Extinction debt: a challenge for biodiversity conservation. Trends in Ecology and Evolution 24, 564-571.
157. Laliberté, E. & Tylianakis, J. M. (2010). Deforestation homogenizes tropical parasitoid-host networks. Ecology 91, 1740-1747.
158. Laliberté, E., Wells, J. A., DeClerck, F., Metcalfe, D. J., Catterall, C. P., Queiroz, C., Aubin, I., Bonser, S. P., Ding, Y., Fraterrigo, J. M., McNamara, S., Morgan, J. W., Merlos, D. S., Vesk, P. A. & Mayfield, M. M. (2010). Land-use intensification reduces functional redundancy and response diversity in plant communities. Ecology Letters 13, 76-86.
159. Lamb, E. G. & Cahill, J. F. (2008). When competition does not matter: grassland diversity and community composition. American Naturalist 171, 777-787.
160. Landis, D. A., Gardiner, M. M., van der Werf, W. & Swinton, S. M. (2008). Increasing corn for biofuel production reduces biocontrol services in agricultural landscapes. Proceedings of the National Academy of Sciences 105, 20552-20557.
161. Landis, D. A. & Marino, P. C. (1999). Landscape structure and extra-field processes: impact on management of pests and beneficials. In Handbook of Pest Management (ed. J. R. Ruberson), pp. 79-104. Marcel Dekker Inc., New York.
162. Landis, D. A., Wratten, S. D. & Gurr, G. M. (2000). Habitat management to conserve natural enemies of arthropod pests in agriculture. Annual Review of Entomology 45, 175-201.
163. Larrivée, M., Fahrig, L. & Drapeau, P. (2008). Edge effects created by wildfire and clear-cutting on boreal forest ground-dwelling spiders. Forest Ecology and Management 255, 1434-1445.
164. Larsen, T. H., Williams, N. M. & Kremen, C. (2005). Extinction order and altered community structure rapidly disrupt ecosystem functioning. Ecology Letters 8, 538-547.
165. Laurance, W. F. (1991). Ecological correlates of extinction proneness in Australian tropical rainforest mammals. Conservation Biology 5, 79-89.
166. Laurance, W. F. (2002). Hyperdynamism in fragmented habitats. Journal of Vegetation Science 13, 595-602.
167. Laurance, W. F. (2008). Theory meets reality: how habitat fragmentation research has transcended island biogeographic theory. Biological Conservation 141, 1731-1744.
168. Laurance, W. F., Camargo, J. L. C., Luizão, R. C. C., Laurance, S. G., Pimm, S. L., Bruna, E. M., Stouffer, P. C., Williamson, G. B., Benítez-Malvido, J. & Vasconcelos, H. L. (2011). The fate of Amazonian forest fragments: a 32-year investigation. Biological Conservation 144, 56-67.
169. Laurance, W. F., Delamonica, P., Laurance, S. G., Vasconcelos, H. L. & Lovejoy, T. E. (2000). Conservation-rainforest fragmentation kills big trees. Nature 404, 836.
170. Laurance, W. F., Nascimento, H. E. M., Laurance, S. G., Andrade, A., Ewers, R., Harms, K. E., Luizão, R. C. C. & Ribeiro, J. E. (2007). Habitat fragmentation, variable edge effects, and the landscape-divergence hypothesis. PLoS One 2, e1017.
171. Laurance, W. F. & Yensen, E. (1991). Predicting the impacts of edge effects in fragmented habitats. Biological Conservation 55, 77-92.
172. Lavandero, B. I., Wratten, S. D., Didham, R. K. & Gurr, G. (2006). Increasing floral diversity for selective enhancement of biological control agents: a double-edged sward? Basic and Applied Ecology 7, 236-243.
173. Lavorel, S. & Garnier, E. (2002). Predicting changes in community composition and ecosystem functioning from plant traits: revisiting the Holy Grail. Functional Ecology 16, 545-556.
174. Lawton, J. H. (1995). Population dynamic principles. In Extinction Risks (eds J. H. Lawton and R. M. May), pp. 147-163. Oxford University Press, Oxford.
175. Lawton, J. H. (1999). Are there general laws in ecology? Oikos 84, 177-192.
176. Lawton, J. H. & Brown, V. K. (1994). Redundancy in ecosystems. In Biodiversity and Ecosystem Function (eds E. D. Schulze and H. A. Mooney), pp. 255-270. Springer, Berlin.
177. Lee, J. C., Menalled, F. D. & Landis, D. A. (2001). Refuge habitats modify impact of insecticide disturbance on carabid beetle communities. Journal of Applied Ecology 38, 472-483.
178. Lee, J. S. H., Lee, I. Q. W., Lim, S. L.-H., Huijbregts, J. & Sodhi, N. S. (2009). Changes in dung beetle communities along a gradient of tropical forest disturbance in South-east Asia. Journal of Tropical Ecology 25, 677-680.
179. Lehtinen, R. M., Ramanamanjato, J.-B. & Raveloarison, J. G. (2003). Edge effects and extinction proneness in a herpetofauna from Madagascar. Biodiversity and Conservation 12, 1357-1370.
180. Leibold, M. A., Holyoak, M., Mouquet, N., Amarasekare, P., Chase, J. M., Hoopes, M. F., Holt, R. D., Shurin, J. B., Law, R., Tilman, D., Loreau, M. & Gonzalez, A. (2004). The metacommunity concept: a framework for multi-scale community ecology. Ecology Letters 7, 601-613.
181. Leimu, R. & Fischer, M. (2008). A meta-analysis of local adaptation in plants. PLoS One 3, e4010.1-8.
182. Lembo, T., Hampson, K., Haydon, D. T., Craft, M., Dobson, A., Dushoff, J., Ernest, E., Hoare, R., Kaare, M., Mlengeya, T., Mentzel, C. & Cleaveland, S. (2008). Exploring reservoir dynamics: a case study of rabies in the Serengeti ecosystem. Journal of Applied Ecology 45, 1246-1257.
183. Leps, J. (2004). What do the biodiversity experiments tell us about consequences of plant species loss in the real world? Basic and Applied Ecology 5, 529-534.
184. Levin, S. A. (1992). The problem of pattern and scale in ecology. Ecology 73, 1943-1967.
185. Levin, S. A. (2000). Multiple scales and the maintenance of biodiversity. Ecosystems 3, 498-506.
186. Levins, R. (1969). Some genetic and demographic consequences of environmental heterogeneity for biological control. Bulletin of the Entomological Society of America 15, 237-240.
187. Lindberg, N. & Bengtsson, J. (2006). Recovery of forest soil fauna diversity and composition after repeated summer droughts. Oikos 114, 496-506.
188. Lindenmayer, D. B., Cunnungham, S., MacGregor, C., Crane, M., Michael, D., Fischer, J., Montague-Drake, R., Felton, A. & Manning, A. (2008). Temproal changes in vertebrates during landscape transformation: a large-scale "natural experiment". Ecological Monographs 78, 567-590.
189. Lindenmayer, D. B. & Fischer, J. (2006). Habitat Fragmentation and Landscape Change. Island Press, Washington, DC.
190. Lindenmayer, D. B. & Fischer, J. (2007). Tackling the habitat fragmentation panchestron. Trends in Ecology and Evolution 22, 127-132.
191. Lindenmayer, D. B., Possingham, H. P., Lacy, R. C., McCarthy, M. A. & Pope, M. L. (2003). How accurate are population models? Lessons from landscape-scale population tests in a fragmented system. Ecology Letters 6, 41-47.
192. Lonsdorf, E., Kremen, C., Ricketts, T., Winfree, R., Williams, N. & Greenleaf, S. (2009). Modelling pollination services across agricultural landscapes. Annals of Botany 103, 1589-1600.
193. Loreau, M. (2000). Are communities saturated? On the relationship between alpha, beta and gamma diversity. Ecology Letters 3, 73-76.
194. Loreau, M., Mouquet, N. & Gonzalez, A. (2003). Biodiversity as spatial insurance in heterogeneous landscapes. Proceedings of the National Academy of Sciences 100, 12765-12770.
195. Lortie, C. J., Brooker, R. W., Choler, P., Kikvidze, Z., Michalet, R., Pugnaire, F. I. & Callaway, R. M. (2004). Rethinking plant community theory. Oikos 107, 433-438.
196. Lundberg, J. & Moberg, F. (2003). Mobile link organisms and ecosystem functioning: implications for ecosystem resilience and management. Ecosystems 6, 87-98.
197. Maas, B., Dwi Putra, D., Waltert, M., Clough, Y., Tscharntke, T. & Schulze, C. H. (2009). Six years of habitat modification in a tropical rainforest margin of Indonesia do not affect bird diversity but endemic forest species. Biological Conservation 142, 2665-2671.
198. MacArthur, R. H. & Wilson, E. O. (1967). The Theory of Island Biogeography. Princeton University Press, Princeton.
199. McCann, K. S., Rasmussen, J. B. & Umbanhowar, J. (2005). The dynamics of spatially coupled food webs. Ecology Letters 8, 513-523.
200. McGrady-Steed, J., Harris, P. M. & Morin, P. J. (1997). Biodiversity regulates ecosystem predictability. Nature 390, 162-165.
201. MEA (Millennium Ecosystem Assessment) (2005). Ecosystems and Human Well-Being: Synthesis. Island Press, Washington, DC.
202. Meehan, T. D., Werling, B. P., Landis, D. A. & Gratton, C. (2011). Agricultural landscape simplification and insecticide use in the Midwestern United States. Proceedings of the National Academy of Sciences PNAS, 108, 11500-11505.
203. Melbourne, B. A., Sears, A. L. W., Donahue, M. J. & Chesson, P. (2005). Applying scale transition theory to metacommunities in the field. In Metacommunities: Spatial Dynamics and Ecological Communities (eds M. Holyoak, M. A. Leibold and R. D. Holt), pp. 307-330. University of Chicago Press, Chicago.
204. Meyer, B., Jauker, F. & Steffan-Dewenter, I. (2009). Contrasting responses of species richness and density of hoverflies (Diptera: Syrphidae) to agricultural land-use change. Basic and Applied Ecology 10, 178-186.
205. Mortelli, A., Fagiani, S., Battisti, C., Capizzi, D. & Boitani, L. (2010). Independent effects of habitat loss, habitat fragmentation and structural connectivity on forest-dependent birds. Diversity and Distribution 16, 941-951.
206. Naeem, S. & Li, S. (1997). Biodiversity enhances ecosystem reliability. Nature 390, 507-509.
207. Noss, R. F. (1983). A regional landscape approach to maintain diversity. Bioscience 33, 700-706.
208. van Nouhuys, S. & Hanski, I. (2002). Multitrophic interactions in space: metacommunity dynamics in fragmented landscapes. In Multitrophic Level Interactions (eds T. Tscharntke and B. A. Hawkins), pp. 124-147. Cambridge University Press, Cambridge.
209. Öberg, S., Mayr, S. & Dauber, J. (2008). Landscape effects on recolonisation patterns of spiders in arable fields. Agriculture Ecosystems and Environment 123, 211-218.
210. Östman, Ö., Ekbom, B. & Bengtsson, J. (2001). Landscape heterogeneity and farming practice influence biological control. Basic and Applied Ecology 2, 365-371.
211. Parsche, S., Fründ, J. & Tscharntke, T. (2011). Experimental environmental change and mutualistic vs. antagonistic plant flower-visitor interactions. Perspectives in Plant Ecology, Evolution and Systematics 13, 27-35.
212. Pärtel, M., Szava-Kovats, R. & Zobel, M. (2011). Dark diversity: shedding light on absent species. Trends in Ecology & Evolution 26, 124-128.
213. Paschke, M., Abs, C. & Schmid, B. (2002). Effects of population size and pollen diversity on reproductive success and offspring size in the narrow endemic Cochlearia bavarica (Brassicaceae). American Journal of Botany 89, 1250-1259.
214. Peintinger, M., Bergamini, A. & Schmid, B. (2003). Species-area relationships and nestedness of four taxonomic groups in fragmented wetlands. Basic and Applied Ecology 4, 385-394.
215. Pereira, H. M. & Daily, G. C. (2006). Modeling biodiversity dynamics in countryside landscapes. Ecology 87, 1877-1885.
216. Perfecto, I. & Vandermeer, J. (2010). The agricultural matrix as an alternative to the land-sparing/agricultural intensification model: facing the food and biodiversity crises. Proceedings of the National Academy of Science 107, 5786-5791.
217. Perfecto, I., Vandermeer, J. & Wright, A. (2009). Nature's Matrix. Linking Agriculture, Conservation and Food Sovereignty. Earthscan, London.
218. Perović, D. J., Gurr, G. M., Raman, A. & Nicol, H. I. (2010). Effect of landscape composition and arrangement on biological control agents in a simplified agricultural system: a cost-distance approach. Biological Control 52, 263-270.
219. Petchey, O. L. & Gaston, K. J. (2006). Functional diversity: back to basics and looking forward. Ecology Letters 9, 741-758.
220. Peters, R. H. (1983). The Ecological Implications of Body Size. Cambridge University Press, Cambridge.
221. Peterson, G., Allen, C. R. & Holling, C. S. (1998). Ecological resilience, biodiversity, and scale. Ecosystems 1, 6-18.
222. Polis, G. A., Anderson, W. B. & Holt, R. D. (1997). Toward an integration of landscape and food web ecology: the dynamics of spatially subsidized food webs. Annual Review of Ecology, Evolution and Systematics 28, 289-316.
223. Polis, G. A., Power, M. E. & Huxel, G. R. (2004). Food Webs at the Landscape Level. The University of Chicago Press, Chicago.
224. Pope, S. E., Fahrig, L. & Merriam, H. G. (2000). Landscape complementation and metapopulation effects on leopard frog populations. Ecology 81, 2498-2508.
225. Priess, J. A., Mimler, M., Klein, A.-M., Schwarze, S., Tscharntke, T. & Steffan-Dewenter, I. (2007). Linking deforestation scenarios to pollination services and economic returns in coffee agroforestry systems. Ecological Applications 17, 407-417.
226. Prugh, L. R., Hodges, K. E., Sinclaer, A. R. E. & Brashares, J. S. (2008). Effect of habitat area and isolation on fragmented animal populations. Proceedings of the National Academy of Science 105, 20770-20775.
227. Pullin, A. S. (2002). Conservation Biology. Cambridge University Press, Cambridge.
228. Purvis, A. (2008). Phylogenetic approaches to the study of extinction. Annual Review of Ecology, Evolution and Systematics 39, 301-319.
229. Pywell, R. F., Warman, E. A., Hulmes, L., Hulmes, S., Nuttall, P., Sparks, T. H., Critchley, C. N. R. & Sherwood, A. (2006). Effectiveness of new agri-environment schemes in providing foraging resources for bumblebees in intensively farmed landscapes. Biological Conservation 129, 192-206.
230. Quinn, J. F. & Harrison, S. P. (1988). Effects of habitat fragmentation and isolation on species richness: evidence from biogeographic patterns. Oecologia 75, 132-140.
231. Rand, T. A. & Tscharntke, T. (2007). Contrasting effects of natural habitat loss on generalist and specialist aphid natural enemies. Oikos 116, 1353-1362.
232. Rand, T. A., Tylianakis, J. M. & Tscharntke, T. (2006). Spillover edge effects: the dispersal of agriculturally subsidized insect predators into adjacent natural habitats. Ecology Letters 9, 603-614.
233. Rand, T. A., van Veen, F. J. F. & Tscharntke, T. (2011). Landscape complexity differentially benefits generalized fourth, over specialized third, trophic level natural enemies. Ecography in press. (doi: 10.1111/j.1600-0587.2011.07016.x).
234. Rantalainen, M. L., Fritze, H., Haimi, J., Pennanen, T. & Setala, H. (2005). Species richness and food web structure of soil decomposer community as affected by the size of habitat fragment and habitat corridors. Global Change Biology 11, 1614-1627.
235. Rantalainen, M. L., Haimi, J., Fritze, H., Pennanen, T. & Setala, H. (2008). Soil decomposer community as a model system in studying the effects of habitat fragmentation and habitat corridors. Soil Biology & Biochemistry 40, 853-863.
236. Rantalainen, M. L., Haimi, J., Fritze, H. & Setala, H. (2006). Effects of small-scale habitat fragmentation, habitat corridors and mainland dispersal on soil decomposer organisms. Applied Soil Ecology 34, 152-159.
237. Reiners, W. A. & Driese, K. L. (2004). Transport Processes in Nature: Progagation of Ecological Influences through Environmental Space. Cambridge University Press, Cambridge.
238. Ricketts, T. H. (2001). The matrix matters: effective isolation in fragmented landscapes. The American Naturalist 158, 87-99.
239. Ricketts, T., Regetz, J., Steffan-Dewenter, I., Cunningham, S. A., Kremen, C., Bogdanski, A., Gemmill-Herren, B., Greenleaf, S. S., Klein, A.-M., Mayfield, M. M., Morandin, L. A., Ochieng, A. & Viana, F. B. (2008). Landscape effects on crop pollination services: are there general patterns? Ecology Letters 11, 499-515.
240. Ries, L., Fletcher, R. J., Battin, J. & Sisk, T. D. (2004). Ecological responses to habitat edges: mechanisms, models, and variability explained. Annual Review of Ecology, Evolution and Systematics 35, 491-522.
241. Roland, J. & Taylor, P. D. (1995). Herbivore-natural enemy interactions in fragmented and continuous forests. In Population Dynamics: New Approaches and Synthesis (eds N. Cappucino and P. W. Price), pp. 195-208. Academic Press, San Diego.
242. Roschewitz, I., Gabriel, D., Tscharntke, T. & Thies, C. (2005). The effects of landscape complexity on arable weed species diversity in organic and conventional farming. Journal of Applied Ecology 42, 873-882.
243. Rosenzweig, M. L. (1995). Species Diversity in Space and Time. University of Cambridge Press, Cambridge.
244. Rundlöf, M., Bengtsson, J. & Smith, H. G. (2008). Local and landscape effects of organic farming on butterfly species richness and abundance. Journal of Applied Ecology 45, 813-820.
245. Rundlöf, M. & Smith, H. G. (2006). The effect of organic farming on butterfly diversity depends on landscape context. Journal of Applied Ecology 43, 1121-1127.
246. Ryall, K. L. & Fahrig, V. (2006). Response of predators to loss and fragmentation of prey habitat: a review of theory. Ecology 87, 1086-1093.
247. Rytwinski, T. & Fahrig, L. (2011). Reproductive rate and body size predict road impacts on mammal abundance. Ecological Applications 21, 589-600.
248. Saunders, D. A., Hobbs, R. J. & Margules, C. R. (1991). Biological consequences of ecosystem fragmentation: a review. Conservation Biology 5, 18-32.
249. Scheid, B. E., Thies, C. & Tscharntke, T. (2011). Enhancing rape pollen beetle parasitism within sown flower fields along a landscape complexity gradient. Agricultural and Forest Entomology 13, 173-179.
250. Scherber, C., Eisenhauer, N., Weisser, W. W., Schmid, B., Voigt, W., Schulze, E. D., Roscher, C., Weigelt, A., Allan, E., Beßler, H., Bonkowski, M., Buchmann, N., Buscot, F., Clement, L. W., Ebeling, A., Engels, C., Fischer, M. S., Halle, S., Kertscher, I., Klein, A.-M., Koller, R., König, S., Kowalski, E., Kummer, V., Kuu, A., Lange, M., Lauterbach, D., Middelhoff, C., Migunova, V. D., Milcu, A., Müller, R., Partsch, S., Petermann, J. S., Renker, C., Rottstock, T., Sabais, A., Scheu, S., Schumacher, J., Temperton, V. M. & Tscharntke, T. (2010). Bottom-up effects of plant diversity on multitrophic interactions in a biodiversity experiment. Nature 468, 553-556.
251. Schmidt, M. H., Thies, C., Nentwig, W. & Tscharntke, T. (2008). Contrasting responses of arable spiders to the landscape mdifferent spatial scales. Journal of Biogeography 35, 157-166.
252. Schmidt, M. H. & Tscharntke, T. (2005a). Landscape context of sheetweb spider (Araneae: Linyphiidae) abundance in cereal fields. Journal of Biogeography 32, 467-473.
253. Schmidt, M. H. & Tscharntke, T. (2005b). The role of perennial habitats for Central European farmland spiders. Agriculture Ecosystems and Environment 105, 235-242.
254. Schmitz, O. J. (2008). Effects of predator hunting mode on grassland ecosystem function. Science 319, 952-954.
255. Schneider, K., Scheu, S. & Maraun, M. (2007). Microarthropod density and diversity respond little to spatial isolation. Basic Applied Ecology 8, 26-35.
256. Sekercioglu, C. H., Loarie, S. R., Brenes, F. O., Ehrlich, P. R. & Daily, G. C. (2007). Persistence of forest birds in the Costa Rican agricultural countryside. Conservation Biology 21, 482-494.
257. Simberloff, D. (1986). Design of nature reserves. In Wildlife Conservation Evaluation (ed. M. B. Usher), pp. 315-337. Chapman and Hall Ltd, London.
258. Simberloff, D. S. & Abele, V. (1976). Island biogeography theory and conservation practice. Science 191, 285-286.
259. Simianer, H. (2005). Using expected allele number as objective function to design between and within breed conservation of farm animal biodiversity. Journal of Animal Breeding and Genetics 122, 177-187.
260. Smith, H. G., Dänhardt, J., Lindström, A. & Rundlöf, M. (2010). Consequences of organic farming and landscape heterogeneity for species richness and abundance of farmland birds. Oecologia 162, 1071-1079.
261. Smith, A. C., Koper, N., Francis, C. M. & Fahrig, L. (2009). Confronting collinearity: comparing methods for disentangling the effects of habitat loss and fragmentation. Landscape Ecology 24, 1271-1285.
262. Sodhi, N. S., Brook, B. W. & Bradshaw, C. J. A. (2007). Tropical Conservation Biology. Blackwell Publishing, Oxford.
263. Sodhi, N. S. & Ehrlich, P. R. (2010). Conservation Biology for All. Oxford University Press, Oxford.
264. Sousa, J. P., Bolger, T., da Gama, M. M., Lukkari, T., Ponge, J. F., Simon, C., Traser, G., Vanbergen, A. J., Brennan, A., Dubs, F., Ivitis, E., Keating, A., Stofer, S. & Watt, A. D. (2006). Changes in Collembola richness and diversity along a gradient of land-use intensity: a pan European study. Pedobiologia 50, 147-156.
265. Srivastava, D. S. & Vellend, M. (2005). Biodiversity-ecosystem function research: is it relevant to conservation? Annual Review of Ecology Evolution and Systematics 36, 267-294.
266. Steffan-Dewenter, I., Kessler, M., Barkmann, J., Bos, M. M., Buchori, D., Erasmi, S., Faust, H., Gerold, G., Glenk, K., Gradstein, S. R., Guhardja, E., Harteveld, M., Hertel, D., Höhn, P., Kappas, M., Köhler, S., Leuschner, C., Maertens, M., Marggraf, R., Migge-Kleian, S., Mogea, J., Pitopang, R., Schaefer, M., Schwarze, S., Sporn, S. G., Steingrebe, A., Tjitrosoedirdjo, S. S., Tjitrosoemito, S., Twele, A., Weber, R., Woltmann, L., Zeller, M. & Tscharntke, T. (2007). Tradeoffs between income, biodiversity, and ecosystem functioning during tropical rainforest consersion and agroforestry intensification. Proceedings of the National Academy of Sciences 104, 4973-4978.
267. Steffan-Dewenter, I., Münzenberg, U., Bürger, C., Thies, C. & Tscharntke, T. (2002). Scale-dependent effects of landscape context on three pollinator guilds. Ecology 83, 1421-1432.
268. Steffan-Dewenter, I. & Tscharntke, T. (1997). Early succession of butterfly and plant communities on set-aside fields. Oecologia 109, 294-302.
269. Steffan-Dewenter, I. & Tscharntke, T. (2000). Butterfly community structure in fragmented habitats. Ecology Letters 3, 449-456.
270. Swift, T. L. & Hannon, S. J. (2010). Critical thresholds associated with habitat loss: a review of the concept, evidence, and applications. Biological Reviews 85, 35-53.
271. Swift, M. J., Izac, A. M. N. & van Noordwijk, M. (2004). Biodiversity and ecosystem services in agricultural landscapes-are we asking the right questions? Agriculture Ecosystems and Environment 104, 113-134.
272. Symstad, A. J., Tilman, D., Willson, J. & Knops, J. M. H. (1998). Species loss and ecosystem functioning: effects of species identity and community composition. Oikos 81, 389-397.
273. Terborgh, J., Lopez, L., Nuñez, V. P., Rao, M., Shahabuddin, G., Orijuela, G., Riveros, M., Ascanio, R., Adler, G. H., Lambert, T. D. & Balbas, L. (2001). Ecological meltdown in predator-free forest fragments. Science 294, 1923-1926.
274. Thies, C., Roschewitz, I. & Tscharntke, T. (2005). The landscape context of cereal aphid-parasitoid interactions. Proceedings of the Royal Society of London B 272, 203-210.
275. Thies, C., Steffan-Dewenter, I. & Tscharntke, T. (2003). Effects of landscape context on herbivory and parasitism at different spatial scales. Oikos 101, 18-25.
276. Thies, C., Steffan-Dewenter, I. & Tscharntke, T. (2008). Interannual landscape changes influence plant-herbivore-parasitoid interactions. Agriculture Ecosystems and Environment 125, 266-268.
277. Thies, C. & Tscharntke, T. (1999). Landscape structure and biological control in agroecosystems. Science 285, 893-895.
278. Tittler, R., Villard, M.-A. & Fahrig, L. (2009). How far do songbirds disperse? Ecography 32, 1051-1061.
279. Tscharntke, T. (1992). Fragmentation of Phragmites habitats, minimum viable population size, habitat suitability, and local extinction of moths, midges, flies, aphids, and birds. Conservation Biology 6, 530-536.
280. Tscharntke, T., Batáry, P. & Dormann, C. F. (2011). Set-aside management: how do succession, sowing patterns and landscape context affect biodiversity? Agriculture Ecosystems and Environment 143, 37-44.
281. Tscharntke, T., Bommarco, R., Clough, Y., Crist, T. O., Kleijn, D., Rand, T. A., Tylianakis, J. M., van Nouhuys, S. & Vidal, S. (2007a). Conservation biological control and enemy diversity on a landscape scale. Biological Control 43, 294-309 (reprint in: Biological Control 2008 45, 238-253).
282. Tscharntke, T. & Brandl, R. (2004). Plant-insect interactions in fragmented landscapes. Annual Review of Entomology 49, 405-430.
283. Tscharntke, T., Clough, Y., Bhagwat, S. A., Buchori, D., Faust, H., Hertel, D., Hölscher, D., Juhrbandt, J., Kessler, M., Perfecto, I., Scherber, C., Schroth, G., Veldkamp, E. & Wanger, T. C. (2011). Multifunctional shade-tree management in tropical agroforestry landscapes-a review. Journal of Applied Ecology 48, 619-629.
284. Tscharntke, T., Clough, Y., Wanger, T. C., Jackson, L., Motzke, I., Perfecto, I., Vandermeer, J. & Whitbread, A. (2012.)Global food security, biodiversity conservation and the future of agricultural intensification. Biological Conservation (under revision).
285. Tscharntke, T., Hochberg, M. E., Rand, T. A., Resh, V. H. & Krauss, J. (2007b). Author sequence and credit for contributions in multiauthored publications. PLoS Biology 5, 13-14.
286. Tscharntke, T., Klein, A.-M., Kruess, A., Steffan-Dewenter, I. & Thies, C. (2005a). Landscape perspectives on agricultural intensification and biodiversity-ecosystem service management. Ecology Letters 8, 857-874.
287. Tscharntke, T. & Kruess, A. (1999). Habitat fragmentation and biological control. In Theoretical Approaches to Biological Control (eds B. A. Hawkins and H. V. Cornell), pp. 190-205. Cambridge University Press, Cambridge.
288. Tscharntke, T., Rand, T. & Bianchi, F. (2005b). The landscape context of trophic interactions: insect spillover across the crop-noncrop interface. Annales Zoologici Fennici 42, 421-432.
289. Tscharntke, T., Sekercioglu, C. H., Dietsch, T. V., Sodhi, N. S., Hoehn, P. & Tylianakis, J. M. (2008). Landscape constraints on functional diversity of birds and insects in tropical agroecosystems. Ecology 89, 944-951.
290. Tscharntke, T., Steffan-Dewenter, I., Kruess, A. & Thies, C. (2002a). Characteristics of insect populations on habitat fragments-a mini review. Ecological Research 17, 229-239.
291. Tscharntke, T., Steffan-Dewenter, I., Kruess, A. & Thies, C. (2002b). Contribution of small habitat fragments to conservation of insect communities of grassland-cropland landscapes. Ecological Applications 12, 354-363.
292. Tubelis, D. P., Lindenmayer, D. B. & Cowling, A. (2004). Novel patch-matrix interactions: patch width influences matrix-use by birds. Oikos 107, 634-644.
293. Tylianakis, J. M., Didham, R. K., Bascompte, J. & Wardle, D. A. (2008a). Global change and species interactions in terrestrial ecosystems. Ecololgy Letters 11, 1351-1363.
294. Tylianakis, J. M., Didham, R. K. & Wratten, S. D. (2004). Improved fitness of aphid parasitoids receiving resource subsidies. Ecology 85, 658-666.
295. Tylianakis, J. M., Klein, A.-M. & Tscharntke, T. (2005). Spatiotemporal variation in the diversity of Hymenoptera across a tropical habitat gradient. Ecology 86, 3296-3302.
296. Tylianakis, J. M., Rand, T. A., Kahmen, A., Klein, A.-M., Buchmann, N., Perner, J. & Tscharntke, T. (2008b). Resource heterogeneity moderates the biodiversity-function relationship in real world ecosystems. PLoS Biology 6, e122; 947-956.
297. Tylianakis, J. M. & Romo, C. M. (2010). Natural enemy diversity and biological control: making sense of the context-dependency. Basic and Applied Ecology 11, 657-668.
298. Tylianakis, J. M., Tscharntke, T. & Klein, A.-M. (2006). Diversity, ecosystem function and stability of parasitoid-host interactions across a tropical habitat gradient. Ecology 87, 3047-3057.
299. Tylianakis, J. M., Tscharntke, T. & Lewis, O. T. (2007). Habitat modification alters the structure of tropical host-parasitoid food webs. Nature 445, 202-205.
300. Urban, M. C., Leibold, M. A., Amarasekare, P., De Meester, L., Gomulkiewicz, R., Hochberg, M. E., Klausmeier, C. A., Loeuille, N., de Mazancourt, C., Norberg, J., Pantel, J. H., Strauss, S. Y., Vellend, M. & Wade, M. J. (2008). The evolutionary ecology of metacommunities. Trends in Ecology and Evolution 23, 311-317.
301. Vance, M. D., Fahrig, L. & Flather, C. H. (2003). Relationship between minimum habitat requirements and annual reproductive rates in forest breeding birds. Ecology 84, 2643-2653.
302. Van de Voorde, T. F. J., Van der Putten, W. H., Gamper, H. A., Hol, W. H. G. & Bezemer, T. M. (2010). Comparing arbuscular mycorrhizal communities of individual plants in a grassland biodiversity experiment. New Phytologist 186, 746-754.
303. Van der Putten, W. H., Bardgett, R. D., de Ruiter, P. C., Hol, W. H. G., Meyer, K. M., Bezemer, T. M., Bradford, M. A., Christensen, S., Eppinga, M. B., Fukami, T., Hemerik, L., Molofsky, J., Schädler, M., Scherber, C., Strauss, S. Y., Vos, M. & Wardle, D. A. (2009). Empirical and theoretical challenges in aboveground-belowground ecology. Oecologia 161, 1-14.
304. Van der Wal, A., De Boer, W., Smant, W., Boschker, H. T. S., Bloem, J., Kardol, P., Van der Putten, W. H. & Van Veen, J. A. (2006). Fungal biomass development in a chronosequence of land abandonment. Soil Biology and Biochemistry 38, 51-60.
305. Veddeler, D., Klein, A.-M. & Tscharntke, T. (2006). Contrasting responses of bee communities to coffee flowering at different spatial scales. Oikos 112, 594-601.
306. Veddeler, D., Tylianakis, J., Tscharntke, T. & Klein, A.-M. (2010). Natural enemy diversity reduces temporal variability in wasp but not bee parasitism. Oecologia 162, 755-762.
307. Ward, D. (2006). Long-term effects of herbivory on plant diversity and functional types in arid ecosystems. In Large Herbivore Ecology, Ecosystem Dynamics and Conservation (eds K. Danell, R. Bergström, P. Duncan and J. Pastor), pp. 142-169. Cambridge University Press, Cambridge.
308. Wardle, D. A. (2002). Communities and Ecosystems: Linking the Aboveground and Belowground Components. Princeton University Press, Princeton.
309. Wardle, D. A. & van der Putten, W. H. (2002). Biodiversity, ecosystem functioning and above-ground-below-ground linkages. In Biodiversity and Ecosystem Functioning (eds M. Loreau, S. Naeem and P. Inchausti), pp. 157-168. Oxford University Press, Oxford.
310. Wardle, D. A., Yeates, G. W., Barker, G. M., Bellingham, P. J., Bonner, K. I. & Williamson, W. M. (2003). Island biology and ecosystem functioning in epiphytic soil communities. Science 301, 1717-1720.
311. Westphal, C., Steffan-Dewenter, I. & Tscharntke, T. (2003). Mass-flowering crops enhance pollinator densities at a landscape scale. Ecology Letters 6, 961-965.
312. Westphal, C., Steffan-Dewenter, I. & Tscharntke, T. (2006). Bumblebee species experience agricultural landscapes at different spatial scales. Oecologia 149, 289-300.
313. Whittingham, M. J. (2011). The future of agri-environment schemes: biodiversity gains and ecosystem service delivery? Journal of Applied Ecology 48, 509-513.
314. Wiens, J. A. (1989). Spatial scaling in ecology. Functional Ecology 3, 385-397.
315. Wiens, J. A. (1996). Wildlife in patchy environments: metapopulations, mosaics, and management. In Metapopulations and Wildlife Conservation (ed. D. R. McCullough), pp. 53-84. Island Press, Washington, DC.
316. Wiens, J. A., Moss, M. R., Turner, M. G. & Mladenoff, D. J. (2007). Foundation Papers in Landscape Ecology. Columbia University Press, New York.
317. Williams, N. M., Crone, E. E., Roulston, T. H., Minckley, R. L., Packer, L. & Potts, S. G. (2010). Ecological and life history traits predict bee species responses to environmental disturbances. Biological Conservation 143, 2280-2291.
318. Williams, N. M. & Kremen, C. (2007). Resource distributions among habitats determine solitary bee offspring production in a mosaic landscape. Ecological Applications 17, 910-921.
319. Wilsey, B. J. & Potvin, C. (2000). Biodiversity and ecosystem functioning: importance of species evenness in an old field. Ecology 81, 887-892.
320. Winfree, R., Aguilar, R., Vazques, D. P., Lebhuhn, G. & Aizen, M. A. (2009). A meta-analysis of bees' responses to anthropogenic disturbance. Ecology 90, 2068-2076.
321. Winfree, R. & Kremen, C. (2009). Are ecosystem services stabilized by differences among species? A test using crop pollination. Proceedings of the Royal Society B 276, 229-237.
322. Winfree, R., Williams, N. M., Dushoff, J. & Kremen, C. (2007). Wild bees provide insurance against ongoing honey bee losses. Ecology Letters 10, 1105-1113.
323. Wissinger, S. A. (1997). Cyclic colonization in predictably ephemeral habitats: a template for biological control in annual crop systems. Biological Control 10, 4-15.
324. With, K. A. & Crist, T. O. (1995). Critical threshold in species responses to landscape structure. Ecology 76, 2446-2459.
325. Yaacobi, G., Ziv, Y. & Rosenzweig, M. L. (2007). Habitat fragmentation may not matter to species diversity. Proceedings of the Royal Society B 274, 2409-2412.
326. Yachi, S. & Loreau, M. (1999). Biodiversity and ecosystem functioning in a fluctuating environment: the insurance hypothesis. Proceedings of the National Academy of Science 96, 1463-1468.
327. Young, H. S., McCauley, D. J., Dunbar, R. B. & Dirzo, R. (2010). Plants cause ecosystem nutrient depletion via the interruption of bird-derived spatial subsidies. Proceedings of the National Academy of Science 107, 2072-2077.
328. Zavaleta, E. S. & Hulvey, K. B. (2004). Realistic species losses disproportionately reduce grassland resistance to biological invaders. Science 306, 1175-1177.
329. Zhang, W., Ricketts, T. H., Kremen, C., Carney, K. & Swinton, S. M. (2007). Ecosystem services and dis-services to agriculture. Ecological Economics 64, 253-260.