Lichen epiphyte diversity: A species, community and trait-based review

Perspectives in Plant Ecology, Evolution and Systematics

Volumn 14, Issue 2, 2012, Pages 131-152

  Ellis, Christopher J ^a  

^a ROYAL BOTANIC GARDEN EDINBURGH   (United Kingdom)

Author keywords

Dispersal limitation; Ecosystem function; Forests; Life history; Niche; Succession

Indexed keywords

BOREAL FOREST; DISPERSAL; ECOSYSTEM FUNCTION; EPIPHYTE; FACILITATION; FOREST CANOPY; FOREST ECOSYSTEM; LICHEN; LIFE HISTORY TRAIT; NICHE; SPECIES DIVERSITY; SPECIES RICHNESS; TAXONOMY; TEMPERATE FOREST;

EID: 84859628224     PISSN: 14338319     EISSN: 16180437     Source Type: Journal    
DOI: 10.1016/j.ppees.2011.10.001     Document Type: Review

References (298)

1. Adams D.B., Risser P.G. Some factors influencing the frequency of bark lichens in north central Oklahoma. American Journal of Botany 1971, 58:752-757.
2. Affeld K., Sulliven J., Worner S.P., Didham R.K. Can spatial variation in epiphyte diversity and community structure be predicted from sampling vascular epiphytes alone?. Journal of Biogeography 2008, 35:2274-2288.
3. André H.M. Associations between corticolous microarthropod communities and epiphytic cover on bark. Holarctic Ecology 1985, 8:113-119.
4. Antoine M.E. An ecophysiological approach to quantifying nitrogen fixation by Lobaria oregana. The Bryologist 2004, 107:82-87.
5. Antoine M.E., McCune B. Contrasting fundamental and realized ecological niches with epiphytic lichen transplants in an old-growth Pseudotsuga forest. The Bryologist 2004, 107:163-173.
6. Aptroot A. Lichenized and saprobic fungal biodiversity of a single Elaeocarpus tree in Papua New Guinea, with the report of 200 species of ascomycetes associated with one tree. Fungal Diversity 2001, 6:1-11.
7. Aragón G., Martínez I., Izquierdo P., Belinchón R., Escurado A. Effects of forest management on epiphytic lichen diversity in Mediterranean forests. Applied Vegetation Science 2010, 13:183-194.
8. Armstrong R.A. Dispersal in a population of the lichen Hypogymnia physodes. Environmental and Experimental Botany 1987, 27:357-363.
9. Armstrong R.A. Dispersal, establishment and survival of soredia and fragments of the lichen, Hypogymnia physodes (L.) Nyl. New Phytologist 1990, 114:239-245.
10. Armstrong R.A. Soredial dispersal from individual soralia in the lichen Hypogymnia physodes (L.) Nyl. Environmental and Experimental Botany 1992, 32:55-63.
11. Arseneau M.-J., Sirois L., Ouellet J.-P. Effects of altitude and tree height on the distribution and biomass of fruticose arboreal lichens in an old growth balsam fir forest. écoscience 1997, 4:206-213.
12. Asplund J., Gauslaa Y. Mollusc grazing limits growth and early development of the old forest lichen Lobaria pulmonaria in broadleaved deciduous forests. Oecologia 2008, 155:93-99.
13. Asplund J., Larsson P., Vatne S., Gauslaa Y. Gastropod grazing shapes the vertical distribution of epiphytic lichens in forest canopies. Journal of Ecology 2010, 98:218-225.
14. Asta J., Erhardt W., Ferretti M., Fornasier F., Kirschbaum U., Nimis P.L., Purvis O.W., Pirintsos S., Scheidegger C., Van Haluwyn C., Wirth V. Mapping lichen diversity as an indicator of environmental quality. Monitoring with Lichens 2002, 273-279. Kluwer Academic, Dordrecht. P.L. Nimis, C. Scheidegger, P.A. Wolseley (Eds.).
15. Aude E., Poulsen R.S. Influence of management on the species composition of epiphytic cryptogams in Danish Fagus forests. Applied Vegetation Science 2000, 3:81-88.
16. Barkman J.J. Phytosociology and Ecology of Cryptogamic Epiphytes 1958, Van Corcum & Comp. N.V., Assen.
17. Bates J.W. Influence of chemical and site factors on Quercus and Fraxinus epiphytes at Loch Sunart, western Scotland: a multivariate analysis. Journal of Ecology 1992, 80:163-179.
18. Bates J.W., Brown D.W. Epiphyte differentiation between Quercus petraea and Fraxinus excelsior trees in a maritime area of south west England. Vegetatio 1981, 48:61-70.
19. Beals E.W. Ordination of some corticolous cryptogamic communities in south-central Wisconsin. Oikos 1965, 16:1-8.
20. Belinchón R., Martínez I., Escudero A., Aragón G., Valladares F. Edge effects on epiphytic communities in a Mediterranean Quercus pyrenaica forest. Journal of Vegetation Science 2007, 18:81-90.
21. Belinchón R., Martínez I., Otálora M.A.G., Aragón G., Dimas J., Escudero A. Fragment quality and matrix affect epiphytic performance in a Mediterranean forest landscape. American Journal of Botany 2009, 96:1974-1982.
22. Benner J.W. Epiphytes preferentially colonize high-phosphorus host trees in unfertilized Hawaiian montane forests. The Bryologist 2011, 114:335-345.
23. Benner J.W., Conroy S., Lunch C.K., Toyoda N., Vitousek P.M. Phosphorus fertilization increases the abundance and nitrogenase activity of the cyanolichen Pseudocyphellaria crocata in Hawaiian montane forests. Biotropica 2007, 39:400-405.
24. Benner J.W., Vitousek P.M. Development of a diverse epiphyte community in response to phosphorus fertilization. Ecology Letters 2007, 10:628-636.
25. Benson S., Coxson D.S. Lichen colonization and gap structure in wet-temperate rainforests of northern interior British Columbia. The Bryologist 2002, 105:673-692.
26. Berglund H., Jonsson B.G. Predictability of plant and fungal species richness of old-growth boreal forest islands. Journal of Vegetation Science 2001, 12:857-866.
27. Berglund H., Jonsson B.G. Verifying an extinction debt among lichens and fungi in northern Swedish boreal forests. Conservation Biology 2005, 19:338-348.
28. Berryman S., McCune B. Estimating epiphytic macrolichen biomass from topography, stand structure and lichen community data. Journal of Vegetation Science 2006, 17:157-170.
29. Binder M.D., Ellis C.J. Conservation of the rare British lichen Vulpicida pinastri: changing climate, habitat loss and strategies for mitigation. The Lichenologist 2008, 40:63-79.
30. Birks H.J.B. Quantitative palaeoenvironmental reconstructions. Statistical Modelling of Quaternary Science Data 1995, 161-254. Quaternary Research Association, Cambridge. D. Maddy, J.S. Brew (Eds.).
31. Boucher V.L., Nash T.H. The role of the fruticose lichen Ramalina menziesii in the annual turnover of biomass and macronutrients in a blue oak woodland. Botanical Gazette 1990, 151:114-118.
32. Boudreault C., Coxson D.S., Vincent E., Bergeron Y., Marsh J. Variation in epiphytic lichen and bryophyte composition and diversity along a gradient of productivity in Populus tremuloides stands of northeastern British Columbia, Canada. écoscience 2008, 15:101-112.
33. Boudreault C., Gauthier S., Bergeron Y. Epiphytic lichens and bryophytes on Populus tremuloides along a chronosequence in the southwestern boreal forest of Québec, Canada. The Bryologist 2000, 103:725-738.
34. Brunialti G., Frati L., Aleffi M., Marignani M., Rosati L., Burrascano S., Ravera S. Lichens and bryophytes as indicators of old-growth features in Mediterranean forests. Plant Biosystems 2010, 144:221-233.
35. Cácares M.E.S., Lücking R., Rambold G. Photophyte specificity and environmental parameters versus stochasticity as determinants for species compositon of corticolous crustose lichen communities in the Atlantic rain forest of northeastern Brazil. Mycological Progress 2007, 6:117-136.
36. Campbell J., Bradfield G.E., Prescott C.E., Fredeen A.L. The influence of overstorey Populus on epiphytic lichens in subboreal spruce forests of British Columbia. Canadian Journal of Forest Research 2010, 40:143-154.
37. Campbell J., Coxson D.S. Canopy microclimate and arboreal lichen loading in subalpine spruce-fir forest. Canadian Journal of Botany 2001, 79:537-555.
38. Campbell J., Fredeen A.L. Lobaria pulmonaria abundance as an indicator of macrolichen diversity in interior cedar-hemlock forests of east central British Columbia. Canadian Journal of Botany 2004, 82:970-982.
39. Campbell J., Fredeen A.L. Contrasting the abundance, nitrogen, and carbon of epiphytic macrolichen species between host trees and soil types in a sub-boreal forest. Canadian Journal of Botany 2007, 85:31-42.
40. Campbell J., Fredeen A.L., Prescott C.E. Decomposition and nutrient release from four epiphytic lichen litters in sub-boreal spruce forests. Canadian Journal of Forest Research 2010, 40:1473-1484.
41. Caruso A., Rudolphi J., Thor G. Lichen species diversity and substrate amounts in young planted boreal forests: a comparison between slash and stumps of Picea abies. Biological Conservation 2008, 141:47-55.
42. Casano L.M., del Campo E.M., García-Breijo F.J., Reig-Armiñana J., Gasulla F., del Hoyo A., Guéra A., Barreno E. Two Trebouxia algae with different physiological performances are ever-present in lichen thalli of Ramalina farinacea. Coexistence or competition?. Environmental Microbiology 2011, 13:806-818.
43. Cleavitt N.L., Dibble A.C., Werier D.A. Influence of tree composition upon epiphytic macrolichens and bryophytes in old forests of Acadia National Park, Maine. The Bryologist 2009, 112:467-487.
44. Clement J.P., Shaw D.C. Crown structure and the distribution of epiphyte functional group biomass in old-growth Pseudotsuga menziesii trees. écoscience 1999, 6:243-254.
45. Coppins A.M., Coppins B.J. Indices of Ecological Continuity for Woodland Epiphytic Lichen Habitats in the British Isles 2002, British Lichen Society, London.
46. Coppins B.J., Coppins A.M. Lichens - the biodiversity value of western woodlands. Botanical Journal of Scotland 2005, 57:141-153.
47. Coppins B.J., Coppins A.M. The lichens of Scottish native pinewoods. Forestry 2006, 79:249-259.
48. Cornelissen J.H.C., Ter Steege H. Distribution and ecology of epiphytic bryophytes and lichens in dry evergreen forest of Guyana. Journal of Tropical Ecology 1989, 5:131-150.
49. Coxson D.S., Coyle M. Niche partitioning and photosynthetic response of alectorioid lichens from subalpine spruce-fir forest in north-central British Columbia, Canada: the role of canopy microclimate gradients. The Lichenologist 2003, 35:157-175.
50. Crites S., Dale M.R.T. Diversity and abundance of bryophytes, lichens, and fungi in relation to woody substrate and successional stage in aspen mixedwood boreal forests. Canadian Journal of Botany 1998, 76:641-651.
51. Culberson W.L. The corticolous communities of lichens and bryophytes in the upland forests of northern Wisconsin. Ecological Monographs 1955, 25:215-231.
52. Degelius G. Biological studies on the epiphytic vegetation on twigs of Fraxinus excelsior. Acta Horti Gotoburgensis 1964, 27:11-55.
53. Dettki H., Esseen P.-A. Epiphytic macrolichens in managed and natural forest landscapes: a comparison at two spatial scales. Ecography 1998, 21:613-624.
54. Dettki H., Klintberg P., Essen P.-A. Are epiphytic lichens in young forests limited by local dispersal?. écoscience 2000, 7:317-325.
55. Doering M., Coxson D. Riparian alder ecosystems as epiphytic lichen refugia in sub-boreal spruce forests of British Columbia. Botany 2010, 88:144-157.
56. Edwards R.Y., Soos J., Ritcey R.W. Quantitative observation on epidendric lichens used as food by caribou. Ecology 1960, 41:425-431.
57. Ellis C.J., Coppins B.J. Contrasting functional traits maintain lichen epiphyte diversity in response to climate and autogenic succession. Journal of Biogeography 2006, 33:1643-1656.
58. Ellis C.J., Coppins B.J. Reproductive strategy and the compositional dynamics of crustose lichen communities on aspen (Populus tremula L.) in Scotland. The Lichenologist 2007, 39:377-391.
59. Ellis C.J., Coppins B.J. 19th Century woodland structure controls stand-scale epiphyte diversity in present-day Scotland. Diversity and Distributions 2007, 13:84-91.
60. Ellis C.J., Coppins B.J. Quantifying the role of multiple landscape-scale drivers controlling epiphyte composition and richness in a conservation priority habitat (juniper scrub). Biological Conservation 2009, 142:1291-1301.
61. Ellis C.J., Yahr R., Coppins B.J. Archaeobotanical evidence for a massive loss of epiphyte species richness during industrialization in southern England. Proceedings of the Royal Society B 2011, 278:3482-3489.
62. Ellyson W.J.T., Sillett S.C. Epiphyte communities on Sitka spruce in an old-growth redwood forest. The Bryologist 2003, 106:197-211.
63. Esseen P.-A. Host specificity and ecology of epiphytic macrolichens in some central Swedish spruce forests. Wahlenbergia 1981, 7:73-80.
64. Esseen P.-A., Renhorn K.-E., Petterson R.B. Epiphytic lichen biomass in managed and old-growth boreal forests: effect of branch quality. Ecological Applications 1996, 6:228-238.
65. Essen P.-A., Renhorn K.-E. Edge effects on an epiphytic lichen in fragmented forests. Conservation Biology 1998, 12:1307-1317.
66. Fedrowitz K., Kaasalainen U., Rikkinen J. Genotype variability of Nostoc symbionts associated with three epiphytic Nephroma species in a boreal forest landscape. The Bryologist 2011, 114:220-230.
67. Ferry B.W., Lodge E. Distribution and successsion of lichens associated with Prunus spinosa at Dungeness, England. The Lichenologist 1996, 28:129-143.
68. Flaherty E.A., Ben-David M., Smith W.P. Diet and food availability: implications for foraging and dispersal of Prince of Wales northern flying squirrels across managed landscapes. Journal of Mammalogy 2010, 91:79-91.
69. Forman R.T.T. Canopy lichens with blue-green algae: a nitrogen source in Colombian rain forest. Ecology 1975, 56:1176-1184.
70. Foucard T. Svenska Skorplavar 2001, Interpublishing, Stockholm.
71. Fritz ö. Vertical distribution of epiphytic bryophytes and lichens emphasizes the importance of old beeches in conservation. Biodiversity and Conservation 2009, 18:289-304.
72. Fritz O., Brunet J., Caldiz M. Interacting effects of tree charactersictics on the occurrence of rare epiphytes in a Swedish boreal forest. The Bryologist 2009, 112:488-505.
73. Fritz O., Gustafsson L., Larsson K. Does forest continuity matter in conservation? A study of epiphytic lichens and bryophytes in beech forests of southern Sweden. Biological Conservation 2008, 141:655-668.
74. Fritz O., Heilmann-Clausen J. Rot holes create key microhabitats for epiphytic lichens and bryophytes on beech (Fagus sylvatica). Biological Conservation 2010, 143:1008-1016.
75. Fritz ö., Niklasson M., Churski M. Tree age is a factor for the conservation of epiphytic lichens and bryophytes in beech forests. Applied Vegetation Science 2008, 12:93-106.
76. Gauslaa Y. The ecology of Lobarion pulmonariae and Parmelion caperatae in Quercus dominated forests in south-west Norway. The Lichenologist 1985, 17:117-140.
77. Gauslaa Y. The Lobarion, an epiphytic lichen community of ancient forests threatened by acid rain. The Lichenologist 1995, 27:59-76.
78. Gauslaa Y., Holien H. Acidity of boreal Picea abies-canopy lichens and their substratum, modified by local soils and airborne acidic depositions. Flora 1998, 193:249-257.
79. Gauslaa Y., Solhaug K.A. Photoinhibition in lichens depends on cortical characteristics and hydration. The Lichenologist 2004, 36:133-143.
80. Gerson U., Seaward M.R.D. Lichen-invertebrate associations. Lichen Ecology 1977, 69-119. Academic Press, London. M.R.D. Seaward (Ed.).
81. Gibb H., Ball J.P., Johansson T., Atlegrim O., Hjältén J., Danell K. Effects of management on coarse woody debris volume and composition in boreal forests in northern Sweden. Scandinavian Journal of Forest Research 2005, 20:213-222.
82. Gignac L.D., Dale M.R.T. Effects of fragment size and habitat heterogeneity on cryptogam diversity in the low-boreal forest of western Canada. The Bryologist 2005, 108:50-66.
83. Gough L.P. Cryptogam distributions on Pseudotsuga menziesii and Abies lasiocarpa in the Front Range, Boulder County, Colorado. The Bryologist 1975, 78:124-145.
84. Goward T., Arseneau A. Cyanolichen distribution in young unmanaged forests: a drip-zone effect?. The Bryologist 2000, 103:28-37.
85. Grime J.P. Vegetation classsification by reference to strategies. Nature 1974, 250:26-31.
86. Grime J.P. Evidence for the existence of three primary strategies in plants and its relevance to ecological and evolutionary theory. The American Naturalist 1977, 111:1169-1194.
87. Gu W.-D., Kuusinen M.T.K., Hanski I. Spatial pattern in the occurrence of the lichen Lobaria pulmonaria in managd and virgin boreal forests. Ecography 2001, 24:139-150.
88. Gunnarson B., Hake M., Hultengren S. A functional relationship between species richness of spiders and lichens in spruce. Biodiversity and Conservation 2004, 13:685-693.
89. Gustafsson L., Eriksson I. Factors of importance for the epiphytic vegetation of aspen Populus tremula with special emphasis on bark chemistry and soil chemistry. Journal of Applied Ecology 1995, 32:412-424.
90. Gustafsson L., Fiskesjö A., Ingelög T., Pettersson B., Thor G. Factors of importance to some lichen species of deciduous broad-leaved woods in southern Sweden. The Lichenologist 1992, 24:255-266.
91. Hale M.E. Vertical distribution of cryptogams in a virgin forest in Wisconsin. Ecology 1952, 33:398-406.
92. Hale M.E. Vertical distribution of cryptogams in a red maple swamp in Connecticut. The Bryologist 1965, 68:193-197.
93. Hale M.E. The Biology of Lichens 1983, Edward Arnold, London.
94. Hannah L., Carr J.L., Lankerani A. Human disturbance and natural habitat: a biome level analysis of a global data set. Biodiversity and Conservation 1995, 4:128-155.
95. Hanski I. Metapopulation Ecology 1999, Oxford University Press, Oxford.
96. Harris G.P. The ecology of corticolous lichens. I. The zonation on oak and birch in south Devon. Journal of Ecology 1971, 59:431-439.
97. Hauck M., Spribille T. The Mn/Ca and Mn/Mg ratios in bark as possible causes for the occurrence of Lobarion lichens on conifers in the dripzone of Populus in western North America. The Lichenologist 2002, 34:527-532.
98. Hauck M., Spribille T. The significance of precipitation and substrate chemistry for epiphytic lichen diversity in spruce-fir forests of the Salish Mountains, northwestern Montana. Flora 2005, 200:547-562.
99. Hayward G.D., Rosentreter R. Lichens as nesting material for northern flying squirrels in the northern Rocky Mountains. Journal of Mammalogy 1994, 75:663-673.
100. Hazell P., Gustafsson L. Retention of trees at final harvest - evaluation of a conservation technique using epiphytic bryophyte and lichen transplants. Biological Conservation 1999, 90:133-142.
101. Hedenås H., Blomberg P., Ericson L. Significance of old aspen (Populus tremula) trees for the occurrence of lichen photobionts. Biological Conservation 2007, 135:380-387.
102. Hedenås H., Bolukh V.O., Jonsson B.G. Spatial distribution of epiphytes on Populus tremula in relation to dispersal mode. Journal of Vegetation Science 2003, 14:233-242.
103. Hedenås H., Ericson L. Epiphytic macrolichens as conservation indicators: successional sequence in Populus tremula stands. Biological Conservation 2000, 93:43-53.
104. Hedenås H., Ericson L. Species occurrence at the stand level cannot be understood without considering the landscape context: cyanolichens on aspen in boreal Sweden. Biological Conservation 2008, 141:710-718.
105. Henderson A., Hackett D.J. Lichen and algal camouflage and dispersal in the psocid nymph Trichadenotecnum fasciatum. The Lichenologist 1986, 18:199-200.
106. Hilmo O. Distribution and succession of epiphytic lichens on Picea abies branches in a Boreal forest, central Norway. The Lichenologist 1994, 26:149-169.
107. Hilmo O. Growth and morphological response of old-forest lichens transplanted into a young and an old Picea abies forest. Ecography 2002, 25:329-335.
108. Hilmo O., Holien H. Epiphytic lichen response to the edge environment in a boreal Picea abies forest in central Norway. The Bryologist 2002, 105:48-56.
109. Hilmo O., Rocha L., Holien H., Gauslaa Y. Establishment success of lichen diaspores in young and old boreal rainforests: a comparison between Lobaria pulmonaria and L. scrobiculata. The Lichenologist 2011, 43:241-255.
110. Hilmo O., Såstad S.M. Colonization of old-forest lichens in a young and an old boreal Picea abies forest: an experimental approach. Biological Conservation 2001, 102:251-259.
111. Hinds H.R. Vertical distribution of lichens on aspen in Michigan. The Bryologist 1970, 73:626-628.
112. Holien H. Influence of site and stand factors on the distribution of crustose lichens of the Caliciales in a suboceanic spruce forest area in central Norway. The Lichenologist 1996, 28:315-330.
113. Holien H. The lichen flora of Picea abies in a suboceanic spruce forest area in central Norway with emphasis on the relationship to site and stand parameters. Nordic Journal of Botany 1997, 17:55-76.
114. Holien H. Lichens in spruce forest stands of different successional stages in central Norway with emphasis on diversity and old-growth species. Nova Hedwigia 1998, 66:283-324.
115. Hosokawa T., Odani N. The daily compensation period and vertical ranges of epiphytes in a beech forest. Journal of Ecology 1957, 45:901-906.
116. Hosokawa T., Odani N., Tagawa H. Causality of the distribution of corticolous species in forests with special reference to the physio-ecological approach. The Bryologist 1964, 67:396-411.
117. Hubbell S.P. The Unified Neutral Theory of Biodiversity and Biogeography 2001, Princeton University Press, Princeton.
118. Humphrey J.W., Davey S., Peace A.J., Ferris R., Harding K. Lichens and bryophyte communities of planted and semi-natural forests in Britain: the influence of site type, stand structure and deadwood. Biological Conservation 2002, 107:165-180.
119. Hutchinson G.E. Concluding remarks. Cold Spring Harbour Symposia on Quantitative Biology 1957, 22:415-427.
120. Hyvärinen M., Halonen P., Kauppi M. Influence of stand age and structure on the epiphytic lichen vegetation in the middle-boreal forests of Finland. The Lichenologist 1992, 24:165-180.
121. Ihlen P.G., Gjerde I., Sætersdal M. Structural indicators of richness and rarity of epiphytic lichens on Corylus avellana in two different forest types within a nature reserve in south-western Norway. The Lichenologist 2001, 33:215-229.
122. Jansová I., Soldán Z. The habitat factors that affect the composition of bryophyte and lichen communities on fallen logs. Preslia 2006, 78:67-86.
123. Jarman S.J., Kantvilas G. Epiphytes on an old Huon pine tree (Lagarostrobos franklinii) in Tasmanian rainforest. New Zealand Journal of Botany 1995, 33:65-78.
124. Jesberger J.A., Sheard J.W. A quantitative and multivariate analysis of corticolous lichen communities in the southern boreal forest of Saskatchewan. Canadian Journal of Botany 1973, 51:185-201.
125. Johansson P. Consequences of disturbance on epiphytic lichens in boreal and near boreal forests. Biological Conservation 2008, 141:1933-1944.
126. Johansson P., Ehrlén J. Influence of habitat quantity, quality and isolation on the distribution and abundance of two epiphytic lichens. Journal of Ecology 2003, 91:213-221.
127. Johansson P., Rydin H., Thor G. Tree age relationships with epiphytic lichen diversity and lichen life history traits on ash in southern Sweden. écoscience 2007, 14:81-91.
128. Johansson V., Bergman K.-O., Lättman H., Milberg P. Tree and site quality preferences of six epiphytic lichens growing on oaks in southeastern Sweden. Annales Botanici Fennici 2009, 46:496-506.
129. Johansson V., Snäll T., Johansson P., Ranius T. Detection probability and abundance estimation of epiphytic lichens based on height-limited surveys. Journal of Vegetation Science 2010, 21:332-341.
130. John E. Distribution patterns and interthalline interactions of epiphytic foliose lichens. Canadian Journal of Botany 1992, 70:818-823.
131. John E., Dale M.R.T. Neighbour relations within a community of epiphytic lichens and bryophytes. The Bryologist 1995, 98:29-37.
132. Jonsson B.G., Jonsell M. Exploring potential biodiversity indicators in boreal forests. Biodiversity and Conservation 1999, 8:1417-1433.
133. Jönsson M.T., Thor G., Johansson P. Environmental and historical effects on lichen diversity in managed and unmanaged wooded meadows. Applied Vegetation Science 2011, 14:120-131.
134. Josefsson T., Hellberg E., östlund L. Influence of habitat history on the distribution of Usnea longissima in boreal Scandinavia: a methodological study. The Lichenologist 2005, 37:555-567.
135. Jüriado I., Liira J., Csencsics D., Widmer I., Adolf C., Kohv K., Scheidegger C. Dispersal ecology of the endangered woodland lichen Lobaria pulmonaria in managed hemiboreal forest landscape. Biodiversity and Conservation 2011, 20:1803-1819.
136. Jüriado I., Liira J., Paal J., Suija A. Tree and stand level variables influencing diversity of lichens on temperate broad-leaved trees in boreo-nemoral floodplain forests. Biodiversity and Conservation 2009, 18:105-125.
137. Jüriado I., Paal J., Liira J. Epiphytic and epixylix lichen species diversity in Estonian natural forests. Biodiversity and Conservation 2003, 12:1587-1607.
138. Jüriado I., Suija A., Liira J. Biogeographical determinants of lichen species diversity on islets in the west-Estonian archipelago. Journal of Vegetation Science 2006, 17:125-134.
139. Kalgutkar R.M., Bird C.D. Lichens found on Larix lyallii and Pinus albicaulis in southwestern Alberta, Canada. Canadian Journal of Botany 1969, 47:627-648.
140. Kalwij J.M., Wagner H.H., Scheidegger C. Effects of stand-level disturbance on the spatial distribution of a lichen indicator. Ecological Applications 2005, 15:2015-2024.
141. Kantvilas G., Jarman S.J. Lichens and bryophytes on Eucalyptus obliqua in Tasmania: management implications in production forests. Biological Conservation 2004, 117:359-373.
142. Kantvilas G., Minchin P.R. An analysis of epiphytic lichen communities in Tasmanian cool temperate rainforest. Vegetatio 1989, 84:99-112.
143. Kapusta P., Szarek-Łukaszewska G., Kiszka J. Spatial analysis of species richness in a disturbed ecosystem (Niepołomice Forest, S Poland). The Lichenologist 2004, 36:249-260.
144. Kenkel N.C., Bradfield G.E. Epiphytic vegetation on Acer macrophyllum: a multivariate study of species-habitat relationships. Vegetatio 1986, 68:43-53.
145. Keon D.B., Muir P.S. Growth of Usnea longissima across a variety of habitats in the Oregon Coast Range. The Bryologist 2002, 105:233-242.
146. Kermit T., Gauslaa Y. The vertical gradient of bark pH of twigs and macrolichens in a Picea abies canopy not affected by acid rain. The Lichenologist 2001, 33:353-359.
147. Kershaw K.A. Preliminary observations on the distribution and ecology of epiphytic lichens in Wales. The Lichenologist 1964, 2:263-276.
148. Kirby K.J., Reid C.M., Thomas R.C., Goldsmith F.B. Preliminary estimates of fallen dead wood and standing dead trees in managed and unmanged forests in Britain. Journal of Applied Ecology 1998, 35:148-155.
149. Kivistö L., Kuusinen M. Edge effects on the epiphytic lichen flora of Picea abies in middle boreal Finland. The Lichenologist 2000, 32:387-398.
150. Knops J.M.H., Nash T.H., Schlesinger W.H. The influence of epiphytic lichens on the nutrient cycling of an oak woodland. Ecological Monographs 1996, 66:159-179.
151. Komposch H., Hafellner J. Diversity and vertical distribution of lichens in a Venezuelan tropical lowland rain forest. Selbyana 2000, 21:11-24.
152. Kruys N., Fries C., Jonsson B.G., Lämås T., Ståhl G. Wood-inhabiting cryptogams on dead Norway spruce (Picea abies) trees in managed Swedish boreal forests. Canadian Journal of Forest Research 1999, 29:178-186.
153. Kruys N., Jonsson B.G. Insular patterns of calicioid lichens in a boreal old-growth forest-wetland mosaic. Ecography 1997, 20:605-613.
154. Kuusinen M. Epiphytic lichen diversity on Salix caprea in old-growth southern and middle boreal forests of Finland. Annales Botanici Fennici 1994, 31:77-92.
155. Kuusinen M. Epiphyte flora and diversity on basal trunks of six old-growth forest tree species in southern and middle boreal Finland. The Lichenologist 1996, 28:443-463.
156. Kuusinen M., Siitonen J. Epiphytic lichen diversity in old-growth and managed Picea abies stands in southern Finland. Journal of Vegetation Science 1998, 9:283-292.
157. Lakatos M., Rascher U., Büdel B. Functional characteristics of corticolous lichens in the understorey of a tropical lowland rain forest. New Phytologist 2006, 172:679-695.
158. Lamit L.J., Bowker M.A., Holeski L.M., Næsborg R.R., Wooley S.C., Zinkgraf M., Lindroth R.L., Whitham T.G., Gehring C.A. Genetically-based trait variation within a foundation tree species influences a dominant bark lichen. Fungal Ecology 2011, 4:103-109.
159. Lang G.E., Reiners W.A., Heier R.K. Potential alteration of precipitation chemistry by epiphytic lichens. Oecologia 1976, 25:229-241.
160. Lang G.E., Reiners W.A., Pike L.H. Structure and biomass dynamics of epiphytic lichen communities of balsam fir forests in New Hampshire. Ecology 1980, 61:541-550.
161. Lättman H., Lindblom L., Mattson J.-E., Milberg P., Skage M., Ekman S. Estimating the dispersal capacity of the rare lichen Cliostomum corrugatum. Biological Conservation 2009, 142:1870-1878.
162. Lawrey J.D. Lichen herbivore preference: a test of two hypotheses. American Journal of Botany 1983, 70:1188-1194.
163. Lawrey J.D. Biological role of lichen substances. The Bryologist 1986, 89:111-122.
164. Lawrey J.D. Biotic interactions in lichen community development: a review. The Lichenologist 1991, 23:205-214.
165. Leppik E., Jüriado I., Liira J. Changes in stand structure due to the cessation of traditional land use in wooded meadows impoverish epiphytic lichen communities. The Lichenologist 2011, 43:257-274.
166. Lesica P., McCune B., Cooper S.V., Hong W.S. Differences in lichen and bryophyte communities between old-growth and managed second-growth forests in the Swan Valley, Montana. Canadian Journal of Botany 1991, 69:1745-1755.
167. Lewis J.E.J., Ellis C.J. Taxon-compared with trait-based analysis of epiphytes, and the role of tree species and tree age in community composition. Plant Ecology and Diversity 2010, 3:203-210.
168. Lie M.H., Arup U., Grytnes J.-A., Ohlson M. The importance of host tree age, size and growth rate as determinants of epiphytic lichen diversity in boreal spruce forests. Biodiversity and Conservation 2009, 18:3579-3596.
169. Liira J., Sepp T., Parrest O. The forest structure and ecosystem quality in conditions of anthropogenic disturbance along productivity gradients. Forest Ecology and Management 2007, 250:34-46.
170. Liu C., Ilvesniemi H., Westman C.J. Biomass of arboreal lichens and its vertical distribution in a boreal coniferous forest in central Finland. The Lichenologist 2000, 32:495-504.
171. Löbel S., Snäll T., Rydin H. Species richness patterns and metapopulation processes - evidence from epiphyte communities in boreo-nemoral forests. Ecography 2006, 29:169-182.
172. Lõhmus A., Lõhmus P. Epiphyte communities on the tree trunks of retention trees stabilise in 5 years after timber harvesting, but remain threatened due to tree loss. Biological Conservation 2010, 143:891-898.
173. Lõhmus A., Lõhmus P., Vellak K. Substratum diversity explains landscape-scale covariation in the species-richness of bryophytes and lichens. Biological Conservation 2007, 135:405-414.
174. Lõhmus P., Lõhmus A. Snags, and their lichen flora in old Estonian peatland forests. Annales Botanici Fennici 2001, 38:265-280.
175. Lommi S., Berglund H., Kuusinen M., Kuuluvainen T. Epiphytic lichen diversity in late-successional Pinus sylvestris forests along local and regional forest utilization gradients in eastern boreal Fennoscandia. Forest Ecology and Management 2010, 259:883-892.
176. Loppi S., Frati L. Influence of tree substrate on the diversity of epiphytic lichens: comparison between Tilia platyphyllos and Quecus ilex (Central Itlay). The Bryologist 2004, 107:340-344.
177. MacArthur R.H., Wilson E.O. The Theory of Island Biogeography 1967, Princeton University Press, Princeton.
178. Mandl N., Lehnert M., Kessler M., Gradstein S.R. A comparison of alpha and beta diversity patterns of ferns, bryophytes and macrolichens in tropical montane forests of southern Ecuador. Biodiversity and Conservation 2010, 19:2359-2369.
179. Marmor L., Tõrra T., Randlane T. The vertical gradient of bark pH and epiphytic macrolichen biota in relation to alkaline air pollution. Ecological Indicators 2010, 10:1137-1143.
180. McCune B. Gradients in epiphyte biomass in three Pseudotsuga-Tsuga forests of different ages in western Oregon and Washington. The Bryologist 1993, 96:405-411.
181. McCune B., Antos J.A. Epiphyte communities of the Swan Valley, Montana. The Bryologist 1982, 85:1-12.
182. McCune B., Caldwell B.A. A single phosphorus treatment doubles growth of cyanobacterial lichen transplants. Ecology 2009, 90:567-570.
183. McCune B., Hutchinson J., Berryman S. Concentration of rare epiphytic lichens along large streams in a mountainous watershed in Oregon, U.S.A. The Bryologist 2002, 105:439-450.
184. McCune B., Rosentreter R., Ponzetti J.M., Shaw D.C. Epiphyte habitats in an old conifer forest in western Washington, U.S.A. The Bryologist 2000, 103:417-427.
185. McMullin R.T., Duinker P.N., Richardson D.H.S., Cameron R.P., Hamilton D.C., Newmaster S.G. Relationships between the structural complexity and lichen community in coniferous forests of southwestern Nova Scotia. Forest Ecology and Management 2010, 260:744-749.
186. MEA, 2005. Ecosystems and Human Well Being. Biodiversity Synthesis in Millennium Ecosystem Assessment, Washington.
187. Meier E., Paal J. Cryptogams in Estonian alvar forests: species composition and their substrata in stands of different age and management intensity. Annales Botanici Fennici 2009, 46:1-20.
188. Mikhailova I., Trubina M., Vorobeichik E., Scheidegger C. Influence of environmental factors on the local-scale distribution of cyanobacterial lichens: case study in the North Urals, Russia. Folia Cryptogamica Estonica 2005, 41:45-54.
189. Milne J., Louwhoff S. Vertical distribution of bryophytes and lichens on a myrtle beech, Nothofagus cunninghamii (Hook.) Oerst. Hikobia 1999, 13:23-30.
190. Mitchell D. Spring and fall diet of the endangered West Virginia northern flying squirrel (Glaucomys sabrinus fuscus). American Midland Naturalist 2001, 146:439-443.
191. Moe B., Botnen A. A quantitative study of the epiphytic vegetation on pollarded trunks of Fraxinus excelsior at Havrå, Osterøy, western Norway. Plant Ecology 1997, 129:157-177.
192. Moe B., Botnen A. Epiphytic vegetation on pollarded trunks of Fraxinus excelsior in four different habitats at Grnde, Leikanger, western Norway. Plant Ecology 2000, 151:143-159.
193. Moen J., Jonsson B.G. Edge effects on liverworts and lichens in forest patches in a mosaic of boreal forest and wetland. Conservation Biology 2003, 17:380-388.
194. Moning C., Müller J. Critical forest age thresholds for the diversity of lichens, molluscs and birds in beech (Fagus sylvatica L.) dominated forests. Ecological Indicators 2009, 9:922-932.
195. Moning C., Werth S., Dziock F., Bässler C., Bradtka J., Hothorn T., Müller J. Lichen diversity in temperate montane forests is influenced by forest structure more than climate. Forest Ecology and Management 2009, 258:745-751.
196. Morley S.E., Gibson M. Successional changes in epiphytic rainforest lichens: implications for the management of rainforest communities. The Lichenologist 2010, 42:311-321.
197. Nascimbene J., Brunialti G., Ravera S., Frati L., Caniglia G. Testing Lobaria pulmonaria (L.) Hoffm. as an indicator of lichen conservation importance of Italian forests. Ecological Indicators 2010, 10:353-360.
198. Nascimbene J., Marini L., Caniglia G., Cester D., Nimis P.L. Lichen diversity on stumps in relation to wood decay in subalpine forests of northern Italy. Biodiversity and Conservation 2008, 17:2661-2670.
199. Nascimbene J., Marini L., Motta R., Nimis P.L. Lichen diversity of coarse woody debris in a Pinus-Larix stand in the Italian Alps. The Lichenologist 2008, 40:153-163.
200. Nascimbene J., Marini L., Motta R., Nimis P.L. Influence of tree age, tree size and crown structure on lichen communities in mature Alpine spruce forests. Biodiversity and Conservation 2009, 18:1509-1522.
201. Nascimbene J., Marini L., Nimis P.L. Influence of forest management on epiphytic lichens in a temperate beech forest of northern Italy. Forest Ecology and Management 2007, 247:43-47.
202. Nascimbene J., Marini L., Nimis P.L. Influence of tree species on epiphytic macrolichens in temperate mixed forests of northern Italy. Canadian Journal of Forest Research 2009, 39:785-791.
203. Nascimbene J., Marini L., Nimis P.L. Epiphytic lichen diversity in old-growth and managed Picea abies stands in Alpine spruce forests. Forest Ecology and Management 2010, 260:603-609.
204. Neitlich P.N., McCune B. Hotspots of epiphytic lichen diversity in two young managed forests. Conservation Biology 1996, 11:172-182.
205. Nilsson S.G., Arup U., Baranowski R., Ekman S. Tree-dependent lichens and beetles as indicators in conservation forests. Conservation Biology 1995, 9:1208-1215.
206. Nimis P.L. Phytogeography and ecology of epiphytic lichens at the southern rim of the Clay Belt (N-Ontario, Canada). The Bryologist 1985, 88:315-324.
207. Nordén B., Appelqvist T. Conceptual problems of ecological continuity and its bioindicators. Biodiversity and Conservation 2001, 10:779-791.
208. Nordén B., Paltto H., Götmark F., Wallin K. Indicators of biodiversity, what do they indicate? Lessons for conservation of cryptogams in oak-rich forest. Biological Conservation 2007, 135:369-379.
209. Normann F., Weigelt P., Gehrig-Downie C., Gradstein S.R., Sipman H.J.M., Obregon A., Bendix J. Diversity and vertical distribution of epiphytic macrolichens in lowland rain forest and lowland cloud forest of French Guiana. Ecological Indicators 2010, 10:1111-1118.
210. öckinger E., Niklasson M., Nilsson S.G. Is local distribution of the epiphytic lichen Lobaria pulmonaria limited by dispersal capacity or habitat quality?. Biodiversity and Conservation 2005, 14:759-773.
211. Ohlson M., Söderström L., Hörnberg G., Zackrisson O., Hermansson J. Habitat qualities versus long-term continuity as determinants of biodiversity in boreal old-growth swamp forests. Biological Conservation 1997, 81:221-231.
212. Oksanen J. Impact of habitat, substrate and microsite classes on the epiphyte vegetation: interpretation using exploratory and canonical correspondence analysis. Annales Botanici Fennici 1988, 25:59-71.
213. Ott S. Differences in the developmental rates of lichens. Annales Botanici Fennici 1987, 24:385-393.
214. Ott S. The juvenile develoment of lichen thalli from vegetative diaspores. Symbiosis 1987, 3:57-74.
215. Ozanne C.M.P., Anhuf D., Boulter S.L., Keller M., Kitching R.L., Körner C., Meinzer F.C., Mitchell A.W., Nakashizuka T., Sliva Dias P.L., Stork N.E., Wright S.J., Yoshimura M. Biodiversity meets the atmosphere: a global view of forest canopies. Science 2003, 301:183-186.
216. öztürk ş., Güvenç ş. Comparison of the epiphytic lichen communities growing on various tree species on Mt. Uludaĝ (Bursa, Turkey). Turkish Journal of Botany 2010, 34:449-456.
217. Paltto H., Nordén B., Götmark F. Partial cutting as a conservation alternative for oak (Quercus spp.) forest - response of bryophytes and lichens on deadwood. Forest Ecology and Management 2008, 256:536-547.
218. Paltto H., Thomasson I., Nordén B. Multispecies and multiscale conservation planning: setting quantitative targets for red-listed lichens on ancient oaks. Conservation Biology 2010, 24:758-768.
219. Peck J.E., McCune B. Remnant trees and canopy lichen communities in western Oregon: a retrospective approach. Ecological Applications 1997, 7:1181-1187.
220. Perhans K., Appelgren L., Jonsson F., Nordin U., Söderström B., Gustaffson L. Retention patches as potential refugia for bryophytes and lichens in managed forest landscapes. Biological Conservation 2009, 142:1125-1133.
221. Peterson E.B., McCune B. The importance of hotspots for lichen diversity in forests of western Oregon. The Bryologist 2003, 106:246-256.
222. Petterson R.B., Ball J.P., Renhorn K.-E., Esseen P.-A., Sjöberg K. Invertebrate communities in boreal forest canopies as influenced by forestry and lichens with implications for passerine birds. Biological Conservation 1995, 74:57-63.
223. Pike L.H. The importance of lichens in mineral cycling. The Bryologist 1978, 81:247-257.
224. Pike L.H., Denison W.C., Tracy D.M., Sherwood M.A., Rhoades F.M. Floristic survey of epiphytic lichens and bryophytes growing on old-growth conifers in western Oregon. The Bryologist 1975, 78:389-402.
225. Pike L.H., Rydell R.A., Denison W.C. A 400-year-old Douglas fir tree and its epiphytes: biomass, surface area, and their distributions. Canadian Journal of Forest Research 1977, 7:680-699.
226. Pirintsos S.A., Diamantopoulos J., Stamou G.P. Analysis of the vertical distribution of epiphytic lichens on Pinus nigra (Mount Olympus, Greece) along an latitudinal gradient. Vegetatio 1993, 109:63-70.
227. Pöykko H., Hyvärinen M., Bačkor M. Removal of lichen secondary metabolites affects food choice and survival of lichenivorous moth larvae. Ecology 2005, 86:2623-2632.
228. Price K., Hochachka G. Epiphytic lichen abundance: effects of stand age and composition in coastal British Columbia. Ecological Applications 2001, 11:904-913.
229. Pykälä J. Effects of new forestry practices on rare epiphytic macrolichens. Conservation Biology 2004, 18:831-838.
230. Rambo T.R. Habitat preferences of an arboreal forage lichen in a Sierra Nevada old-growth mixed-conifer forest. Canadian Journal of Forest Research 2010, 40:1034-1041.
231. Rambo T.R. Structure and composition of corticolous epiphyte communities in a Sierra Nevada old-growth mixed-conifer forest. The Bryologist 2010, 113:55-71.
232. Ranius T., Eliasson P., Johansson P. Large-scale occurrence patterns of red-listed lichens and fungi on old oaks are influenced both by current and historical habitat density. Biodiversity and Conservation 2008, 17:2371-2381.
233. Ranius T., Johansson P., Berg N., Niklasson M. The influence of tree age and microhabitat quality on the occurrence of crustose lichnes associated with old oaks. Journal of Vegetation Science 2008, 19:653-662.
234. Reiners W.A., Olson R.K. Effects of canopy components on throughfall chemistry: an experimental approach. Oecologia 1984, 63:320-330.
235. Renhorn K.-E., Esseen P.-A., Palmqvist K., Sunberg B. Growth and vitality of epiphytic lichens. I. Responses to microclimate along a forest edge-interior gradient. Oecologia 1997, 109:1-9.
236. Rheault H., Drapeau P., Bergeron Y., Esseen P.-A. Edge effects on epiphytic lichens in managed black spruce forests of eastern North America. Canadian Journal of Forest Research 2003, 33:23-32.
237. Rhoades F.M. Biomass of epiphytic lichens and bryophytes on Abies lasiocarpa on a Mt. Baker lava flow, Washington. The Bryologist 1981, 84:39-47.
238. Richardson D.H.S., Young C.M. Lichens and vertebrates. Lichen Ecology 1977, Academic Press, London. M.R.D. Seaward (Ed.).
239. Rikkinen J., Oksanen J., Lohtander K. Lichen guilds share related cyanobacterial symbionts. Science 2002, 297:357.
240. Rodwell J.S. British Plant Communities, volume 1. Woodlands and Scrub 1991, Cambridge University Press, Cambridge.
241. Rogers R.W. Succession and survival strategies in lichen populations on a palm trunk. Journal of Ecology 1988, 76:759-776.
242. Rogers R.W. Ecological strategies of lichens. The Lichenologist 1990, 22:149-162.
243. Rolstad J., Gjerde I., Gunderson V.S., Sætersdal M. Use of indicator species to assess forest continuity: a critique. Conservation Biology 2002, 16:253-257.
244. Rolstad J., Gjerde I., Storaunet K.O., Rolstad E. Epiphytic lichens in Norwegian coastal spruce forest: historic logging and present forest structure. Ecological Applications 2001, 11:421-436.
245. Rolstad J., Rolstad E. Does tree age predict the occurrence and abundance of Usnea longissima in multi-aged submontane Picea abies stands?. The Lichenologist 1999, 31:613-625.
246. Rose F. The epiphytes of oak. The British Oak 1974, 250-273. E.W. Classey, London. M.G. Morris, F.H. Perrring (Eds.).
247. Rose F. Lichenological indicators of age and environmental continuity in woodlands. Lichenology: Progress and Problems 1976, 279-307. Academic Press, London and New York. D.H. Brown, D.L. Hawksworth, R.H. Bailey (Eds.).
248. Rose F. Ancient British woodlands and their epiphytes. British Wildlife 1993, 5:83-93.
249. Ruchty A., Rosso A.L., McCune B. Changes in epiphyte communities as the shrub, Acer circinatum, dvelops and ages. The Bryologist 2001, 104:274-281.
250. Sætersdal M., Gjerde I., Blom H. Indicator species and the problem of spatial inconsistency in nestedness patterns. Biological Conservation 2005, 122:305-316.
251. Scheidegger C. Early development of transplanted isidioid soredia of Lobaria pulmonaria in an endangered population. The Lichenologist 1995, 27:361-374.
252. Scheidegger C., Groner U., Keller C., Stofer S. Biodiversity assessment tools - lichens. Monitoring with Lichens - Monitoring Lichens 2002, 359-365. Kluwer Academic Publishers, Dordrecht. P.L. Nimis, C. Scheidegger, P.A. Wolseley (Eds.).
253. Schmitt C.K., Slack N.G. Host specificity of epiphytic lichens and bryophytes: a comparison of the Adirondack Mountains (New York) and the southern Blue Ridge Mountains (North Carolina). The Bryologist 1990, 93:257-274.
254. Selva S.B. Lichen diversity and stand continuity in the northern hardwoods and spruce-fir forests on northern New England and western New Brunswick. The Bryologist 1994, 97:424-429.
255. Shirazi A.M., Muir P.S., McCune B. Environmental factors influencing the distribution of the lichens Lobaria oregana and L. pulmonaria. The Bryologist 1996, 99:12-18.
256. Shorrocks B., Marsters J., Ward I., Evennett P.J. The fractal dimension of lichens and the distribution of arthropod body lengths. Functional Ecology 1991, 5:457-460.
257. Sillett S.C. Growth rates of two epiphytic cyanolichen species at the edge and in the interior of a 700-year-old Douglas fir forest in the western Cascades of Oregon. The Bryologist 1994, 97:321-324.
258. Sillett S.C. Branch epiphyte assemblages in the forest interior and on the clearcut edge of a 700-year-old Douglas fir canopy in western Oregon. The Bryologist 1995, 98:301-312.
259. Sillett S.C., Goslin M.N. Distribution of epiphytic macrolichens in relation to remnant trees in a multiple-age Douglas-fir forest. Canadian Journal of Forest Research 1999, 29:1204-1215.
260. Sillett S.C., McCune B. Survival and growth of cyanolichen transplants in Douglas-fir forest canopies. The Bryologist 1998, 101:20-31.
261. Sillett S.C., McCune B., Peck J.E., Rambo T.R. Four years of epiphyte colonization in Douglas-fir forest canopies. The Bryologist 2000, 103:661-669.
262. Sillett S.C., McCune B., Peck J.E., Rambo T.R., Ruchty A. Dispersal limitations of epiphytic lichens result in species dependent on old-growth forests. Ecological Applications 2000, 10:789-799.
263. Sillett S.C., Van Pelt R. Trunk reiteration promotes epiphytes and water storage in an old-growth redwood forest canopy. Ecological Monographs 2007, 77:335-359.
264. Smith C.W., Aptroot A., Coppins B.J., Fletcher A., Gilbert O.L., James P.W., Wolseley P.A. The Lichens of Great Britain and Ireland 2010, The British Lichen Society, London.
265. Söderström L. The occurrence of epixylic bryophyte and lichen species in an old natural and a managed forest stand in northeast Sweden. Biological Conservation 1988, 45:169-178.
266. Söderström L. Sequence of bryophytes and lichens in relation to substrate variables of decaying confiferous wood in northern Sweden. Nordic Journal of Botany 1988, 8:89-97.
267. Sollins P., Grier C.C., McCorison F.M., Cromack K., Fogel R., Fredriksen R.L. The internal element cycles of an old-growth Douglas-fir ecosystem in western Oregon. Ecological Monographs 1980, 50:261-285.
268. Spier L., Van Dobben H.F., Van Dort K. Is bark pH more important than tree spcies in determining the composition of nitrophytic or acidophytic lichen floras?. Environmental Pollution 2010, 158:3607-3611.
269. Spribille T., Thor G., Bunnell F.L., Goward T., Björk C.R. Lichens on dead wood: species-substrate relationships in the epiphytic lichen floras of the Pacific northwest and Fennoscandia. Ecography 2008, 31:741-750.
270. St.Clair L.L., Rushforth S.R., Brotherson J.D. The influence of microhabitat on diversity, distribution and abundance of corticolous lichens in Zion National Park, Utah and Navajo National Monument, Arizona. Mycotaxon 1986, 26:253-262.
271. Stofer S., Bergamini A., Aragón G., Carvalho P., Coppins B.J., Davey S., Dietrich M., Farkas E., Kärkkäinen K., Keller C., Lökös L., Lommi S., Máguas C., Mitchell R., Pinho P., Rico V.J., Truscott A.-M., Wolseley P.A., Watt A., Scheidegger C. Species richness of lichen funtional groups in relation to land use intensity. The Lichenologist 2006, 38:331-353.
272. Stone D.F. Epiphyte succession on Quercus garryana branches in the Willamette Vally of western Oregon. The Bryologist 1989, 92:81-94.
273. Stubbs C.S. Patterns of distribution and abundance of corticolous lichens and their invertebrate associates on Quercus rubra in Maine. The Bryologist 1989, 92:453-460.
274. Svensson M., Johansson P., Thor G. Lichens of wooden barns and Pinus sylvestris snags in Dalarna, Sweden. Annales Botanici Fennici 2005, 42:351-363.
275. Tapper R. Dispersal and changes in the local distributions of Evernia prunastri and Ramalina farinacea. New Phytologist 1976, 77:725-734.
276. Tewari M., Upreti N., Pandey P., Singh S.P. Epiphytic succession on tree trunks in a mixed oak-cedar forest, Kumaun Himalaya. Vegetatio 1985, 63:105-112.
277. Thor G., Johansson P., Jönsson M.T. Lichen diversity and red-listed lichen species relationships with tree species and diameter in wooded meadows. Biodiversity and Conservation 2010, 19:2307-2328.
278. Tibell L. Crustose lichens as indicators of forest continuity in boreal coniferous forests. Nordic Journal of Botany 1992, 12:427-450.
279. Tibell L. Distribution patterns and dispersal strategies of Caliciales. Botanical Journal of the Linnean Society 1994, 116:159-202.
280. Topham P.B. Colonization, growth, succession and competition. Lichen Ecology 1977, 31-68. Academic Press, London. M.R.D. Seaward (Ed.).
281. Uliczka H., Angelstam P. Occurrence of epiphytic macrolichens in relation to tree species and age in managed Boreal forest. Ecography 1999, 22:396-405.
282. Uliczka H., Angelstam P. Assessing conservation values of forest stands based on specialised lichens and birds. Biological Conservation 2000, 95:343-351.
283. Van Leerdam A., Zagt R.J., Veneklaas E.J. The distribution of epiphyte growth-forms in the canopy of a Columbian cloud-forest. Vegetatio 1990, 87:59-71.
284. Veneklaas E.J., Zagt R.J., Van Leerdam A., Van Ek R., Broekhoven A.J., Genderen V. Hydrological properties of the epiphyte mass of a montane tropical forest, Colombia. Vegetatio 1990, 89:183-192.
285. Walser J.-C. Molecular evidence for the limited dispersal of vegetative propagules in the epiphytic lichen Lobaria pulmonaria. American Journal of Botany 2004, 91:1273-1276.
286. Walser J.-C., Holderegger R., Gugerli F., Hoebe S.E., Scheidegger C. Microsatellites reveal regional population differentiation in Lobaria pulmonaria, an epiphytic lichen. Molecular Ecology 2005, 14:457-467.
287. Walser J.-C., Zoller S., Büchler U., Scheidegger C. Species-specific detection of Lobaria pulmonaria (lichenized ascomycete) diaspores in litter samples trapped in snow cover. Molecular Ecology 2001, 10:2129-2138.
288. Wannebo-Nilsen K., Bjerke J.W., Beck P.S.A., Tømmervik H. Epiphytic macrolichens in spruce plantations and native birch forests along a coast-inland gradient in north Norway. Boreal Environment Research 2010, 15:43-57.
289. Wein R.W., Speer J.E. Lichen biomass in Acadian and boreal forests of Cape Breton Island, Nova Scotia. The Bryologist 1975, 78:328-333.
290. Werth S., Sork V.L. Identity and genetic structure of the photobiont of the epiphytic lichen Ramalina manziesii on three oak species in southern California. American Journal of Botany 2010, 97:821-830.
291. Werth S., Wagner H.H., Gugerli F., Holderegger R., Csencsics D., Kalwij J.M., Scheidegger C. Quantifying dispersal and establishment limitation in a population of an epiphytic lichen. Ecology 2006, 87:2037-2046.
292. Werth S., Wagner H.H., Holderegger R., Kalwij J.M., Scheidegger C. Effect of disturbance on the genetic diversity of an old-forest associated lichen. Molecular Ecology 2006, 15:911-921.
293. Whittaker R.H. Evolution and measurement of species diversity. Taxon 1972, 21:213-251.
294. Williams C.B., Sillett S.C. Epiphyte communities on redwood (Sequoia sempervirens) in northwestern California. The Bryologist 2007, 110:420-452.
295. Wolf J.H.D. Non-vascular epiphyte diversity patterns in the canopy of an upper montane rain forest (2550-3670m), Central Cordillera, Columbia. Selbyana 1995, 16:185-195.
296. Yarranton G.A. Distribution and succession of epiphytic lichens on black spruce near Cochrane, Ontario. The Bryologist 1972, 75:462-480.
297. Young P.J., Greer V.L., Six S.K. Characteristics of bolus nests of red squirrels in the Pinaleño and White mounatins of Arizona. The Southwestern Naturalist 2002, 47:267-275.
298. Zenner E.K. Does old-growth condition imply high live-tree structural complexity?. Forest Ecology and Management 2004, 195:243-258.