Systematics and Paleobiology of Hippotherium malpassii n. sp. (Equidae, Mammalia) from the latest Miocene of Baccinello V3 (Tuscany, Italy);Sistematica e paleobiologia di hippotherium malpassii n. sp. (Equidae, Mammalia) del miocene superiore di baccinello V3 (Toscana, Italia)

Bollettino della Societa Paleontologica Italiana

Volumn 50, Issue 3, 2011, Pages 175-208

  Bernor, Ray L ^a   Kaiser, Thomas M ^b   Nelson, Sherry V ^c   Rook, Lorenzo ^d  

^a HOWARD UNIVERSITY COLLEGE OF MEDICINE   (United States)

^b UNIVERSITY OF HAMBURG   (Germany)

^c UNIVERSITY OF NEW MEXICO   (United States)

^d UNIVERSITY OF FLORENCE   (Italy)

Author keywords

Baccinello cinigiano basin; Hippotherium malpassii n.sp.; Italy; Late miocene; Messinian; Paleoecology; Systematics; Tuscany; V3 fauna

Indexed keywords

GRASS; HABITAT TYPE; HORSE; MESSINIAN; NEW RECORD; NEW SPECIES; PALEOBIOLOGY; PALEOECOLOGY; PALEOENVIRONMENT; PALEOGEOGRAPHY; POPULATION STRUCTURE; SKELETON; TAXONOMY; WOODLAND;

EUROPE;

EQUIDAE; HIPPOTHERIUM PRIMIGENIUM; MAMMALIA; VALLESIA;

EID: 84855514216     PISSN: 03757633     EISSN: None     Source Type: Journal    
DOI: 10.4435/BSPI.2011.16     Document Type: Article

References (122)

1. Abbazzi L. (2001). Cervidae and Moschidae (Mammalia, Artiodactyla) from the Baccinello V3 faunal assemblage (Late Miocene, Late Turolian, Grosseto, Central Italy). Rivista Italiana di Paleontologia e Stratigrafia, 107: 10-123.
2. Abbazzi L., Delfino M., Gallai G., Trebini L. & Rook L. (2008). New data on the vertebrate assemblage of Fiume Santo (North-western Sardinia, Italy), and overview on the Late Miocene Tusco-sardinian paleobioprovince. Palaeontology, 51: 425-451.
3. Agustí J., Carbrera L., Garcés M., Krijgsman W., Oms O. & Parés J.M. (2001). A calibrated mammal scale for the Neogene of Western Europe. State of the Art. Earth Sciences Review, 52: 247-260.
4. Angelone C. & Rook L. (2011). Alilepus meini nov. sp. (Leporidae, Lagomorpha) from the Early Messinian of Tuscany (central-western Italy). Géobios, 44: 151-156.
5. Armour-Chelu M. & Bernor R.L. (2011). Equidae. In Harrison T. (ed.), Paleontology and geology of Laetoli: Human evolution in context (Fossil hominins and the associated fauna, Vol. 2), Springer Dordrecht: 295-326.
6. Armour-Chelu M., Bernor R.L. & Mittmann H.-W. (2006). Hooijer's hypodigm for "Hipparion" cf. ethiopicum (Equidae, Hipparioninae) from Olduvai, Tanzania and comparative material from the East African Plio-Pleistocene. In Nagel D. & van den Hoek Ostende L.W. (eds), Abhandlungen der Bayerischen Akademie der Wissenschaften, 30:15-24.
7. Benvenuti M., Bertini A. & Rook L. (1994). Facies analysis, vertebrate paleontology and palynology in the Late Miocene Baccinello-Cinigiano basin (southern Tuscany). Memorie della Società Geologica Italiana, 48: 415-423.
8. Benvenuti M., Papini M. & Rook L. (2001). Mammal biochronology, UBSU and paleoenvironment evolution in a post-collisional basin: evidence from the Late Miocene Baccinello-Cinigiano basin in southern Tuscany. Italy. Bollettino della Società Geologica Italiana, 120 (1995): 97-118.
9. Benvenuti M., Papini M. & Testa G. (1999). Sedimentary facies analysis in paleoclimatic reconstructions. Examples from the upper Miocene-Pliocene successions of southcentral Tuscany (Italy). In Agustí J., Rook L. & Andrews P. (eds.), The Evolution of Neogene Terrestrial Ecosystems in Europe, Cambridge University Press, Cambridge: 355-377.
10. Bernor R.L. (1985). Systematic and evolutionary relationships of the Hipparionine horses from Maragheh, Iran (Late Miocene, Turolian Age). Paleovertebrata, 15: 173-269.
11. Bernor R.L. (2007). The latest Miocene Hipparionine (Equidae) from Lemudong'o, Kenya. Kirtlandia, 56: 148-151.
12. Bernor R.L. & Armour-Chelu M. (1999). Towards an evolutionary history of African hipparionine horses. In Bromage T. & Schrenk F. (eds), African Biogeography, Climate Change and Early Hominid Evolution, Oxford University Press, New York: 189-215.
13. Bernor R. L., Armour-Chelu M., Gilbert H., Kaiser T.M. & Schulz E. (2010). Equidae. In Werdelin L. & Sanders W.L. (eds), Cenozoic mammals of Africa. University of California Press, Berkeley: 685-721.
14. Bernor R.L., Armour-Chelu M., Kaiser T. M. & Scott R.S. (2003a). An Evaluation of the Late MN 9 (Late Miocene, Vallesian Age), Hipparion Assemblage from Rudabánya (Hungary): Systematic Background, Functional Anatomy and Paleoecology. Coloquios de Paleontología, Vol. Ext., 1: 35-45.
15. Bernor R.L., Fahlbusch V., Andrews P., de Bruijn H., Fortelius M., Rögl F., Steininger F.F. & Werdelin L. (1996). The evolution of Western Eurasian Neogene mammal faunas: a chronologic, systematic, biogeographic, and paleoenvironmental synthesis. In Bernor R.L., Fahlbusch V. & Mittman H.W. (eds), The Evolution of Western Eurasian Neogene Mammal Faunas, Columbia University Press, New York: 449-469.
16. Bernor R.L., Fortelius M. & Rook L. (2001). Evolutionary biogeography and paleoecology of the "Oreopithecus bambolii Faunal Zone" (Late Miocene, Tusco-Sardinian Province). Bollettino della Società Paleontologica Italiana, 40: 139-148.
17. Bernor R.L. & Franzen J. (1997). The hipparionine horses from the Turolian Age (Late Miocene) locality of Dorn Dürkheim, Germany. Courier Forschungsinstitut Senckenberg, 197: 117-185.
18. Bernor R.L. & Haile-Selassie Y. (2009). Equidae. In Haile-Selassie Y. & WoldeGabriel G. (eds), Ardipithecus kadabba: Late Miocene Evidence from the Middle Awash, Ethiopia, University of California Press, Berkeley and Los Angeles: 397-428.
19. Bernor R. L. & Harris J.M. (2003). Systematics and evolutionary biology of the Late Miocene and Early Pliocene hipparionine horses from Lothagam, Kenya. In Harris J.M. & Leakey M.G. (eds), Lothagam: The Dawn of Humanity in Eastern Africa, Columbia University Press, New York: 387-438.
20. Bernor R.L., Heissig K. & Tobien H. (1987). Early Pliocene Perissodactyla from Sahabi, Libya. In Boaz N.T., El-Arnauti A., Gaziry A., de Heinzelin J. & Boaz D.D. (eds), Neogene Paleontology and Geology of Sahabi, Alan R. Liss, New York: 233-254.
21. Bernor R.L. & Hussain S. T. (1985). An assessment of the systematic, phylogenetic and biogeographic relationships of Siwalik hipparionine horses. Journal of Vertebrate Paleontology, 5: 32-87.
22. Bernor R.L. & Kaiser T.M. (2006). Systematics and paleoecology of the earliest Pliocene equid Eurygnathohippus hooijeri n. sp. from Langebaanweg, South Africa. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 103: 147-183.
23. Bernor R.L., Kaiser T.M., Kordos L. & Scott R.S. (1999). Stratigraphic Context, Systematic Position and Paleoecology of Hippotherium sumegense Kretzoi, 1984 from MN 10 (Late Vallesian of the Pannonian Basin). Mitteilungen der Bayrischen Staatssammlung für Paläontologie und historische Geologie, 39: 1-35.
24. Bernor R.L., Kaiser T.M. & Nelson S.V. (2004a). The oldest Ethiopian hipparion (Equinae, Perissodactyla) from Chorora: Systematics, paleodiet and paleoclimate. Courier Forschunginstitut Senckenberg, 246: 213-226.
25. Bernor R.L., Kaiser T.M. & Wolf D. (2008). Revisiting Sahabi equid species diversity, biogeographic patterns and diet preferences. In El-Arnauti A., Salem M., Pavlakis P. & Boaz N. (eds), Circum- Mediterranean Geology and Biotic Evolution During the Neogene Period: The Perspective from Libya. Garyounis Scientific Bulletin, Special Issue, 5: 159-167.
26. Bernor R.L., Kordos L., Rook L., Agustí J., Andrews P., Armour-Chelu M., Begun D.R., Cameron D.W., Damuth J., Daxner-Höck G., de Bonis L., Fejfar O., Fessaha N., Fortelius M., Franzen J., Gasparik M., Gentry A., Heissig K., Hernyak N., Kaiser T., Koufos G.D., Krolopp E., Jánossy D., Llenas M., Meszáros L., Müller P., Renne P., Rocek Z., Sen S., Scott R., Szyndlar Z., Topál Gy., Ungar P.S., Utescher T., Van Dam J.A., Werdelin L. & Ziegler R. (2004b). Recent advances on multidisciplinary research at Rudabánya, Late Miocene (MN9), Hungary: A compendium. Palaeontographica Italica, 89: 3-36.
27. Bernor R.L., Kovar J., Lipscomb D., Rögl F. & Tobien H. (1988). Systematic, stratigraphic and paleoenvironmental contexts of first appearing Hipparion in the Vienna Basin, Austria. Journal of Vertebrate Paleontology, 8: 427-452.
28. Bernor R.L., Mittmann H.-W., Kretzoi M. & Tobien H. (1993). A preliminary systematic assessment of the Rudabánya hipparions. Mitteilungen der Bayerischen Staatssammlung für Paläontologie und historische Geologie, 33: 1-20.
29. Bernor R.L., Qiu Z, & Hayek L. (1990). Systematic revision of Chinese hipparion species described by Sefve, 1927. American Museum Novitates, 2984: 1-60.
30. Bernor R.L. & Scott R.S. (2003). New interpretations of the systematics, biogeography and paleoecology of the Sahabi hipparions (latest Miocene), Libya. Geodiversitas, 25: 297-319.
31. Bernor R.L., Scott R. S., Fortelius M., Kappelman J. & Sen S. (2003b). Systematics and evolution of the Late Miocene Hipparions from Sinap, Turkey. In Fortelius M., Kappelman J., Sen S. & Bernor R.L. (eds), The Geology and Paleontology of the Miocene Sinap Formation, Turkey, Columbia University Press, New York: 220-281.
32. Bernor R.L., Scott R.S. & Haile-Selassie Y. (2005). A contribution to the evolutionary history of Ethiopian hipparionine horses: Morphometric evidence from the postcranial skeleton. Geodiversitas, 27: 133-158.
33. Bernor R.L., Tobien H., Hayek L.-A. C. & Mittmann H.-W. (1997). Hippotherium primigenium (Equidae, Mammalia) from the late Miocene of Höwenegg (Hegau, Germany). Andrias, 10: 1-230.
34. Bernor R.L., Tobien H. & Woodburne M.O. (1989). Patterns of Old World hipparionine evolutionary diversification. In Lindsay E., Fahlbusch V. & Mein P. (eds), European Neogene Mammal Chronology. Plenum Press, New York: 263-319.
35. Bernor R.L. & White T.D. (2009). Systematics and biogeography of "Cormohipparion" africanum, early Vallesian (MN9, ca. 10.5 MA) of Bou Hanifa, Algeria. Museum of Northern Arizona Bulletin, 65: 635-653.
36. Bernor R.L., Woodburne M.O. & Van Couvering J.A. (1980). A contribution to the chronology of some Old World Miocene faunas based on hipparionine horses. Géobios, 13: 705-739.
37. Bossio A., Costantini A., Foresi L., Mazzei R., Monterforti B., Salvatorini G. & Sandrelli F. (1991). Notizie preliminari sul Pliocene del bacino del medio Ombrone e della zona di Roccastrada. Atti della Società Toscana di Scienze Naturali, Memorie A, 98: 259-269.
38. Breymeyer A.I. & Van Dyne G.M. (1980). Grasslands, systems analysis and man. Cambridge: Cambridge University Press.
39. Casanovas-Vilar I., van Dam J.A., Moyá-Solá S. & Rook L. (2011). Late Miocene insular mice from the Tusco-Sardinian palaeobioprovince provide new insights on the palaeoecology of the Oreopithecus faunas. Journal of Human Evolution, 61: 42-49.
40. Casanovas-Vilar I., van Dam J.A., Trebini L. & Rook L. (2011). The rodents from the Late Miocene Oreopithecus-bearing site of Fiume Santo (Sardinia, Italy). Géobios, 44: 173-187.
41. Chesi F., Delfino M. & Rook L. (2009). Late Miocene Mauremys (Testudines, Geoemydidae) from Tuscany (Italy): evidence of terrapin persistence after a mammal turnover. Journal of Paleontology, 83: 379-388.
42. Codron D.J., Sponheimer M., Lee-Thorp J.A., Robinson T., Grant C.C. & deRuiter D. (2005). Assessing diet in savanna herbivores using stable carbon isotope rations of faeces. Koedoe, 48: 115-124.
43. De Terra H. (1956). New approaches to the problem of man's origin. Science, 124: 1282-1285.
44. Eisenmann V. (1995). What metapodial morphometry has to say about some Miocene hipparions. In Vrba E.S., Denton G.H., Partridge T.C. & Burckle L.H. (eds), Paleoclimate and Evolution, with Emphasis on Human Origins, Yale University Press, New Haven: 148-163.
45. Eisenmann V., Alberdi M.T., De Giuli C. & Staesche U. (1988). Studying Fossil Horses. Volume I - Methodology. In Woodburne M.O. & Sondaar P.Y. (eds), Collected papers after the "New York International Hipparion Conference, 1981", Brill, Leiden: 1-71.
46. Engesser B. (1983). Die jungtertiären Klinsäuger des Gebietes der Maremma (Toskana, Italien. 1.Teil: Gliridae (Rodentia, Mammalia). Eclogae Geologicae Helvetiae, 76: 763-780.
47. Engesser B. (1989). The Late Tertiary small mammals of the Maremma region (Tuscany, Italy): II part. Muridae and Cricetidae (Rodentia, Mammalia). Bollettino della Società Paleontologica Italiana, 29: 227-252.
48. Eronen J. & Rook L. (2004). The Mio-Pliocene European primate fossil record: dynamics and habitat tracking. Journal of Human Evolution, 4: 323-341.
49. Estes R. (1991). The Behavior Guide to African Mammals, including Hoofed Mammals, Carnivores, Primates. Los Angeles, University of California Press: 1-611.
50. Esu D. & Girotti O. (1989). Late Miocene and Early Pliocene continental and oligohaline molluscan faunas of Italy. Bollettino della Società Paleontologica Italiana, 28: 253-263.
51. Fortelius M. & Solounias N. (2000). Functional characterization of ungulate molars using the abrasion-attrition wear gradient: a new method for reconstructing paleodiets. American Museum Novitates, 3301: 1-36.
52. Gabunia L. (1959). The Hipparion history (according to SSSR Neogene material). Izdatelstvo Akademii Nauk SSSR, Moscou: 5-565 [in Russian].
53. Gagnon M. & Chew A.E. (2000). Dietary preferences in extant African Bovidae. Journal of Mammology, 81: 490-511.
54. Gallai G. & Rook L. (2011). Propotamochoerus provincialis (Gervais, 1859) (Suidae, Mammalia) from the latest Miocene (late Messinian; MN13) of Monticino Quarry (Brisighella, Emilia-Romagna, Italy). Bollettino della Società Paleontologica Italiana, 50: 29-34.
55. Getty R. (1982). The Anatomy of Domestic Animals. Saunders Company, Philadelphia: 1-1211.
56. Gilbert H.W. & Bernor R.L. (2008). Equidae. In: Gilbert H.W. & Asfaw B. (eds), Homo erectus - Pleistocene Evidence from the Middle Awash, Ethiopia. University of California Press, Berkeley: 133-166.
57. Gillet S., Lorenz H.G. & Woltersdorf F. (1965). Introduction à l'étude du Miocène supérieur de Baccinello (environs de Grosseto, Italie). Bullettin Service Carte Géologique de Alsace et Lorraine, 18: 31-42.
58. Gray J.E. (1821). On the natural arrangement of vertebrose animals. London Medical Repository, 15(1): 296-310.
59. Gromova V. (1952). Le genre Hipparion (transalded from Russian by St. Aubin). Bureau de Recherches Geologiques et Minieres, Orléans, CEDP, 12: 1-288.
60. Gruvaeus G. & Wainer H. (1972). Two additions to hierarchical cluster analysis. British Journal of Mathematical and Statatistical Psychology, 25: 200-206.
61. Hartigan J.A. (1975). Clustering algorithms, (John Wiley & Sons), New York.
62. Hay O.P. (1902). Bibliography and catalogue of the fossil Vertebrata of North America. U.S. Geological Survey Bulletin, 179: 1-868.
63. Hofmann R.R. (1989). Evolutionary steps of ecophysiological adaptation and diversification of ruminants: a comparative view of their digestive system. Oecologia, 78: 443-457.
64. Hulbert R.C., Jr. (1988). Cormohipparion and Hipparion (Mammalis, Perissodactyla, Equidae) from the Late Neogene of Florida. Bulletin of the Florida State Museum, Biolgical Sciences, 33: 229-338.
65. Hulbert R.C., Jr. & MacFadden B.J. (1991). Morphological transformation and cladogenesis at the base of the adaptive radiation of Miocene hypsodont horses. American Museum Novitates, 3000: 1-61.
66. Hürzeler J. & Engesser B. (1976). Les faunes des mammifères néogènes du Bassin de Baccinello (Grosseto, Italie). Comptes Rendus de l'Academie de Sciences de Paris, sér. 2D, 283: 333-336.
67. Janis C.M. (1988). An estimation of tooth volume and hypsodonty indices in ungulate mammals, and the correlation of these factors with dietary prefernce. In Russell D.E., Santoro J.P. & Sigogneau-Russell D. (eds), Teeth Revisited: Proceedings of the 7th International Symposium Dental Morphology, Paris 1986, 367-387, Mémoires du Museum national d'Histoire Naturelle, (Sér. C), Paris.
68. Janis C.M. & Ehrhard D. (1988). Correlation of relative muzzle width and relative incisor width with dietary preference in ungulates. Zoological Journal of the Linnean Society, 92: 267-284.
69. Kaiser T.M. (2003). The dietary regimes of two contemporaneous populations of Hippotherium primigenium (Perissodactyla, Equidae) from the Vallesian (upper Miocene) of Southern Germany. Palaeogeography, Palaeoclimatology, Palaeoecology, 198: 381-402.
70. Kaiser T.M. & Bernor R.L. (2007). The Baltavar Hippotherium: A mixed feeding Upper Miocene hipparion (Equidae, Perissodactyla) from Hungary (East-Central Europe); In Nagel D. & van den Hoek Ostende L.W. (eds), Abhandlungen der Bayerischen Akademie der Wissenschaften, 30: 241-267.
71. Kaiser T. M., Bernor R.L., Scott R.S., Franzen J.L. & Solounias N. (2003). New Interpretations of the Systematics and Palaeoecology of the Dorn-Dürkheim 1 Hipparions (Late Miocene, Turolian Age [MN 11]) Rheinhessen, Germany. Senckenbergiana lethaea, 83: 103-133.
72. Kaiser T.M. & Fortelius M. (2003). Differential mesowear in occluding upper and lower molars - opening mesowear analysis for lower molars and premolars in hypsodont equids. Journal of Morphology, 258: 67-83.
73. Kaiser T.M. & Solounias N. (2003). Extending the tooth mesowear method to extinct and extant equids. Geodiversitas, 25: 321-345.
74. Kaiser T.M., Solounias N., Fortelius M., Bernor R.L. & Schrenk F. (2000). Tooth mesowear analysis on Hippotherium primigenium from the Vallesian Dinotheriensande (Germany) - A blind test study. Carolinea, 58: 103-114.
75. Kingdon J. (1982). East African Mammals (Bovids). Vol. 3 Part C. Academic Press, London.
76. Kostopolous D. & Bernor R.L. (in press). The Maragheh bovids (Mammalia, Artiodactyla): systematic revision and biostratigraphic-biozoogeographic interpretation. Geodiversitas.
77. Koufos G.D. (1987a). Study of Pikermi hipparions Part I: Generalities and taxonomy. Bulletin du Museum Nationale d'Histoire Naturelle, Paris, ser. 4, 9, sec. C, 2: 197-252.
78. Koufos G.D. (1987b). Study of Pikermi hipparions Part II: Comparisons and odontograms. Bulletin du Museum Nationale d'Histoire Naturelle, Paris, ser. 4, 9, sec. C, 2: 327-363.
79. Koufos G.D. & Vlachou T. (2005). Equidae (Mammalia, Perissodactyla) from the late Miocene of Akkasdagi, Turkey. Geodiversitas, 27 (4): 633-705.
80. Ligios S., Benvenuti M., Gliozzi E., Papini M. & Rook L. (2008). Late Miocene palaeoenvironmental evolution of the Baccinello-Cinigiano Basin (Tuscany, central Italy) and new autoecological data on rare fossil fresh-to brackish-water ostracods. Palaeogeography Palaeoclimatology Palaeoecology, 264: 277-287.
81. Lorenz H.G. (1968). Stratigraphisches und mikropaläontologisches Untersuchungen des Braunkohlengebietes von Baccinello (Grosseto, Italien). Rivista Italiana di Paleontologia e Stratigrafia, 74: 147-270.
82. Lourens L., Hilgen F., Shackleton N.J., Laskar J. & Wilson J. (2004). Orbital tuning calibrations and conversions for the Neogene Period. In Gradstein F., Ogg J. & Smith A. (eds), A Geologic Time Scale. Cambridge University Press, Cambridge: 469-471.
83. MacFadden B. (1980). The Miocene horse Hipparion from North America and from the type locality in Soputhern France. Palaeontology 23(3): 617-635.
84. MacFadden B. (1984). Systematics and phylogeny of Hipparion, Neohipparion, Nanhippus, and Cormohipparion (Mammalia, Equidae) from the Miocene and Pliocene of the New World. Bulletin of the American Museum of Natural History, 179: 1-196.
85. MacFadden B.J. & Woodburne M.O. (1982). Systematics of the Neogene Siwalik hipparions (Mammalia, Equidae) based on cranial and dental morphology. Journal of Vertebrate Paleontology, 2: 185-218.
86. Marino B. & McElroy M. (1991). Isotopic composition of atmospheric CO2 inferred from carbon in plant cellulose. Nature, 349: 127-131.
87. Martini P.I. & Sagri M. (1993). Tectono-sedimentary characteristic of LateMiocene-Quaternary extensional basins of the Northern Appenines. Italy. Earth Science Review, 34: 197-233.
88. Meyer H.V. (1833). Fossile Säugethiere. Nova Acta Caesaris Leopoldi, 16 (2): 1-423.
89. Nelson S. (2005). Paleoseasonality inferred from equid teeth and intra-tooth isotopic variability. Palaeogeography, Palaeoclimatology, Palaeoecology, 222: 122-144.
90. Nickel R., Schummer A., Seiferle E., Wilkens H., Wille K.-H. & Frewein J. (1986). The Anatomy of the Domestic Animals, Volume 1: The Locomotor System. Verlag Paul Parey-Springer Verlag, Berlin.
91. Nowak R.M. (1999). Walker's Mammals of the World, 6th ed. John Hopkins University Press, Baltimore, London.
92. Owen R. (1848). Description of teeth and portions of jaws of two extinct anthracotherioid quadrupeds discovered in the Eocene deposits on the N. W. coast of the Isle of Wight. Quarterly Journal of the Geological Society of London, 4: 103-141.
93. Qiu Z., Weilong H. & Zhihui G. (1988). Chinese hipparionines from the Yushe Basin. Palaeontologica Sinica, Series C, 175: 1-250.
94. Quade J., Cerling T., Andrews P. & Alpagut B. (1995). Paleodietary reconstruction of Miocene faunas from Pasalar, Turkey using stable carbon and oxygen isotopes of fossil tooth enamel. Journal of Human Evolution, 28: 373-384.
95. Rook L. (1999). Late Turolian Mesopithecus (Mammalia, Primates, Colobinae) from Italy. Journal of Human Evolution, 36: 535-547.
96. Rook L. (2009). The Italian fossil primate record: an update and perspective for future research. Bollettino della Società Paleontologica Italiana, 48: 67-77.
97. Rook L., Abbazzi L., Chesi F., Delfino M., Ferretti M.P. & Gallai G. (2008). The Italian record of latest Miocene continental vertebrates. Bollettino della Società Paleontologica Italiana, 47: 191-194.
98. Rook L., Ficcarelli G. & Torre D. (1991). Messinian carnivores from Italy. Bollettino della Società Paleontologica Italiana, 30: 7-22.
99. Rook L., Gallai G. & Torre D. (2006). Lands and endemic mammals in the Late Miocene of Italy: Constraints for paleogeographic outlines of Tyrrhenian area. Palaeogeography Palaeoclimatology Palaeoecology, 238: 263-269.
100. Rook L., Harrison T. & Engesser B. (1996), The taxonomic status and biochronological implications of new finds of Oreopithecus from Baccinello. Journal of Human Evolution, 30: 3-27.
101. Rook L. & Martínez-Navarro B. (2004). Viverra howelli n. sp., a new viverrid (Carnivora, Mammalia) from the Baccinello-Cinigiano basin (latest Miocene, Italy). Rivista Italiana di Paleontologia e Stratigrafia, 110: 719-723.
102. Rook L., Oms O., Benvenuti M. & Papini M. (2011). Magnetostratigraphy of the Late Miocene Baccinello-Cinigiano basin (Tuscany, Italy) and the age of Oreopithecus bambolii faunal assemblages. Palaeogeography Palaeoclimatology Palaeoecology, 305: 286-294.
103. Rook L., Renne P., Benvenuti M. & Papini M. (2000). Geochronology of Oreopithecus bearing succession at Baccinello (Italy) and the extinction pattern of European Miocene hominoids. Journal of Human Evolution, 39: 577-582.
104. Rook L. & Rustioni M. (1991). Tapirus cf. arvernensis from the Late Turolian Baccinello V3 Faunal Assemblage (Grosseto, Italy). Bollettino della Società Paleontologica Italiana, 30: 325-327.
105. Rook L. & Torre D. (1995). Celadensia grossetana nov. sp., (Cricetidae, Rodentia) from the late Turolian Baccinello-Cinigiano basin (Southern Tuscany, Italy). Géobios, 28: 379-382.
106. Scott R.S. (2004). The Comparative Paleoecology of Late Miocene Eurasian Hominoids. Unpublished Ph.D. Dissertation thesis, The University of Texas at Austin.
107. Scott R.S., Armour-Chelu M. & Bernor R.L. (2005b). Evidence for Two Hipparion Species at Rudabánya II. In Bernor R.L., Kordos L. & Rook L. (eds), Multidisciplinary Research at Rudabánya. Palaeontographia Italica, 90: 213-216.
108. Scott R.S., Bernor R.L. & Raba W. (2005a). Hipparionine Horses of the Greater Pannonian Basin: Morphometric Evidence from the Postcranial Skeleton. In Bernor R.L., Kordos L. & Rook L. (eds), Multidisciplinary Research at Rudabánya. Palaeontographia Italica, 90: 195-210.
109. Scott R.S., Fortelius M., Huttunen K. & Armour-Chelu M.J. (2003). Abundance of Hipparion. In Fortelius M., Kappelman J., Sen S. & Bernor R.L. (eds), The Geology and Paleontology of the Miocene Sinap Formation, Turkey. Columbia University Press, New York: 380-397.
110. Skinner J. D. & Smithers R.H.N. (1990). The mammals of the southern African subregion. Pretoria. University of Pretoria.
111. Sponheimer M., Lee-Thorp J.A., DeRuither J.D.J., Smith J.M., van der Merwe N.J., Reed K., Grant C.C., Ayliffe L.K., Robinson T.F., Heidelberger C. & Marcus W. (2003). Diets of Southern African bovidae: Stable isotope evidence. Journal of Mammalogy, 84: 471-479.
112. Steinmann G. & Döderlein L. (1890). Elemente der Paläontologie. Wilhelm Engelmann, Leipzig: 1-848.
113. Stoppani A. (1880). Cenni sulle nuove miniere di lignite in territorio di Cana (Toscana). Tipografia degli Ingegneri, Milano: 1-8.
114. Van Wieren S.E. (1996). Digestive strategies in ruminants and nonruminants. Landbouwuniversiteit Wageningen, Den Haag: 1-191.
115. Vlachou T.D. & Koufos G.D. (2009). Equidae. In Koufos G.D. & Nagel D. (eds), The Late Miocene Mammal Faunas of the Mytilinii Basin, Samos Island, Greece: New Collection. Beiträge zur Paläontologie, 31: 207-281.
116. Webb S.D. & Hulbert R.C. Jr. (1986). Systematics and evolution of Pseudohipparion (Mammalia, Equidae) from the late Neogene of the Gulf Coastal Plain and the Great Plains. In Flanagan K.M. & Lillegraven J.A. (eds), Vertebrates, Phylogeny, and Philosophy. University of Wyoming Contributions in Geology, spec. paper, 3: 237-272.
117. Wolf D., Kaiser T.M., Nelson S.M., Nelson S.V., Semprebon G., Schwartz H. & Bernor R.L. (2010). The systematics and paleodiet of a diverse equid assemblage from Makuyuni, Northern Tanzania. Palaeodiversity, 3: 249-269.
118. Wood H.E. (1937). Perissodactyl suborders. Journal of Mammalogy, 18(1): 1-106
119. Woodburne M.O. (1989). Hipparion horses: a pattern of endemic evolution and intercontinental dispersal. In Prothero D.R. & Schoch R.M. (eds), The Evolution of the Perissodactyls. Oxford Monographs on Geology and Geophysics, 15: 197-233.
120. Woodburne M.O. (2007). Phyletic diversification of the Cormohipparion occidentale Complex (Mammalia, Perissodactyla, Equidae), Late Miocene, North America, and the origin of the Old World Hippotherium Datum. Bulletin of the American Museum of Natural History, 306: 1-138.
121. Woodburne M.O. & Bernor L.R. (1980). On superspecific groups of some Old World hipparionine horses. Journal of Paleontology, 54: 1319-1348.
122. Woodburne M.O., MacFadden B.J. & Skinner M.F. (1981). The North American "Hipparion" datum, and implications for the Neogene Old World. Géobios, 14: 1-32.