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Volumn 68, Issue 9, 2011, Pages 1615-1624

Influence of pinniped-caused injuries on the survival of adult Chinook salmon (Oncorhynchus tshawytscha) and steelhead trout (Oncorhynchus mykiss) in the Columbia River basin

Author keywords

[No Author keywords available]

Indexed keywords

ABUNDANCE; CONSERVATION PLANNING; DENSITY DEPENDENCE; ENDANGERED SPECIES; INJURY; INTERSPECIFIC INTERACTION; MIGRATION; PINNIPED; POPULATION DECLINE; PREDATION; RADIOTELEMETRY; SALMONID; SPECIES CONSERVATION; SUCCESSION; SURVIVAL;

EID: 80052532814     PISSN: 0706652X     EISSN: 12057533     Source Type: Journal    
DOI: 10.1139/f2011-064     Document Type: Article
Times cited : (15)

References (50)
  • 1
    • 0034255819 scopus 로고    scopus 로고
    • The nature of predation: Prey dependent, ratio dependent, or neither?
    • doi:10.1016/S0169-5347(00)01908-X. PMID:10884706
    • Abrams, P.A., and Ginzburg, L.R. 2000. The nature of predation: prey dependent, ratio dependent, or neither? Trends Ecol. Evol. 15(8): 337-341. doi:10.1016/S0169-5347(00)01908-X. PMID:10884706
    • (2000) Trends Ecol. Evol , vol.15 , Issue.8 , pp. 337-341
    • Abrams, P.A.1    Ginzburg, L.R.2
  • 2
    • 71549151643 scopus 로고    scopus 로고
    • Oceanic, riverine, and genetic influences on spring Chinook salmon migration timing
    • doi:10.1890/08-0477.1. PMID:20014573
    • Anderson, J.J., and Beer, W.N. 2009. Oceanic, riverine, and genetic influences on spring Chinook salmon migration timing. Ecol. Appl. 19(8): 1989-2003. doi:10.1890/08-0477.1. PMID:20014573
    • (2009) Ecol. Appl , vol.19 , Issue.8 , pp. 1989-2003
    • Anderson, J.J.1    Beer, W.N.2
  • 3
    • 0034586859 scopus 로고    scopus 로고
    • Trends and potential interactions between pinnipeds and fisheries of New England and the U.S.west coast
    • Baraff, L.S., and Loughlin, T.R. 2000. Trends and potential interactions between pinnipeds and fisheries of New England and the U.S. west coast. Mar. Fish. Rev. 62: 1-40
    • (2000) Mar. Fish. Rev , vol.62 , pp. 1-40
    • Baraff, L.S.1    Loughlin, T.R.2
  • 7
    • 0000644514 scopus 로고
    • The relation of size to the rate of oxygen consumption and sustained swimming speed of sockeye salmon (Oncorhynchus nerka)
    • doi:10.1139/f65-128
    • Brett, J.R. 1965. The relation of size to the rate of oxygen consumption and sustained swimming speed of sockeye salmon (Oncorhynchus nerka). J. Fish. Res. Board Can. 22(6): 1491-1501. doi:10.1139/f65-128
    • (1965) J. Fish. Res. Board Can , vol.22 , Issue.6 , pp. 1491-1501
    • Brett, J.R.1
  • 8
    • 0020895366 scopus 로고
    • Abundance, movements, and feeding-habits of harbor seals, Phoca vitulina, at Netarts and Tillamook Bays
    • Brown, R.F., and Mate, B.R. 1983. Abundance, movements, and feeding-habits of harbor seals, Phoca vitulina, at Netarts and Tillamook Bays, Oregon. Fish Bull. 81: 291-301
    • (1983) Oregon. Fish Bull , vol.81 , pp. 291-301
    • Brown, R.F.1    Mate, B.R.2
  • 10
    • 35348956487 scopus 로고    scopus 로고
    • Slow dam passage in adult Columbia River salmonids associated with unsuccessful migration: Delayed negative effects of passage obstacles or condition-dependent mortality?
    • doi:10.1139/f07-065
    • Caudill, C.C., Daigle, W.R., Keefer, M.L., Boggs, C.T., Jepson, M. A., Burke, B.J., Zabel, R.W., Bjornn, T.C., and Peery, C.A. 2007. Slow dam passage in adult Columbia River salmonids associated with unsuccessful migration: delayed negative effects of passage obstacles or condition-dependent mortality? Can. J. Fish. Aquat. Sci. 64(7): 979-995. doi:10.1139/f07-065
    • (2007) Can. J. Fish. Aquat. Sci , vol.64 , Issue.7 , pp. 979-995
    • Caudill, C.C.1    Daigle, W.R.2    Keefer, M.L.3    Boggs, C.T.4    Jepson, M.A.5    Burke, B.J.6    Zabel, R.W.7    Bjornn, T.C.8    Peery, C.A.9
  • 11
    • 0029503182 scopus 로고
    • The condition of fish escaping from fishing gears-a review
    • doi:10.1016/0165-7836(94)00301-C
    • Chopin, F.S., and Arimoto, T. 1995. The condition of fish escaping from fishing gears-a review. Fish. Res. 21(3-4): 315-327. doi:10. 1016/0165-7836(94)00301-C
    • (1995) Fish. Res , vol.21 , Issue.3-4 , pp. 315-327
    • Chopin, F.S.1    Arimoto, T.2
  • 12
    • 0030304330 scopus 로고    scopus 로고
    • Predator foraging capabilities and prey antipredator behaviours: Pre- versus postcapture constrains on sizedependent predator-prey interactions
    • doi:10.2307/3546209
    • Christensen, B. 1996. Predator foraging capabilities and prey antipredator behaviours: pre- versus postcapture constrains on sizedependent predator-prey interactions. Oikos, 76(2): 368-380. doi:10.2307/3546209
    • (1996) Oikos , vol.76 , Issue.2 , pp. 368-380
    • Christensen, B.1
  • 13
    • 0031948404 scopus 로고    scopus 로고
    • Frequency of pinniped-caused scars and wounds on adult spring-summer chinook and sockeye salmon returning to the Columbia River
    • doi:10.1577/1548-8675(1998)018<0046:FOPCSA>2.0.CO;2
    • Fryer, J.K. 1998. Frequency of pinniped-caused scars and wounds on adult spring-summer chinook and sockeye salmon returning to the Columbia River. N. Am. J. Fish. Manage. 18(1): 46-51. doi:10.1577/1548-8675(1998)018<0046:FOPCSA>2.0.CO;2
    • (1998) N. Am. J. Fish. Manage , vol.18 , Issue.1 , pp. 46-51
    • Fryer, J.K.1
  • 14
    • 33845700944 scopus 로고    scopus 로고
    • Behavioral thermoregulation and slowed migration by adult fall Chinook salmon in response to high Columbia River water temperatures
    • doi:10.1577/T04-113.1
    • Goniea, T.M., Keefer, M.L., Bjornn, T.C., Peery, C.A., Bennett, D. H., and Stuehrenberg, L.C. 2006. Behavioral thermoregulation and slowed migration by adult fall Chinook salmon in response to high Columbia River water temperatures. Trans. Am. Fish. Soc. 135(2): 408-419. doi:10.1577/T04-113.1
    • (2006) Trans. Am. Fish. Soc , vol.135 , Issue.2 , pp. 408-419
    • Goniea, T.M.1    Keefer, M.L.2    Bjornn, T.C.3    Peery, C.A.4    Bennett, D.H.5    Stuehrenberg, L.C.6
  • 15
    • 0027837377 scopus 로고
    • Variation in the numbers of shags Phalacrocorax aristotelis and common seals Phoca vitulina near the mouth of an Atlantic salmon Salmo salar river at the time of the smolt run
    • doi:10.2307/5205
    • Greenstreet, S.P.R., Morgan, R.I.G., Barnett, S., and Redhead, P. 1993. Variation in the numbers of shags Phalacrocorax aristotelis and common seals Phoca vitulina near the mouth of an Atlantic salmon Salmo salar river at the time of the smolt run. J. Anim. Ecol. 62(3): 565-576. doi:10.2307/5205
    • (1993) J. Anim. Ecol , vol.62 , Issue.3 , pp. 565-576
    • Greenstreet, S.P.R.1    Morgan, R.I.G.2    Barnett, S.3    Redhead, P.4
  • 16
    • 0028010248 scopus 로고
    • Prevalence of marine-mammal tooth and claw abrasions on adult anadromous salmonids returning to the Snake River
    • doi:10.1577/1548-8675(1994) 014<0661:POMMTA>2.3.CO;2
    • Harmon, J.R., Thomas, K.L., McIntyre, K.W., and Paasch, N.N. 1994. Prevalence of marine-mammal tooth and claw abrasions on adult anadromous salmonids returning to the Snake River. N. Am. J. Fish. Manage. 14(3): 661-663. doi:10.1577/1548-8675(1994) 014<0661:POMMTA>2.3.CO;2
    • (1994) N. Am. J. Fish. Manage , vol.14 , Issue.3 , pp. 661-663
    • Harmon, J.R.1    Thomas, K.L.2    McIntyre, K.W.3    Paasch, N.N.4
  • 17
    • 0002366911 scopus 로고
    • The functional response of predators to prey density and its role in mimicry and population regulation
    • doi:10.4039/entm9745fv
    • Holling, C.S. 1965. The functional response of predators to prey density and its role in mimicry and population regulation. Mem.Entomol. Soc. Can. 97(45): 1-60. doi:10.4039/entm9745fv
    • (1965) Mem.Entomol Soc. Can , vol.97 , Issue.45 , pp. 1-60
    • Holling, C.S.1
  • 19
    • 0037285377 scopus 로고    scopus 로고
    • Trends and status of harbor seals in Washington state: 1978-1999
    • doi:10.2307/3803076
    • Jeffries, S., Huber, H., Calambokidis, J., and Laake, J. 2003. Trends and status of harbor seals in Washington state: 1978-1999. J. Wildl. Manage. 67(1): 207-218. doi:10.2307/3803076
    • (2003) J. Wildl. Manage , vol.67 , Issue.1 , pp. 207-218
    • Jeffries, S.1    Huber, H.2    Calambokidis, J.3    Laake, J.4
  • 20
    • 11244355116 scopus 로고    scopus 로고
    • Stock-specific migration timing of adult spring-summer Chinook salmon in the Columbia River basin
    • doi:10.1577/M03-170.1
    • Keefer, M.L., Peery, C.A., Jepson, M.A., Tolotti, K.R., Bjornn, T.C., and Stuehrenberg, L.C. 2004a. Stock-specific migration timing of adult spring-summer Chinook salmon in the Columbia River basin. N. Am. J. Fish. Manage. 24(4): 1145-1162. doi:10.1577/ M03-170.1
    • (2004) N. Am. J. Fish. Manage , vol.24 , Issue.4
    • Keefer, M.L.1    Peery, C.A.2    Jepson, M.A.3    Tolotti, K.R.4    Bjornn, T.C.5    Stuehrenberg, L.C.6
  • 21
    • 1542473708 scopus 로고    scopus 로고
    • Regurgitation rates of intragastric radio transmitters by adult Chinook salmon and steelhead during upstream migration in the Columbia and Snake rivers
    • doi:10.1577/M02-128
    • Keefer, M.L., Peery, C.A., Ringe, R.R., and Bjornn, T.C. 2004b. Regurgitation rates of intragastric radio transmitters by adult Chinook salmon and steelhead during upstream migration in the Columbia and Snake rivers. N. Am. J. Fish. Manage. 24(1): 47-54. doi:10.1577/M02-128
    • (2004) N. Am. J. Fish. Manage , vol.24 , Issue.1 , pp. 47-54
    • Keefer, M.L.1    Peery, C.A.2    Ringe, R.R.3    Bjornn, T.C.4
  • 22
    • 22044442690 scopus 로고    scopus 로고
    • Escapement, harvest, and unknown loss of radio-tagged adult salmonids in the Columbia River - Snake River hydrosystem
    • doi:10.1139/f04-246
    • Keefer, M.L., Peery, C.A., Daigle, W.R., Jepson, M.A., Lee, S.R., Boggs, C.T., Tolotti, K.R., and Burke, B.J. 2005. Escapement, harvest, and unknown loss of radio-tagged adult salmonids in the Columbia River - Snake River hydrosystem. Can. J. Fish. Aquat. Sci. 62(4): 930-949. doi:10.1139/f04-246
    • (2005) Can. J. Fish. Aquat. Sci , vol.62 , Issue.4 , pp. 930-949
    • Keefer, M.L.1    Peery, C.A.2    Daigle, W.R.3    Jepson, M.A.4    Lee, S.R.5    Boggs, C.T.6    Tolotti, K.R.7    Burke, B.J.8
  • 23
    • 63249093340 scopus 로고    scopus 로고
    • Transporting juvenile salmonids around dams impairs adult migration
    • doi:10.1890/07-0710., 1.PMID:19263886
    • Keefer, M.L., Caudill, C.C., Peery, C.A., and Lee, S.R. 2008. Transporting juvenile salmonids around dams impairs adult migration. Ecol. Appl. 18(8): 1888-1900. doi:10.1890/07-0710.1.PMID:19263886
    • (2008) Ecol. Appl , vol.18 , Issue.8 , pp. 1888-1900
    • Keefer, M.L.1    Caudill, C.C.2    Peery, C.A.3    Lee, S.R.4
  • 24
    • 77955724514 scopus 로고    scopus 로고
    • Prespawn mortality in adult spring Chinook salmon outplanted above barrier dams
    • doi:10.1111/j.1600-0633.2010.00418.x
    • Keefer, M.L., Taylor, G.A., Garletts, D.F., Gauthier, G.A., Pierce, T. M., and Caudill, C.C. 2010. Prespawn mortality in adult spring Chinook salmon outplanted above barrier dams. Ecol. Freshw. Fish, 19(3): 361-372. doi:10.1111/j.1600-0633.2010.00418.x
    • (2010) Ecol. Freshw. Fish , vol.19 , Issue.3 , pp. 361-372
    • Keefer, M.L.1    Taylor, G.A.2    Garletts, D.F.3    Gauthier, G.A.4    Pierce, T.M.5    Caudill, C.C.6
  • 25
    • 0036324161 scopus 로고    scopus 로고
    • Pinniped diet composition: A comparison of estimation models
    • Laake, J.L., Browne, P., DeLong, R.L., and Huber, H.R. 2002. Pinniped diet composition: a comparison of estimation models. Fish Bull. 100: 434-447
    • (2002) Fish Bull , vol.100 , pp. 434-447
    • Laake, J.L.1    Browne, P.2    Delong, R.L.3    Huber, H.R.4
  • 26
    • 2942679759 scopus 로고    scopus 로고
    • Hooking mortality by anatomical location and its use is estimating mortality of spring Chinook salmon caught and released in a river sport fishery
    • doi:10.1577/M02-101.1
    • Lindsay, R.B., Schroeder, R.K., Kenaston, K.R., Toman, R.N., and Buckman, M.A. 2004. Hooking mortality by anatomical location and its use is estimating mortality of spring Chinook salmon caught and released in a river sport fishery. N. Am. J. Fish. Manage. 24(2): 367-378. doi:10.1577/M02-101.1
    • (2004) N. Am. J. Fish. Manage , vol.24 , Issue.2 , pp. 367-378
    • Lindsay, R.B.1    Schroeder, R.K.2    Kenaston, K.R.3    Toman, R.N.4    Buckman, M.A.5
  • 27
    • 0033322323 scopus 로고    scopus 로고
    • Size-dependent predation in piscivores: Interactions between predator foraging and prey avoidance abilities
    • doi:10.1139/cjfas-56-7-1285
    • Lundvall, D., Svanbck, R., Persson, L., and Byström, P. 1999. Size-dependent predation in piscivores: interactions between predator foraging and prey avoidance abilities. Can. J. Fish. Aquat. Sci. 56(7): 1285-1292. doi:10.1139/cjfas-56-7-1285
    • (1999) Can. J. Fish. Aquat. Sci , vol.56 , Issue.7 , pp. 1285-1292
    • Lundvall, D.1    Svanbck, R.2    Persson, L.3    Byström, P.4
  • 28
    • 0346269173 scopus 로고    scopus 로고
    • A large-scale, multispecies status, assessment: Anadromous salmonids in the Columbia River Basin
    • doi:10.1890/1051-0761(2003)13[964:ALMSAA]2.0.CO;2
    • McClure, M.M., Holmes, E.E., Sanderson, B.L., and Jordan, C.E. 2003. A large-scale, multispecies status, assessment: anadromous salmonids in the Columbia River Basin. Ecol. Appl. 13(4): 964-989. doi:10.1890/1051-0761(2003)13[964:ALMSAA]2.0.CO;2
    • (2003) Ecol. Appl , vol.13 , Issue.4 , pp. 964-989
    • McClure, M.M.1    Holmes, E.E.2    Sanderson, B.L.3    Jordan, C.E.4
  • 29
    • 33645813794 scopus 로고    scopus 로고
    • Functional and aggregative responses of harbour seals to changes in salmonid abundance
    • doi:10.1098/rspb.2005.3215. PMID:16555787
    • Middlemas, S.J., Barton, T.R., Armstrong, J.D., and Thompson, P. 2006. Functional and aggregative responses of harbour seals to changes in salmonid abundance. Proc. Biol. Sci. 273(1583): 193-198. doi:10.1098/rspb.2005.3215. PMID:16555787
    • (2006) Proc. Biol. Sci , vol.273 , Issue.1583 , pp. 193-198
    • Middlemas, S.J.1    Barton, T.R.2    Armstrong, J.D.3    Thompson, P.4
  • 31
    • 73649087141 scopus 로고    scopus 로고
    • Marine mammals; environmental assessment on preventing California sea lion foraging and predation on salmonids at Willamette Falls
    • National Marine Fisheries Service
    • National Marine Fisheries Service. 1998. Marine mammals; environmental assessment on preventing California sea lion foraging and predation on salmonids at Willamette Falls, Oregon. Fed. Regist. 63: 56-60
    • (1998) Oregon. Fed. Regist , vol.63 , pp. 56-60
  • 33
    • 0347283802 scopus 로고    scopus 로고
    • National Marine Fisheries Service, section 9.6.1.5.3, action Item 106, Seattle, Wash
    • National Marine Fisheries Service. 2000. Federal Columbia River Power System Biological Opinion (section 9.6.1.5.3, action Item 106). Seattle, Wash
    • (2000) Federal Columbia River Power System Biological Opinion
  • 35
    • 20644471169 scopus 로고    scopus 로고
    • Late-season mortality during migration of radio-tagged adult sockeye salmon (Oncorhynchus nerka) in the Columbia River
    • doi:10.1139/f04-147
    • Naughton, G.P., Caudill, C.C., Keefer, M.L., Bjornn, T.C., Stuehren- berg, L., and Peery, C.A. 2005. Late-season mortality during migration of radio-tagged adult sockeye salmon (Oncorhynchus nerka) in the Columbia River. Can. J. Fish. Aquat. Sci. 62(1): 30-47. doi:10.1139/f04-147
    • (2005) Can. J. Fish. Aquat. Sci , vol.62 , Issue.1 , pp. 30-47
    • Naughton, G.P.1    Caudill, C.C.2    Keefer, M.L.3    Bjornn, T.C.4    Stuehren-berg, L.5    Peery, C.A.6
  • 36
    • 84950439627 scopus 로고
    • Pacific salmon at the crossroads: Stocks at risk from California, Oregon, Idaho, and Washington
    • Nehlsen, W., Williams, J.E., and Lichatowich, J.A. 1991. Pacific salmon at the crossroads: stocks at risk from California, Oregon, Idaho, and Washington. Fisheries, 16: 4-21
    • (1991) Fisheries , vol.16 , pp. 4-21
    • Nehlsen, W.1    Williams, J.E.2    Lichatowich, J.A.3
  • 37
    • 1342288269 scopus 로고    scopus 로고
    • Density-dependent predation by brown bears (Ursus arctos) on sockeye salmon (Oncorhynchus nerka)
    • doi:10.1139/f03-045
    • Quinn, T.P., Gende, S.M., Ruggerone, G.T., and Rogers, D.E. 2003. Density-dependent predation by brown bears (Ursus arctos) on sockeye salmon (Oncorhynchus nerka). Can. J. Fish. Aquat. Sci. 60(5): 553-562. doi:10.1139/f03-045
    • (2003) Can. J. Fish. Aquat. Sci , vol.60 , Issue.5 , pp. 553-562
    • Quinn, T.P.1    Gende, S.M.2    Ruggerone, G.T.3    Rogers, D.E.4
  • 39
    • 0347531547 scopus 로고    scopus 로고
    • Influence of fishway placement on fallback of adult salmon at the Bonneville Dam on the Columbia River
    • doi:10.1577/M02-113
    • Reischel, T.S., and Bjornn, T.C. 2003. Influence of fishway placement on fallback of adult salmon at the Bonneville Dam on the Columbia River. N. Am. J. Fish. Manage. 23(4): 1215-1224. doi:10.1577/M02-113
    • (2003) N. Am. J. Fish. Manage , vol.23 , Issue.4 , pp. 1215-1224
    • Reischel, T.S.1    Bjornn, T.C.2
  • 40
    • 10844238066 scopus 로고
    • Numerical relations between abundance of predators and survival of prey
    • Ricker, W.E. 1952. Numerical relations between abundance of predators and survival of prey. Can. Fish. Cult. 13: 5-9
    • (1952) Can. Fish. Cult , vol.13 , pp. 5-9
    • Ricker, W.E.1
  • 41
    • 0021545820 scopus 로고
    • Abundances and feeding habits of pinnipeds in the Rogue River
    • doi:10.2307/3801787
    • Roffe, T.J., and Mate, B.R. 1984. Abundances and feeding habits of pinnipeds in the Rogue River, Oregon. J. Wildl. Manage. 48(4): 1262-1274. doi:10.2307/3801787
    • (1984) Oregon. J. Wildl. Manage , vol.48 , Issue.4 , pp. 1262-1274
    • Roffe, T.J.1    Mate, B.R.2
  • 42
    • 0004282518 scopus 로고    scopus 로고
    • SAS Institute Inc, SAS Institute Inc., Cary, N.C., USA
    • SAS Institute Inc. 2008. SAS/STAT user's guide version 9.2. SAS Institute Inc., Cary, N.C., USA
    • (2008) SAS/STAT User's Guide Version 9.2
  • 46
    • 42049100283 scopus 로고    scopus 로고
    • US Army Corps of Engineers, USACE, Portland, Ore
    • US Army Corps of Engineers. 2005. Annual fish passage report. USACE, Portland, Ore
    • (2005) Annual Fish Passage Report
  • 47
    • 0015807248 scopus 로고
    • Optimal fish cruising speed
    • doi:10.1038/245048a0
    • Weihs, D. 1973. Optimal fish cruising speed. Nature, 245(5419): 48-50. doi:10.1038/245048a0
    • (1973) Nature , vol.245 , Issue.5419 , pp. 48-50
    • Weihs, D.1
  • 48
    • 0021567480 scopus 로고
    • The ontogenetic niche and species interactions in size-structured populations
    • doi:10.1146/annurev.es.15.110184. 002141
    • Werner, E.E., and Gilliam, J.F. 1984. The ontogenetic niche and species interactions in size-structured populations. Annu. Rev. Ecol. Syst. 15(1): 393-425. doi:10.1146/annurev.es.15.110184. 002141
    • (1984) Annu. Rev. Ecol. Syst , vol.15 , Issue.1 , pp. 393-425
    • Werner, E.E.1    Gilliam, J.F.2
  • 49
    • 34247195096 scopus 로고    scopus 로고
    • Assessment of harbor seal predation on adult salmonids in a Pacific Northwest estuary
    • doi:10.1890/05-1941. PMID:17489243
    • Wright, B.E., Riemer, S.D., Brown, R.F., Ougzin, A.M., and Bucklin, K.A. 2007. Assessment of harbor seal predation on adult salmonids in a Pacific Northwest estuary. Ecol. Appl. 17(2): 338-351. doi:10.1890/05-1941. PMID:17489243
    • (2007) Ecol. Appl , vol.17 , Issue.2 , pp. 338-351
    • Wright, B.E.1    Riemer, S.D.2    Brown, R.F.3    Ougzin, A.M.4    Bucklin, K.A.5
  • 50
    • 73649117186 scopus 로고    scopus 로고
    • Movements of California sea lions captured in the Columbia River
    • doi:10.3955/046.084.0107
    • Wright, B.E., Tennis, M.J., and Brown, R.F. 2010. Movements of California sea lions captured in the Columbia River. Northwest Sci. 84(1): 60-72. doi:10.3955/046.084.0107
    • (2010) Northwest Sci , vol.84 , Issue.1 , pp. 60-72
    • Wright, B.E.1    Tennis, M.J.2    Brown, R.F.3


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