Late Jurassic Lungfishes (Dipnoi) from Uruguay, with comments on the systematics of Gondwanan Ceratodontiforms

Journal of Vertebrate Paleontology

Volumn 30, Issue 4, 2010, Pages 1049-1058

  Soto, Matías ^a   Perea, Daniel ^b  

^a Centro de Investigaciones Tecnológicas   (Uruguay)

^b Facultad de Ciencias   (Uruguay)

Author keywords

[No Author keywords available]

Indexed keywords

CRETACEOUS; FISH; FOSSIL ASSEMBLAGE; GONDWANA; JURASSIC; TAXONOMY;

BRAZIL; SUB-SAHARAN AFRICA; URUGUAY;

CERATODONTIFORMES; CHONDRICHTHYES; DIPNOI; NEOCERATODUS; SEMIONOTIDAE;

EID: 77956024429     PISSN: 02724634     EISSN: 19372809     Source Type: Journal    
DOI: 10.1080/02724634.2010.483540     Document Type: Article

References (109)

1. Agassiz, L. 1838. Recherches sur les Poissons Fossiles, Vol.3. Imprimerie de Petit-Pierre Neuchâtel
2. Ameghino, F. 1898. Sinopsis geol ógico-paleontol ógica de la Argentina; pp. 115-228 in Segundo Censo Nacional de la República Argentina, Territorio 1.
3. Apesteguía, S., F. L. Agnolín, and K. Claeson. 2007. Review of Cretaceous dipnoans from Argentina (Sarcopterygii: Dipnoi) with description of new species. Revista del Museo Argentino de Ciencias Naturales 9:27-40.
4. Arambourg, C., and L. Joleaud. 1943. Vertébrés fossiles du bassin du Niger. Bulletin de la Service des Mines, Gouverment Général de lAfrique Occidentale Française 7:27-84.
5. Arratia, G., and A. Cione. 1996. The Record of Fossil Fishes of Southern South America. Münchner Geowissenschaftliche Abhandlungen 30:9-72.
6. Berg, L. 1940. Classification of fishes, both recent and fossil. Travaux de lInstitut de Zoologie, Academie des Sciences de LURSS 5: 85-517.
7. Bossi, J. 1966. Geología del Uruguay. Departamento de Publicaciones de la Universidad de la República, Montevideo, 469 pp.
8. Bossi, J., L. A. Ferrando, A. Fernández, G. Elizalde, H. Morales, J. Ledesma, E. Carballo, E. Medina, I. Ford, and J. R. Montaña (eds.). 1975. Carta geol ógica del Uruguay. Escala 1/1.000.000. Montevideo, 32 pp.
9. Bouaziz, S., E. Buffetaut, M. Ghanmi, J.-J. Jaeger, M. Martin, J.-M. Mazin, and H. Tong. 1988. Nouvelles découvertes de vert ébrés fossils dan lAlbien du Sud Tunisien. Bulletin de la Sociét é géologique de France 4:335-339.
10. Cappetta, H. 1986. Types dentaires adaptatifs chez les s élaciens actuels etpost-pal éozöiques. Palaeovertebrata 16:57-76.
11. Cappetta, H., E. Buffetaut, and V. Suteethorn. 1990. A new hybodont shark from the Lower Cretaceous of Thailand. Neues Jarbuch f ür Geologie und Paläontologie Monatshefte H 11:659-666.
12. Case, E. C. 1921. A new species of Ceratodus from the Upper Triassic of western Texas. Ocassional Papers of the Museum of Zoology, University of Michigan 101:1-2.
13. Castro, D. F., C. E. V. Toledo, E. P. Sousa, and M. A. Medeiros. 2004. Nova ocorrência de Asiatoceratodus (Osteichthyes, Dipnoiformes) na Formaç ão Alcântara, Eocenomaniano da Bacia de São Luís, MA, Brasil. Revista Brasileira de Paleontologia 7:245-248.
14. Cavin, L., V. Suteethorn, E. Buffetaut, and H. Tong. 2007. A new Thai Mesozoic lungfish (Sarcopterygii, Dipnoi) with an insight into post- Palaeozoic dipnoan evolution. Zoological Journal of the Linnean Society 149:141-177.
15. Cavin, L., L. Boudadb, S. Duffauda, K. Lahcen, J. Le Loeuffa, I. Rougetc, and H. Tonga. 2001. Lévolution pal éoenvironnementale des faunes de poissons du Crétacé supérieur du bassin du Tafilaltet des r égions avoisinantes (Sud-Est du Maroc): implications paléobiogéographiques. C. R. Acad. Sci. Paris, Sciences de la Terre et des planètes 333:677-683.
16. Churcher, C. S. and G. De Iuliis. 2001.A new species of Protopterus and a revision of Ceratodus humei (Dipnoi: eratodontiformes) from the Late Cretaceous Mut Formation of Eastern Dakhleh Oasis, Western Desert if Egypt. Paleontology 44:305-323.
17. Churcher, C. S, G. De Iuliis, and M. R. Kleindienst. 2006. A new genus for the dipnoan species Ceratodus tuberculatus Tabaste, 1963. Geodiversitas 28:635-647.
18. Cione, A. L., S. Gouiric, F. Goin, and D. Poiré. 2007. Atlantoceratodus, a new genus of lungfish from the upper Cretaceous of South America and Africa. Revista del Museo de la Plata, Paleontología 10:1-12.
19. Cunha, F. L., and C. S. Ferreira. 1980. Um dipnoi na Formaç ão Itapecuru (Cenomaniano), Maranhão, Brasil. II Congreso Argentino de Paleontolog ía y Bioestratigrafía and I Congreso Latinoamericano de Paleontología, Buenos Aires, Actas 1:1-9.
20. Cuny, G., L. Cavin, and V. Suteethorn. 2009. A new hybodont with a cutting dentition from the Lower Cretaceous of Thailand. Cretaceous Research 30:515-520.
21. Cuny, G., V. Suteethorn, S. Kamha, and E. Buffetaut. 2008. Hybodont sharks from the Lower Cretaceous Khok Kruat Formation of Thailand, and hybodont diversity during the Early Cretaceous; pp. 93-107 in L. Cavin, A. Longbottom, and M. Richter (eds.), Fishes and the Break-Up of Pangaea. Geological Society, London, Special Publications 295.
22. Cuny, G.,M. Ouaja, D. Srarfi, L. Schmitz, E. Buffetaut, andM. J. Benton, 2004. Fossil sharks from the Early Cretaceous of Tunisia. Revue de Paléobiologie 9:127-142.
23. Da Silva, J. S. 1990. Paleontología de la Formaci ón Tacuaremb ó (Uruguay). Avance 1989. I Congreso Uruguayo de Geología, Montevideo, Actas (2):27-31.
24. Duffin, C. J. 2001. The hybodont shark, Priohybodus dErasmo, 1960 (Early Cretaceous, northern Africa). Zoological Journal of the Linnean Society 133:303-308.
25. Duffin, C. J. and G. Cuny. 2008. Carcharopsis prototypus and the adaptations of single crystallite enameloid in cutting dentitions. Acta Geologica Polonica 58:181-184.
26. Dutra, M. F. A., and M. C. S. L. Malabarba. 2001. Peixes do Albiano- Cenomaniano do Grupo Itapecuru no Estado do Maranhão, Brasil;pp. 191-208 in D. F. Rossetti, A. M. Góes, and W. Truckenbrodt (eds.), O Cretáceo na Bacia de São Luís-Graja ú, MPGE.
27. Gasparini, Z. 1996. Biogeograpic evolution of the South American crocodilians; pp. 159-184 in G. Arratia (ed.), Contributions of Southern South America to Vertebrate Paleontology. Münchner Geowissenschaftliche Abhandlungen (A) 30.
28. Gayet, M., and P. M. Brito. 1989. Ichthyofaune nouvelle du Crétacé Supérieur du Groupe Baur ú (etats de São Paulo et Minas Gerais, Brésil). Géobios 22:841-847.
29. Gill, T. 1872. On the homologies of the shoulder girdle of the dipnoans and other fishes. Annals and Magazine of Natural History 11:173-178.
30. Goodwin, M. B., W. A. Clemens, H. J. Hutchison, C B. Wood, M. S. Zavada, A. Kemp, C. J. Duffin, and C. R. Schaff. 1999. First Mesozoic terrestrial vertebrates with associated palynostratigraphic dates from the Northwestern Ethiopian Plateau. Journal of Vertebrate Paleontology 19:728-741.
31. Haug, E. 1905. Paléontologie; pp. 751-832 in F. Foureau. Documents scientifiques de la Mission Foureau-Lamy dAlger au Congo par le Tchad. Publication de la Sociét é de Géographie, Paris.
32. Holtz, T. R., Jr., R. E. Chapman, and M. C. Lamanna. 2004. Mesozoic biogeography of Dinosauria; pp. 627-642 in D. B. Weishampel, P.Dodson, and H. Osmólska (eds.), The Dinosauria, second edition. University of California Press, Berkeley, California, Los Angeles, London.
33. Jain, S. L. 1983. A review of the genus Lepidotes (Actinopterygii: Semionotiformes) with special reference to the species from the Kota Formation (Lower Jurassic), India. Journal of the Palaeontological Society of India 28:7-42.
34. Kemp, A. 1991. Australian Mesozoic and Cainozoic lungfish; pp. 465-498 in P. Vickers-Rich, J. M. Monaghan, R. F. Baird, and T. Rich (eds.), Vertebrate Paleontology of Australasia. Melbourne, Pioneer DesignStudio
35. Kemp, A. 1993. Ceratodus diutinus, a new ceratodont from Cretaceous and Late Oligocene-Medial Miocene deposits in Australia. Journal of Paleontology 67:883-888.
36. Kemp, A. 1994. Australian Triassic lungfish skulls. Journal of Paleontology 68:647-654.
37. Kemp, A. 1996. Sagenodus (Ceratodus) carlinvillensis (Romer and Smith 1934), (Osteichthyes, Dipnoi), short ridge anomaly and classification of dipnoans. Journal of Vertebrate Paleontology 16:16-19.
38. Kemp, A. 1997a. Four species of Metaceratodus (Osteichthyes: Dipnoi: family Ceratodontidae) from Australian Mesozoic and Cenozoic deposits. Journal of Vertebrate Paleontology 17:26-33.
39. Kemp, A. 1997b. A revision of Australian Mesozoic and Cenozoic lungfish of the family Neoceratodontidae (Osteichthyes: Dipnoi), with a description of four new species. Journal of Paleontology 71: 713-733.
40. Kemp, A. 1998. Skull structure in post-Paleozoic lungfish. Journal of Vertebrate Paleontology 18:43-63.
41. Kemp, A. 2001a. Consequences of traumatic injury in fossil and recent dipnoan dentitions. Journal of Vertebrate Paleontology 21:13-23.
42. Kemp, A. 2001b. Petrodentine in derived dipnoan dentitions. Journal of Vertebrate Paleontology 21:422-437.
43. Kemp, A. 2002. Unique dentition of lungfish. Microscopical Research Techniques 59:435-448.
44. Kemp, A. 2003. Developmental anomalies in the tooth plates and jaw bones of lungfish. Journal of Vertebrate Paleontology 23:517-531.
45. Kemp, A., and R. E. Molnar. 1981. Neoceratodus forsteri from the Lower Cretaceous of New South Wales, Australia. Journal of Paleontology 55:211-217.
46. Kirkland, J. I. 1987. Upper Jurassic and Cretaceous lungfish tooth plates from the Western Interior, the last dipnoan faunas of North America. Hunteria 2(2), 14 pp.
47. Lehman, 1971. Nouveaux Vertébrés fossiles du Trias de la s érie de Zarzäitine. Annales de Paléontologie, Paris 57:71-92.
48. Long, J. A. 1995. The Rise of Fishes. 500 Million Years of Evolution. The Johns Hopkins University Press, Baltimore and London, 223 pp.
49. López-Arbarello, A. 2004. The record of Mesozoic fishes from Gondwana (excluding India and Madagascar); pp. 597-612 in G. Arratia and A. Tintori (eds.), Mesozoic Fishes 3-Systematics, Paleoenvironments and Biodiversity. Verlag Dr. Friedrich Pfeil München.
50. López-Arbarello, A., O. W. M. Rauhut, and K. Möser. 2008. Jurassic fishes of Gondwana. Revista de la Asociaci ón Geol ógica Argentina 63:586-612.
51. Maisey, J. G. 1983. Cranial Anatomy of Hybodus basanus Egerton from the Lower Cretaceous of England. American Museum Novitates 2758:1-64.
52. Maisey, J. G. 1989. Hamiltonichthys mapesi, g. & sp. nov. (Chondrichthyes: Elasmobranchii), from the Upper Pennsylvanian of Kansas. American Museum Novitates 2931:1-42.
53. Maisey, J. G. 2000. Continental break-up and the distribution of fishes of Western Gondwana during the Early Cretaceous. Cretaceous Research 21:281-314.
54. Martin, M. 1979a. Arganodus atlantis et Ceratodus arganensis, deux nouveaux Dipneustes du Trias supérieur continental marocain. Comptes Rendus de lAcadémie des Sciences (Paris) D 289:89-92.
55. Martin, M. 1979b. Les Dipneustes et Actinistiens du Trias supérieur continental marocain. Stuttgarter Beiträge zur Naturkunde Ser. B 69:1-29.
56. Martin, M. 1981a. Les Ceratodontiformes (Dipnoi) de Gadoufaoua (Aptien supérieur, Niger). Bulletin du Musée national dHistoire naturelle (Paris), 4e s ér. 3(C):267-282.
57. Martin, M. 1981b. Les Dipneustes mésozoiques malgaches, leurs affinit és et leur int érêt paléobiogé ographiques. Bulletin de la Sociét é géologique de France XXVIII 6:579-585.
58. Martin, M. 1982a. Nouvelles données sur la phylogénie et la syst ématique des Dipneustes postpaléozoiques. Comptes Rendus de lAcadémie des Sciences (Paris), s ér. II 294:611-614.
59. Martin, M. 1982b. Nouvelles données sur la phylogénie et la syst ématique des Dipneustes postpaléozoiques, consequences stratigraphiques et paléogéographiques. Geobios mémoire spécial 6:53-64.
60. Martin, M. 1984a. Révision des Arganodontidés et des Néocératodontid és (Dipnoi, Ceratodontiformes) du Crétacé africain. Neues Jarbuch f ür Geologie und Paläontologie Abhandlungen 169:225-260.
61. Martin M. 1984b. Deux Lepidosirenidae (Dipnoi) du Cretace du Sahara Protopterus humei (Priem) et Protopterus protopteroides (Tabaste). Paläontologische Zeitschrift 58:265-277.
62. Martin, M. 1997. Protopterus nigeriensis nov. sp., lun des plus anciens protoptères-DIPNOI-(In Beceten, Sénonien du Niger). Comptes Rendus de lAcadémie des Sciences (Paris) 325:635-638.
63. Martin, M., L. Barbieri, and G. Cuny. 1999. The Madagascan Mesozoic ptychoceratodontids (Dipnoi): systematic relationships and paleobiogeographical significance. Oryctos 2:3-16.
64. Martin, M., E. Buffetaut, H. Tong, and V. Suteethorn. 1996. New Jurassic Dipnoans from Thailand. Geological Society of Denmark Online Series 1. Available at
65. Martin, M., D. Sigogneau-Russel, P. Coupatez, and G. Wouters. 1981. Les cératodontidés (Dipnoi) du Rhétien de Saint-Nicolas-de-Port (Meurthe et Moselle). Géobios 14:773-791.
66. McCune, A. R. 1986. A revision of Semionotus (Pisces, Semionotidae) from the Triassic and Jurassic of Europe. Palaeontology 29:213-233.
67. Medeiros, M. A., and C. L. Schultz. 2001. Uma paleocomunidade de vertebrados do Cretáceo médio, bacia de São Luís; pp. 209-221 in D. F. Rossetti, A. M. Góes, and W. Truckenbrodt (eds.), O Cretáceo na bacia de São Luís. Graja ú, Museu Paraense Emílio Goeldi
68. Medeiros, M. A., P. C. Freire, A. A. Pereira, R. A. B. Santos, R. M. Lindoso, A. F. A. Coelho, E. B. Passos, and E. S. J únior. 2007. Another African dinosaur recorded in the Eocenomanian of Brazil and a revision on the paleofauna of the Laje do Coringa; pp. 413-423 in I. S. Carvalho, R. C. T. Cassab, C. Schwanke, M. A. Carvalho, A. C. S. Fernandes, M. A. C. Rodrigues, M. S. S. de Carvalho, M. Arai, andM. E. Q. Oliveira (eds.), Paleontologia: Cenários de Vida, Vol.1. Rio de Janeiro, Interci ência.
69. Mones, Á . 1997. Los vertebrados mesozoicos del Uruguay y sus relaciones con los de áreas vecinas; pp. 205-222 in J. Arroyo Cabrales and O. J. Polaco (coords.), Homenaje al Profesor Ticul Á lvarez. Instituto Nacional de Antropología e Historia, México, Colecci ón Científica 357.
70. Moody, R. T. J., and P. J. C. Sutcliffe. 1991. The Cretaceous deposits of the Iullemmeden Basin of Niger, central West Africa. Cretaceous Research 12:137-157.
71. Mü ller, J. 1845. Ü ber den Bau und die Grenzen der ganoiden und über das nat ürliche System der Fische. Abhandlungen der Akademie der Wissenschaften zu Berlin 1844:117-216.
72. Murray, A. M. 2000. The Palaeozoic, Mesozoic and Early Cenozoic fishes of Africa. Fish and Fisheries 1:111-145.
73. Pascual, R., and P. Bondesio. 1976. Notas sobre vertebrados de la frontera Cretácico-Terciaria. III: Ceratodontidae (Peces, Osteichthyes, Dipnoi) de la Formaci ón Coli Toro y de otras unidades del Cretácico Tardío de Patagonia y sur de Mendoza. Sus implicancias paleobiogeográficas. VI Congreso Geol ógico Argentino, Actas 1: 565-577.
74. Patterson, C. 1975. The distribution of Mesozoic freshwater fishes. Mémoires du Muséum National dHistoire Naturelle 88:156-173.
75. Pause, M. 1979. Ü ber eine fossile Schildkrö te (Podocnemis) aus der mittleren Kreide der südlichen Kharga-Oasen (SW-Ä gypten). Unpublished Diplom-Arbeit, Institut f ür Geologie und Paläontologie der Technischen Universität, Berlin, 42 pp.
76. Perea, D., M. Ubilla, A. Rojas, and C. Goso. 2001. The West Gondwanan ocurrence of the hybodontid shark Priohybodus and the Late Jurassic-Early Cretaceous age of Tacuaremb ó Formation, Uruguay. Palaeontology 44:1227-1235.
77. Perea, D., M. Soto, G. Veroslavsky, S. Martínez, and M. Ubilla, 2009. A Late Jurassic assemblage in Gondwana: biostratigraphy and correlations of Tacuaremb ó Formation, Paraná Basin, Uruguay. Journal of South American Earth Sciences 28:168-179.
78. Peyer, B. 1925. 6. Die Ceratodus-Funde. II. Wirbeltier-Reste der Bahar îje-Stufe (unterstes Cenoman). Ergebnisse der Forschungsreisen Prof. E. Stromer in den Wü sten Ä gyptens. Abhandlungen der Bayerischen Akademie der Wissenschaften, athematischnaturwissenschaftliche 30:5-32.
79. Rauhut, O. W. M., and A. López-Arbarello. 2009. Considerations on the age of the Tiouraen Formation (Iullemmeden Basin, Niger, Africa): implications for Gondwanan Mesozoic terrestrial vertebrate faunas. Palaeogeography Palaeoclimatology Palaeocology 271:259-267.
80. Rees, J. 1998. Early Jurassic selachians from the Hasle Formation on Bornholm, Denmark. Acta Palaeontologica Polonica 43:439-452.
81. Rees, J., and C. J. Underwood. 2008. Hybodont sharks of the English Bathonian and Callovian (Middle Jurassic). Palaeontology 51:117-147.
82. Reif, W.-E. 1988. Evolution of high swimming velocities in sharks-a case of escalation? Neues Jarbuch f ür Geologie und Paläontologie Monatshefte 6:361-379.
83. Richter, M., and C. E. V. Toledo. 2008. The first Triassic lungfish from South America (Santa Maria Formation, Paraná Basin) and its bearing on geological correlations within Pangaea; pp. 43-54 in L. Cavin, A. Longbottom, and M. Richter, M. (eds), Fishes and the Breakup of Pangaea. Geological Society, London, Special Publications 295.
84. Santos, M. E. C., and M. S. S. Carvalho. 2009. Reconstituçoes Paleobiol ógicas. CPRM, Rio de Janeiro, 215 pp.
85. Schultze, H.-P. 1981. A dipnoan tooth plate from the Lower Cretaceous of Kansas, U.S.A. Transactions of the Kansas Academy of Sciences 84:187-195.
86. Schultze, H.-P. 1992a. Fossilium Catalogus: Animalia, Pars 131, Dipnoi. Kugler Publications, Amsterdam, New York, 464 pp.
87. Schultze, H.-P. 1992b. Lungfish from the El Molino (Late Cretaceous) and Santa Lucia formations in South-central Bolivia; pp. 441-448 in R. Suárez Soruco (ed.), Fósiles y Facies de Bolivia, Vol.I. Vertebrados. Revista Técnica de YPFB I:441-448.
88. Schultze, H.-P. 2004. Mesozoic sarcopterygians; pp. 463-492 in G. Arratia and A. Tintori (eds.), Mesozoic Fishes 3-Systematics, Paleoenvironments and Biodiversity. Verlag Dr. Friedrich Pfeil, München.
89. Schultze, H.-P., and C. R. Marshall. 1993. Contrasting the use of functional complexes and isolated characters in lungfish evolution. Memoirs of the Association of Australasian Palaeontologists 15: 211-224.
90. Sereno, P. C., J. A. Wilson, and J. L. Conrad. 2004. New dinosaurs link southern landmasses in the Mid-Cretaceous. Proceedings of the Royal Society of London B 271:1325-1330.
91. Sereno, P. C., J. A. Wilson, H. C. E. Larsson, D. B. D. Dutheil, and H.-D. Sues. 1994. Early Cretaceous dinosaurs from the Sahara. Science 266:267-271.
92. Shen, Y. B., O. F. Gallego, and S. Martínez. 2004. The conchostracan subgenus Orthestheria (Migransia) from the Tacuaremb ó Formation (Late Jurassic-?Early Cretaceous, Uruguay) with notes on its geological age. Journal of South American Earth Sciences 16:631-638.
93. Silva, V. G., and S.A. de Azevedo. 1992.Umdipn óico da Formaç ão Brejo Santo, Jurássico da Chapada do Araripe, Ceará, Brasil. Anais da Academia Brasileira de Ciências 64:419-420.
94. Silva, V. G., and S. A. Azevedo. 1996. Um dipn óico da Formaç ão Brejo Santo, Eocretáceo da Chapada do Araripe, Ceará, Brasil. Acta Geol ógica Leopoldensia 43:43-58.
95. Soto, M. 2004. Los vertebrados de la Formaci ón Tacuaremb ó: implicancias geocronol ógicas, paleobiogeográficas y paleoecol ógicas. Bachelor thesis, Facultad de Ciencias, Universidad de la República, Montevideo, 105 pp.
96. Soto, M. 2005. La especializada dentici ón de Priohybodus (Chondrichthyes, Hybodontidae); p. 108 in VIII Jornadas de Zoología del Uruguay, Montevideo, Actas.
97. Soto, M., and D. Perea. 2008. A ceratosaurid (Dinosauria, Theropoda) from the Late Jurassic-Early Cretaceous of Uruguay. Journal of Vertebrate Paleontology 28:439-444.
98. Stromer, E. 1914. Die Topographie und Geologie der Streke Gharag- Baharîje nebst Ausführungen über die geologische Ä gyptens. Ergebnisse der Forschungsreisen Professor E. Stromer in den Wü sten Ä gyptens. Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathematisch-naturwissenschaftliche Abteilung 11:1-78.
99. Tabaste, N. 1963. É tude des restes de Poissons du CrÉ tacÉ Saharien. Memoires de lInstitut Français dAfrique Noire 68:437-486.
100. Taquet, P. 1976. Géologie et Paléontologie du gisement de Gadoufaoua (Aptien du Niger). Cahiers de Paléontologie 1-191.
101. Thomson, K. S. 1990. Sarcopterygii: Dipnoi; pp. 29-33 in M. T. Antunes, J. G. Maisey, M. M. Marques, B. Schaeffer, and K. S. Thomson (eds.), Triassic Fishes from the Cassange Depression (R. P. de Angola). Ciencias de la Terra, n úmero especial, Lisboa.
102. Toledo, C. E. V., and R. J. Bertini. 2005. Occurrences of the fossil dipnoiformes in Brazil and its stratigraphic and chronological distributions. Revista Brasileira de Paleontologia 8:47-56.
103. Toledo, C. E. V., R. J. Bertini, E. P. Sousa, and M. A. Medeiros. 2005. First occurrence of Protopterus humei (Dipnoiformes) in South America; pp 262-263 in II Congresso Latinoamericano de Paleontologia de Vertebrados, Rio de Janeiro, Boletim de Resumos.
104. Schmidt, D., and C. Werner. 1998. Early Cretaceous coastal plain sediments of the Mugher Mudstone Formation, Abay River Basin, Ethiopia. Zeitblatt, Geologie und Paläontologie, Teil I(1-2):293-309.
105. Vorobyeva, E. I. 1967. Triassic ceratod from South Fergana and remarks on the systematic and phylogeny of ceratodontids. Palaeontologisk Zhurnal 1967:102-111. [Russian. Translated in Paleontological Journal 1967:80-87.]
106. Vorobyeva, E. I., and M. C. Minikh. 1968. Experimental application of biometry to the study of ceratodontid dental plates. Palaeontologisk Zhurnal 1968:76-87.[Russian. Translated in Paleontological Journal 1968:217-222.]
107. Wenz, S. 1980. A propos du genre Mawsonia, Coelacanthe géant du Crétacé inférieur dAfrique et du Brésil. Mémories de la Sociét é géologique de France 39:187-190.
108. Werner, C. 1993. Eine neue Fundstelle terrestricher Wirbeltiere aus der Kreide des Sudan. Berliner geowissenschaftliche Abhandlungen E 13:221-249.
109. Werner, C. 1995. Neue Funde von mesozoichen Wirbeltieren in Aethiopien. Berliner Geowissenschaftliche Abhandlungen E 16:377-383.