Re-assessment of varanid evolution based on new data from Saniwa ensidens leidy, 1870 (Squamata, reptilia)

American Museum Novitates

Volumn , Issue 3630, 2008, Pages 1-15

  Conrad, Jack L ^a   Rieppel, Olivier ^b   Grande, Lance ^b  

^a DIVISION OF INVERTEBRATE ZOOLOGY   (United States)

^b FIELD MUSEUM OF NATURAL HISTORY   (United States)

Author keywords

[No Author keywords available]

Indexed keywords

EID: 59349103969     PISSN: 00030082     EISSN: None     Source Type: Book Series    
DOI: 10.1206/596.1     Document Type: Review

References (73)

1. Ast, J.C. 2001. Mitochondrial DNA evidence and evolution in Varanoidea (Squamata). Cladistics 17:211-226.
2. Ast, J.C. 2002a. Erratum. Cladistics 18: 125.
3. Ast, J.C. 2002b. Evolution in Squamata (Reptilia). Ph.D. dissertation, University of Michigan, 276 pp.
4. Bellairs, A. 1949. Observations on the snout of Varanus, and a comparison with that of other lizards and snakes. Journal of Anatomy 83: 116-146.
5. Bernstein, P. 1999. Morphology of the nasal capsule of Heloderma suspectum with comments on the systematic position of helodermatids (Squmata: Helodermatidae). Acta Zoológica (Stockholm) 80: 219-230.
6. Borsuk-Bialynicka, M. 1983. The early phylogeny of Anguimorpha as implicated by craniological data. Acta Paleontologica Polonica 28: 5-105.
7. Borsuk-Bialynicka, M. 1984. Anguimorphans and related lizards from the Late Cretaceous of the Gobi Desert, Mongolia. Palaeontologia Polonica 46: 5-105.
8. Branch, W.R. 1982. Hemipeneal morphology of platynotan lizards. Journal of Herpetology 16: 16-38.
9. Caldwell, M.W. 1996. Ontogeny and phylogeny of the mesopodial skeleton in mosasauroid reptiles. Zoological Journal of the Linnean Society 116: 407-436.
10. Caldwell, M.W. 1999. Squamate phylogeny and the relationships of snakes and mosasauroids. Zoological Journal of the Linnean Society 125: 115-147.
11. Caldwell, M.W. 2000. On the aquatic squamate Dolichosaurus longicollis Owen, 1850 (Cenomanian, Upper Cretaceous), and the evolution of elongate necks in squamates. Journal of Vertebrate Paleontology 20: 720-735.
12. Caldwell, M.W. 2003. Holotype snout elements of Saniwa ensidens reassigned to cf. Restes sp. indet. (Xenosauridae). Journal of Paleontology 77: 393-396.
13. Camp, CL. 1923. Classification of the lizards. Bulletin of the American Museum of Natural History 48: 289-481.
14. Clos, L.M. 1995. A new species of Varanus (Reptilia, Sauria) from the Miocene of Kenya. Journal of Vertebrate Paleontology 15: 254-267.
15. Conrad, J.L. 2004. Skull, mandible, and hyoid of Shinisaurus crocodilurus AhI (Squamata, Anguimorpha). Zoological Journal of the Linnean Society 141: 399-434.
16. Conrad, J.L. 2008. Phylogeny and systematics of Squamata (Reptilia) based on morphology. Bulletin of the American Museum of Natural History 310: 1-182.
17. Dal Sasso, C., and G. Pinna. 1997. Aphanizocnemus libanensis n. gen. n. sp., a new dolichosaur (Reptilia, Varanoidea) from the Upper Cretaceous of Lebanon. Paleontologia Lombarda 7: 1-31.
18. Erickson, G.M., A. de Ricqles, V. de Buffrénil, R.E. Molnar, and M.K. Bayless. 2003. Vermiform bones and the evolution of giggantism in Megalania - how a reptilian fox became a lion. Journal of Vertebrate Paleontology 23: 966-970.
19. Estes, R. 1983. Sauria terrestria, Amphisbaenia. New York: Gustav Fischer Verlag, 249 pp.
20. Estes, R., K. de Queiroz, and J. Gauthier. 1988. Phylogenetic relationships within Squamata. In R. Estes and G. Pregill (editors), Phylogenetic relationships of the lizard families, 119-281. Stanford: Stanford University Press.
21. Evans, S.E., and L.J. Barbadillo. 1998. An unusual lizard (Reptilia: Squamata) from the Early Cretaceous of Las Hoyas, Spain. Zoological Journal of the Linnean Society 124: 235-265.
22. Evans, S.E., and L.J. Barbadillo. 1999. A short-limbed lizard from the Lower Cretaceous of Spain. Special Papers in Palaeontology Series 60: 73-85.
23. Evans, S.E., Y. Wang, and C. Li. 2005. The early Cretaceous lizard genus Yabeinosaurus from China: resolving an enigma. Journal of Systematic Palaeontology 4: 319-335.
24. Fitch, A.J., A.E. Goodman, and S.C. Donnellan. 2006. A molecular phylogeny of the Australian monitor lizards (Squamata: Varanidae) inferred from mitochondrial DNA sequences. Australian Journal of Zoology 54: 253-269.
25. Fuller, S., P. Baverstock, and D. King. 1998. Biogeographic origins of goannas (Varanidae): a molecular perspective. Molecular Phylogenetics and Evolution 9: 294-307.
26. Gao, K.-Q., and L. Hou. 1996. Systematics and taxonomie diversity of squamates from the Upper Cretaceous Djadochta Formation, Bayan Manahu, Gobi Desert, People's Republic of China. Canadian Journal of Earth Sciences 33: 578-598.
27. Gao, K.-Q., and M.A. Norell. 1998. Taxonomic revision of Carusia (Reptilia: Squamata) from the Late Cretaceous of the Gobi Desert and phylogenetic relationships of anguimorphan lizards. American Museum Novitates 3230: 1-51.
28. Gao, K.-Q., and M.A. Norell. 2000. Taxonomic composition and systematics of Late Cretaceous lizard assemblages from Ukhaa Tolgod and adjacent localities, Mongolian Gobi Desert. Bulletin of the American Museum of Natural History 249: 1-118.
29. Gilmore, C.W. 1928. Fossil lizards of North America. Memoirs of the National Academy of Sciences 22: 1-197.
30. Gilmore, C.W. 1943. Fossil lizards of Mongolia. Bulletin of the American Museum of Natural History 81: 361-384.
31. Goloboff, P.A., J.S. Farris, and K.C. Nixon. 2003. T.N.T.: Tree Analysis Using New Technology, v. 1.0. Goloboff, Farris, and Nixon (www.zmuc.dk/public/ phylogeny).
32. Grande, L., and H.P. Buchheim. 1994. Paleontological and sedimentological variation in Early Eocene Fossil Lake. Contributions to Geology, University of Wyoming 30: 33-56.
33. Hecht, M.K. 1975. The morphology and relationships of the largest known terrestrial lizard, Megalania prisca Owen, from the Pleistocene of Australia. Proceedings of the Royal Society of Victoria 87: 239-249.
34. Hoffstetter, R. 1969. Présence de Varanidae (Reptilia, Sauria) dans le Miocène de Catalogne. Considerations sur l'histoire de la famille. Bulletin du Muséum National d'Histoire Naturelle, 2e série 40: 1051-1064.
35. Jenkins, F., and G. Goslow. 1983. The functional anatomy of the shoulder of the savannah monitor lizard (Varanus exanthematics). Journal of Morphology 175: 195-216.
36. King, D., S. Fuller, and P. Baverstock. 1999. The biogeographic origins of varanid lizards. In H.-G. Horn and W. Böhme (editors), Mertensiella: advances in monitor research II, 43-49. Rhine Brook: Deutesche Gesellschaft für Herpetologie und Terrarienkunde.
37. Landsmeer, J.M.F. 1983. The mechanism of forearm rotation in Varanus exanthematicus. Journal of Morphology 175: 119-130.
38. Lee, M.S.Y. 1995. Possible affinities between Varanus giganteus and Megalania prisca. Memoirs of the Queensland Museum 39: 232.
39. Lee, M.S.Y. 1997. The phylogeny of varanoid lizards and the affinities of snakes. Philosophical Transactions of the Royal Society of London, Series B: Biological Sciences 352: 53-91.
40. Lee, M.S.Y. 1998. Convergent evolution and character correlation in burrowing reptiles: towards a resolution of squamate relationships. Biological Journal of the Linnean Society 65: 369-453.
41. Lee, M.S.Y., and M.W. Caldwell. 2000. Adriosaurus and the affinities of mosasaurs, dolichosaurs, and snakes. Journal of Paleontology 74: 915-937.
42. Leidy, J. 1870. (Descriptions of Emys jeansesi, E. haydeni, Baëna arenosa, and Saniwa ensidens). Proceedings of the Academy of Natural Sciences of Philadelphia 1870: 123-124.
43. Maisano, J.A. 2001a. Lanthanotus borneensis (http://digimorph.org/ specimens/Lanthanotus-borneensis). Digital Morphology, Austin. Accessed 2006.
44. Maisano, J.A. 2001b. Varanus gouldii. (http://digimorph.org/ specimens/Varanus-gouldii). Digital Morphology, Austin. Accessed 2007.
45. Maisano, J.A., C.J. Bell, J. Gauthier, and T. Rowe. 2002. The osteoderms and palpebral in Lanthanotus borneensis (Squamata: Anguimorpha). Journal of Herpetology 36: 678-682.
46. McDowell, S.B., Jr., and C.M. Bogert. 1954. The systematic position of Lanthanotus and the affinities of the anguinomorphan lizards. Bulletin of the American Museum of Natural History 105: 1-142.
47. Mertens, R. 1942a. Die Familie der Warane (Varanidae). Dritter Teil: Taxonomie. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 466: 235-391.
48. Mertens, R. 1942b. Die Familie der Warane (Varanidae). Erster Teil: Allgemeines. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 462: 1-116.
49. Mertens, R. 1942c. Die Familie der Warane (Varanidae). Zweiter Teil: Der Schädel. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 465: 117-234.
50. Molnar, R.E. 1990. New cranial elements of a giant varanid from Queensland. Memoirs of the Queensland Museum 29: 437-444.
51. Molnar, R.E. 2004. Dragons in the dust: the paleobiology of the giant monitor lizard Megalania. Bloomington: Indiana University Press, 211 pp.
52. Norell, M.A., and K.-Q. Gao. 1997. Braincase and phylogenetic relationships of Estesia mongoliensis from the Late Cretaceous of the Gobi Desert and the recognition of a new clade of lizards. American Museum Novitates 3211: 1-25.
53. Norell, M.A., M.C. McKenna, and M.J. Novacek. 1992. Estesia mongoliensis, a new fossil varanoid from the Late Cretaceous Barun Goyot Formation of Mongolia. American Museum Novitates 3045: 1-24.
54. Norell, M.A., K.-Q. Gao, and J.L. Conrad. 2008. A new platynotan lizard (Diapsida: Squamata) from the Late Cretaceous Gobi Desert (Ömnögov), Mongolia. American Museum Novitates 3605: 1-22.
55. Northcutt, R.G. 1978. Forebrain and midbrain organization in lizards and its phylogenetic significance. In N. Greenberg and P.D. MacLean (editors), Behavior and neurology of lizards, 11-64. Rockville: National Institute of Mental Health.
56. Nydam, R.L. 2000. A new taxon of helodermatid-like lizard from the Albian-Cenomanian of Utah, Journal of Vertebrate Paleontology 20: 285-294.
57. Pepin, D.J. 1999. The origin of monitor lizards based on a review of the fossil evidence. In H.-G. Horn and W. Böhme (editors), Mertensiella: advances in monitor research II, 11-42. Rhine Brook: Deutesche Gesellschaft für Herpetologie und Terrarienkunde.
58. Pepin, DJ. 2001. Natural history of monitor (family Varanidae) with evidence from phylogeny, ecology, life history and morphology. Ph.D. dissertation, Washington University, 234 pp.
59. Pianka, E.R. 1995. Evolution of body size: varanid lizards as a model system. American Naturalist 146: 398-414.
60. Pregill, G.K., J.A. Gauthier, and H.W. Greene. 1986. The evolution of helodermatid squamates, with description of a new taxon and an overview of Varanoidea. Transactions of the San Diego Society of Natural History 21: 167-202.
61. Rieppel, O. 1980a. The phylogeny of anguinimorph lizards. Basel: Naturforschenden Gesellshaft, 86 pp.
62. Rieppel, O. 1980b. The postcranial skeleton of Lanthanotus borneensis (Reptilia, Lacertilia). Amphibia-Reptilia 1: 95-112.
63. Rieppel, O. 1983. A comparison of the skull of Lanthanotus boreensis (Reptilia: Varanoidea) with the skull of primitive snakes. Zeitschrift für zoologische Systematik und Evolutions-forschung 21: 142-153.
64. Rieppel, O. 1993. Patterns of diversity in the reptilian skull. In J. Hanken and B.K. Hall (editors), The skull: v. 2. patterns of structural and systematic diversity, 344-390. Chicago: University of Chicago Press.
65. Rieppel, O., and H. Zaher. 2000a. The intramandibular joint in squamates, and the phylogenetic relationships of the fossil snake Pachyrhachis problematicus Haas. Fieldiana (Geology), new series 43: 1-69.
66. Rieppel, O., and H. Zaher. 2000b. The braincases of mosasaurs and Varanus, and the relationships of snakes. Zoological Journal of the Linnean Society 129: 489-514.
67. Rieppel, O., and L. Grande. 2007. The anatomy of the fossil varanid lizard Saniwa ensidens Leidy, 1870, based on a newly discovered complete skeleton. Journal of Paleontology 81: 643-665.
68. Rieppel, O., J.L. Conrad, and J.A. Maisano. 2006. Morphology and phylogenetics of Eosaniwa (Squamata: Reptilia) based on high-resolution x-ray computed tomography. Journal of Vertebrate Paleontology 26(suppl. 3): 115A.
69. Swofford, D.L. 2001. PAUP*: Phylogenetic Analysis Using Parsimony, v. 4.0b10. Smithsonian Institution, Washington, DC Uetz, P. 2007. The Reptile Database (http://www.reptile-database.org). European Molecular Biology Laboratory, Heidelberg. Accessed 2007.
70. Uetz, P. 2008. The Reptile Database (http://www.reptile-database.org). European Molecular Biology Laboratory, Heidelberg. Accessed 2008.
71. Wu, X.-c, D.B. Brinkman, and A.P. Russell. 1996. Sineoamphisbaena hexatabularis, an amphisbaenian (Diapsida: Squamata) from the Upper Cretaceous redbeds at Bayan Mandahu (Inner Mongolia, People's Republic of China), and comments on the phylogenetic relationships of the Amphisbaenia. Canadian Journal of Earth Sciences 33: 541-577.
72. Zaher, H., and O. Rieppel. 1999a. The phylogenetic relationships of Pachyrhachis problematicus, and the evolution of limblessness in snakes (Lepidosauria, Squamata). Comptes rendus de l'Académie des sciences, série IIa, Earth and Planetary Sciences 329: 831-837.
73. Zaher, H., and O. Rieppel. 1999b. Tooth implantation and replacement in squamates, with special reference to mosasaur lizards and snakes. American Museum Novitates 327: 1-19.