Vermetid reefs and their use as palaeobathymetric markers: New insights from the Late Miocene of the Mediterranean (Southern Italy, Crete)

Palaeogeography, Palaeoclimatology, Palaeoecology

Volumn 267, Issue 1-2, 2008, Pages 89-101

  Vescogni, Alessandro ^a   Bosellini, Francesca R ^a   Reuter, Markus ^b   Brachert, Thomas C ^c  

^a UNIVERSITY OF MODENA AND REGGIO EMILIA   (Italy)

^b UNIVERSITY OF GRAZ   (Austria)

^c UNIVERSITY OF LEIPZIG   (Germany)

Author keywords

Late Miocene; Mediterranean; Palaeobathymetry; Reefs; Vermetids

Indexed keywords

BADENIAN; BATHYMETRY; GASTROPOD; HOLOCENE; INTERTIDAL ENVIRONMENT; MESSINIAN; MIOCENE; RECONSTRUCTION; REEF; SEA LEVEL; SHELL; SUBTIDAL ENVIRONMENT; TORTONIAN;

CAMPANIA [ITALY]; CRETE; EURASIA; EUROPE; GREECE; ITALY; MEDITERRANEAN SEA; SALENTO; SALERNO; SOUTHERN EUROPE;

ALGAE; ANTHOZOA; BRYOZOA; CHLOROPHYTA; DENDROPOMA; GASTROPODA; HALIMEDA; PETALOCONCHUS; PORITES; SERPULIDAE; VERMETIDAE;

EID: 50149086947     PISSN: 00310182     EISSN: None     Source Type: Journal    
DOI: 10.1016/j.palaeo.2008.06.008     Document Type: Article

References (111)

1. 14C ages. Mar. Geol. 159 (1999) 323-339
2. Antonioli F., Chemello R., Improta S., and Riggio S. Dendropoma lower intertidal reef formation and their palaeoclimatological significance, NW Sicily. Mar. Geol. 161 (1999) 155-170
3. Antonioli F., Cremona G., Immordino F., Puglisi C., Romagnoli C., Silenzi S., Valpreda E., and Verrubbi V. New data on the Holocenic sea-level rise in NW Sicily (Central Mediterranean Sea). Global Plan. Change 34 (2002) 121-140
4. Barash A., and Zenziper Z. Structural and biological adaptations of Vermetidae (Gastropoda). Boll. Malacol. 21 (1985) 145-176
5. Baroncelli M.A. Ricostruzione paleoecologica di un'associazione a Petaloconchus glomeratus (Vermetidae) del Pliocene della Valle Botto (Piemonte, Italia N.O.). Boll. Mus. Reg. Sci. Nat. Torino 18 (2000) 209-249
6. Betzler C., Martin J.C., and Braga J.C. Non-tropical carbonates related to rocky submarine cliffs (Miocene, Almeria, southern Spain). Sedim. Geol. 131 (2000) 51-65
7. Bosellini F.R. Diversity, composition and structure of Late Eocene Shelf-edge Coral Associations (Nago Limestone, Northern Italy). Facies 39 (1998) 203-226
8. Bosellini F.R., and Perrin C. The coral fauna of Vitigliano: qualitative and quantitative analysis in a back-reef environment (Castro Limestone, Late Oligocene, Salento Peninsula, southern Italy). Boll. Soc. Paleontol. Ital. 33 (1994) 171-181
9. Bosellini A., Bosellini F.R., Colalongo M.L., Parente M., Russo A., and Vescogni A. Stratigraphic architecture of the Salento coast from Capo d'Otranto to S. Maria di Leuca (Apulia, southern Italy). Riv. Ital. Paleontol. Stratigr. 105 (1999) 397-416
10. Bosellini F.R., Russo A., and Vescogni A. Messinian reef-building assemblages of the Salento Peninsula (southern Italy): palaeobathymetric and palaeoclimatic significance. Palaeogeogr., Palaeoclimatol., Palaeoecol. 175 (2001) 7-26
11. Bosellini F.R., Russo A., and Vescogni A. The Messinian Reef Complex of the Salento Peninsula (southern Italy). Stratigraphy, facies and palaeoenvironmental interpretation. Facies 47 (2002) 91-112
12. Boucot A.J. Principles of Benthic Marine Paleoecology (1981), Academic Press London, London
13. Brachert T.C., Vescogni A., Bosellini F.R., Reuter M., and Mertz-Kraus R. High salinity variability during the early Messinian revealed by stable isotope signatures from vermetid and Halimeda reefs of the Mediterranean region. Geologica Romana 40 (2007) 51-66
14. Braga J.C., Martin J.M., and Riding R. Internal structure of segment reefs: Halimeda algal mounds in the Mediterranean Miocene. Geology 24/1 (1996) 35-38
15. Burchette T.P., and Riding R. Attached vermiform gastropods in Carboniferous marginal marine stromatolites and biostromes. Lethaia 10 (1977) 17-28
16. Carpenter P.P. Firsts steps towards a monograph of the Recent species of Petaloconchus, a genus of Vermetidae. Proc. zool. Soc. Lond. for 1857 (1857) 313-317
17. Colgan M.W. Growth-rate reduction and modification of a coral colony by a vermetid mollusc, Dendropoma maxima. Proc. 5th Int. Coral Reef Congress, Tahiti (1985) 205-210
18. Dalongeville R., Laborel J., Pirazzoli P.A., Sanlaville P., Arnold M., Bernier P., Evin J., and Montaggioni L.F. Les variations recentes de la ligne de rivage sur le littoral syrien. Quaternaire 4 (1993) 45-53
19. Daudin F.M. Recueil de memoires et de notes sur les espèces inédites ou peu connues de mollusques, de vers et de zoophytes. Privately printed, Paris (1800)
20. Esteban M. An overview of Miocene Reefs from Mediterranean areas: general trend and facies models. In: Franseen E.K., Esteban M., Ward W.C., and Rouchy J. (Eds). Models for Carbonate Stratigraphy From Miocene Reef Complexes of Mediterranean Region. Concepts in Sedimentology and Paleontology vol. 5 (1996), SEPM, Tulsa 3-53
21. Fevret M., and Sanlaville P. L'utilisation des Vermet dans la détermination des anciens niveaux marins. Méditerranée 4 (1966) 357-364
22. Focke J.W. The effect of a potentially reef-building vermetid-coralline algal community on a eroding limestone coast, Curaçao, Netherland Antilles. Proc. 3rd Int. Coral Reef. Symp., Miami (1977) 239-245
23. Garcin M., and Vachard D. Recifs, trottoirs et autres biolithosores du bassin de San Miguel de Salinas (Messinien des Cordilleres Betiques, Espagne). Abstracts Interim Colloquium, R.C.M.N.S., Miocene Reefs and Carbonate Platforms of the Mediterranean, Marseille, 23 (1994)
24. Garvie C.L. The molluscan macrofauna of the Reklaw Formation, Marquez Member (Eocene: Lower Claibornian) in Texas. Bull. Am. Paleont. 111 353 (1996) 1-177
25. Ginsburg R.N., and Schroeder J.H. Growth and submarine fossilization of algal cup reefs, Bermuda. Sedimentology 20 (1973) 575-614
26. Gould S.J. Petaloconchus sculpturatus alaminatus, a new Pliocene subspecies of vermetid gastropods lacking its defining generic character, with comments on vermetid systematics in general. J. Paleontol. 68 (1994) 1025-1036
27. Grasso M., Lentini F., and Pedley H.M. Late Tortonian-Lower Messinian (Miocene) palaeogeography of SE Sicily: information from two new formations of the Sortino Group. Sedim. Geol. 32 (1982) 279-300
28. Hadfield M.G. Observations on the anatomy and biology of two California vermetid Gastropods. Veliger 12 (1970) 301-309
29. Hadfield M.G., Kay E.A., Gillette M.U., and Lloyd M.C. The Vermetidae (Mollusca, Gastropoda) of the Hawaiian Islands. Mar. Biol. 12 (1972) 81-98
30. Hillis-Colinvaux L. Halimeda growth and diversity on the deep fore reef on Enewetak Atoll. Coral Reefs 5 (1986) 19-21
31. Hine A.C., Hallock P., Harris M.W., Mullins H.T., Belknap D.F., and Jaap W.C. Halimeda bioherms along an open seaway: Miskito Channel, Nicaragua Rise, SW Caribbean Sea. Coral Reefs 6 (1988) 173-178
32. Hubbard D.K., Gill I.P., and Burke R.B. The Role of Framework in Modern Reefs and Its Application to Ancient Reef Systems. In: Stanley Jr. G.D. (Ed). The History and Sedimentology of Ancient Reef Systems. Topics in Geobiology vol. 17 (2001), Kluwer/Plenum, New York 351-386
33. Hughes R.N. Coloniality in Vermetidae (Gastropoda). In: Larwood G., and Rosen B.R. (Eds). Biology and Systematics of Colonial Organisms. Systematics Association Special Volume 11 (1979), Academic Press, London and New York 243-253
34. Hughes R.N. The vermetid gastropods of Rottnest Island, Western Australia. In: Wells F.E., Walker D.I., Kirkman H., and Lethbridge R. (Eds). The Marine Flora and Fauna of Rottnest Island, Western Australia. Proceedings of the 5th International Marine Biological Workshop (1993), Western Australian Museum, Perth 193-205
35. Insalaco E. Upper Jurassic microsolenid biostromes of northern and central Europe: facies and depositional environment. Palaeogeogr., Palaeoclimatol., Palaeoecol. 121 (1996) 169-194
36. Insalaco E. The descriptive nomenclature and classification of growth fabrics in fossil scleractinian reefs. Sediment. Geol. 118 (1998) 159-186
37. Jindrich V. Structure and diagenesis of recent algal-foraminifer reefs, Fernando de Noronha, Brazil. J. Sed. Petr. 53 (1983) 449-459
38. Jones B., and Hunter I.G. Vermetids buildups from Grand Cayman, British West Indies. J. Coast. Res. 11 (1995) 973-983
39. Keen A.M. Vermetid Gastropods and Marine Intertidal Zonation. Veliger 3 (1960) 1-2
40. Keen A.M. A proposed reclassification of the Gastropod Family Vermetidae. Bull. Br. Mus. Nat. Hist. Ser. D 7 (1961) 183-213
41. Keen A.M. Sea Shells of Tropical West America. Ed.II (1971), Stanford University Press, Stanford
42. Kempf M., and Laborel J. Formations de vermets et d'algues calcaires sur les cotes du Brasil. Recl. Trav. Stn. Mar. Endoume 43 (1968) 9-23
43. Keupp H., and Bellas S.M. Neogene development of the sedimentary basins of NW Crete island, Chania Prefecture, South Aegean Arc System (Greece). Berliner Geowiss. Abh. E 34 (2000) 3-117
44. Keupp H., Bellas S.M., Frydas D., and Kohring R. Aghia Irini, ein Neogenprofil auf der Halbinsel Gramvoùssa/NW-Kreta. Berliner Geowiss. Abh. E 13 (1994) 469-481
45. Kiessling W., and Flügel E. Paleoreefs - a database on Phanerozoic Reefs. In: Kiessling W., Flügel E., and Golonka J. (Eds). Phanerozoic Reefs Patterns. Special Publication vol. 72 (2002), SEPM, Tulsa 625-690
46. Laborel J. Are reef-building vermetids disappearing in the South Atlantic?. Proc. 3rd Int. Coral Reef. Symp., Miami (1977) 233-237
47. Laborel J. Vermetid gastropods as sea-level indicators. In: van de Plassche O. (Ed). Sea-level research: a manual for the collection and evaluation of data (1986), Geo Books, Norwich 281-310
48. Laborel J., and Delibrias G. Niveaux marins recents a Vermetidae du litoral ouest africain. Description, datation et comparison avec les niveaux homologues des côtes du Brésil. Ass. Sénégal. Et. Quatern. Afr., Bull. 47 (1976) 97-110
49. Laborel J., and Laborel-Deguen F. Biological indicators of relative sea-level variations and coaseismic displacements in the Mediterranean Region. J. Coast. Res. 10 (1994) 395-415
50. Laborel J., and Laborel-Deguen F. Biological indicators of Holocene sea-level and climatic variations on rocky coasts of tropical and subtropical regions. Quarter. Int. 31 (1996) 53-60
51. Lea H.C. Descriptions of some new fossil shells, from the Tertiary of Petersburg, Virginia. Trans. Amer. phil. Soc. 9 (1843) 229-274
52. Lipkin Y., and Safriel U.N. Intertidal zonation on rocky shores at Mikhmoret (Mediterranean, Israel). J. Ecol. 59 (1971) 1-30
53. Loya Y. Plotless and transect methods. In: Stoddart D.R., and Johannes R.E. (Eds). Coral Reefs: Research Methods (1978), UNESCO, Paris 197-217
54. Marshall J.F., and Davies P.J. Halimeda bioherms of the northern Great Barrier Reef. Coral Reefs 6 (1988) 139-148
55. Meischner D., and Meischner U. Bermuda south shore reef morphology. A preliminary report. Proc. 3rd Int. Coral Reef. Symp., Miami (1977) 243-250
56. Meulenkamp J.E., Dermitzakis M., Georgiadou-Dikeoulia E., Jonkers H.A., and Böger H. Field guide to the Neogene of Crete (1979), University of Athens, Athens
57. Montenat C., Ott D'Estevou P., Laziat J.C., and Chapel J. La signification des faunes marines contemporaines des évaporites messiniennes dans le SE de l'Espagne. Conséquences pour l'interprétation des conditions d'isolement de la Méditerranée occidentale. Géol. Méd. 7 (1980) 81-90
58. Mörch O.A.L. Review of the Vermetidae (Part I). Proc. zool. Soc. Lond. for 1861 (1861) 145-181
59. Morton J.E. Form and function in the evolution of Vermetidae. Bull. Br. Mus. Nat. Hist. 11 (1965) 585-630
60. Orme R., Riding, and R. Halimeda Segment Reefs of the northern Great Barrier Reef. British Sedimentological Research Group, 1995 Annu. Meet. Abstr., Durham (1995) 64
61. Pedley H.M. Miocene reefs distribution and their association in the central Mediterranean region: an overview. In: Franseen E.K., Esteban M., Ward W.C., and Rouchy J. (Eds). Models for Carbonate Stratigraphy From Miocene Reef Complexes of Mediterranean Region. Concepts in Sedimentology and Paleontology vol. 5 (1996), SEPM, Tulsa 73-87
62. Pedley H.M. Miocene reefs facies of the Pelagian Region (Central Mediterranean). In: Franseen E.K., Esteban M., Ward W.C., and Rouchy J. (Eds). Models for Carbonate Stratigraphy From Miocene Reef Complexes of Mediterranean Region. Concepts in Sedimentology and Paleontology vol. 5 (1996), SEPM, Tulsa 247-259
63. Pérès J.M., and Picard J. Noveau Manuel de Bionomie Bentique de la Mer Mediterranee. Recl. Trav. Stn. Mar. Endoume 31 (1964) 137
64. Perrin C., Bosence D.W., and Rosen B. Quantitative approaches to palaeozonation and palaeobathimetry of corals and coralline algae in Cenozoic reefs. In: Bosence D.W., and Allison P.A. (Eds). Marine Palaeoenvironmental Analisys from Fossils. Special Publication vol. 83 (1995), The Geological Society, London 181-229
65. Pirazzoli P.A., Laborel J., Saliège J.F., Erol O., Kayan I., and Person A. Holocene raised shorelines on the Hatay coasts (Turkey): palaeoecological and tectonic implications. Mar. Geol. 96 (1991) 295-311
66. Pirazzoli P.A., Bellevue M., and Laborel J. Coastal indicators of rapid uplift and subsidence: examples from Crete and other Eastern Mediterranean Sites. Z. Geomorphol. 102 (1996) 21-35
67. Pisera A. Paleoecology and lithogenesis of the Middle Miocene (Badenian) algal vermetid reefs from the Roztocze Hills, south eastern Poland. Acta Geol. Pol. 35 (1985) 89-155
68. Pisera A. Miocene reefs of the Paratethys: a review. In: Franseen E.K., Esteban M., Ward W.C., and Rouchy J. (Eds). Models for Carbonate Stratigraphy From Miocene Reef Complexes of Mediterranean Region. Concepts in Sedimentology and Paleontology vol. 5 (1996), SEPM, Tulsa 97-104
69. Pomar L., Ward W.C., and Green D.G. Upper Miocene Reef complex of the Llucmajor area, Mallorca, Spain. In: Franseen E.K., Esteban M., Ward W.C., and Rouchy J. (Eds). Models for Carbonate Stratigraphy From Miocene Reef Complexes of Mediterranean Region. Concepts in Sedimentology and Paleontology vol. 5 (1996), SEPM, Tulsa 191-225
70. Rafinesque C.S. Analyses de la nature ou tableau du univers et des corps organisés (1815), Barravecchia, Palermo
71. Reuter M., and Brachert T.C. Freshwater discharge and sediment dispersal - control on growth, ecological structure and geometry of Late Miocene shallow-water coral ecosystems (early Tortonian, Crete/Greece). Palaeogeogr., Palaeoclim., Palaeoec. 255 (2007) 308-328
72. Reuter M., Brachert T.C., and Kroeger K.F. Shallow-marine carbonates of the tropical - temperate transition zone: effects of hinterland climate and basin physiography (Late Miocene, Crete/Greece). In: Pedley H.M., and Carannante S. (Eds). Coolwater Carbonates (2006), The Geological Society, London
73. Riding R. Structure and composition of organic reefs and carbonate mud mounds: concepts and categories. Earth-Sci. Rev. 58 (2002) 163-231
74. Riegl B., and Piller W.E. Reefs and coral carpets in the northern Red Sea as a model for organism-environment feedback in coral communities and its reflection in growth fabrics. In: Insalaco E., Skelton P.W., and Palmer T.J. (Eds). Carbonate Platform Systems: components and interactions. Special Publication vol. 178 (2002), The Geological Society, London 71-88
75. Roberts H.H., and Macintyre I.G. Preface to Halimeda Special Issue. Coral Reefs 6 (1988) 121
76. Roberts H.H., Aharon P., and Phipps C.V. Morphology and sedimentology of Halimeda bioherms from the eastern Java Sea (Indonesia). Coral Reefs 6 (1988) 161-172
77. Rosen B.R. Reefs and carbonate build-ups. In: Briggs D.E.G., and Crowther P.R. (Eds). Palaeobiology: A Synthesis (1990), Blackwell, Oxford 341-346
78. Rosen B.R. Paleoclimatic implications of the energy hypothesis from Neogene corals of the Mediterranean region. In: Agusti J., Rook L., and Andrews P. (Eds). The evolution of Neogene terrestrial ecosystems in Europe (1999), Cambridge University Press, Cambridge 309-327
79. Rosen B., Aillud G., Bosellini F.R., Clack N., Insalaco E., Valldeperas F.X., and Wilson M. Platy coral assemblages: 210 million years of response to the limiting effect of depth, light and turbidity. Proc. 9th Int. Coral Reef. Symp., Bali (2002) 255-294
80. Safriel U.N. Recent vermetid formation on the Mediterranean coast of Israel. Proc. Malacol. Soc. London 37 (1966) 27-34
81. Safriel U.N. Vermetid gastropods and intertidal reefs in Israel and Bermuda. Science 186 (1974) 1113-1115
82. Safriel U.N. The role of the Vermetid Gastropods in the formation of Mediterranean and Atlantic reefs. Oecologia 20 (1975) 85-101
83. Safriel U.N., and Hadfield M.G. Sibling speciation by life-history divergence in Dendropoma (Gastropoda; Vermetidae). Biol. J. Linnean Soc. 35 (1988) 1-13
84. Saint Martin J.P. Les formations récifales coralliennes du Miocène supérieur d'Algérie et du Maroc. Mem. Mus. Nat. Hist. Nat. Paris Ser. C 56 (1990) 1-366
85. Saint Martin J.P., and Andre J. Les construction coralliennes de la plateforme carbonatée messinienne de Malte (Méditerranée centrale). Géol. Med. 19 (1992) 145-163
86. Sassi A. Saggio geologico sopra il Bacino terziario di Albenga. G. linguist. Sci. Lett. Art. Genova 1 (1827) 467-484
87. Savazzi E. Adaptations of Vermetid and Siliquariid Gastropods. Palaeontology 39 (1996) 157-177
88. Savazzi E. Cemented and Embedded Gastropods. In: Savazzi E. (Ed). Functional Morphology of the Invertebrate Skeleton (1999), John Wiley and Sons Ltd., Chichester 183-195
89. Savazzi E. Morphodynamics of an endolithic vermetid gastropod. Paleontol. Res. 5 (2001) 3-11
90. Schiapparelli S. Contribution to the knowledge of Vermetidae (Mollusca: Gastropoda) from the Ligurian Sea. Boll. Malacol. 31 (1995) 267-276
91. Schiapparelli S., and Cattaneo-Vietti R. Functional morphology of vermetid feeding tubes. Lethaia, 32 (1999) 41-46
92. Schiapparelli S., Albertelli G., and Cattaneo-Vietti R. Phenotypic plasticity of Vermetidae suspension feeding: a potential bias in their use as a biological sea-level indicators. Mar. Ecol. 27 (2006) 44-53
93. Scuderi D. Il genere Dendropoma (Gastropoda: Vermetidae) nel Mediterraneo. Boll. Malacol. 31 (1995) 1-6
94. Scuderi D., Terlizzi A., and Faimali M. Osservazioni su alcuni tratti della biologia riproduttiva dei vermetidi biocostruttori e loro ruolo nell'edificazione dei "trottoir". Biol. Mar. Medit. 5 (1998) 284-289
95. Shier D.E. Vermetid reefs and coastal development in ten thousand islands, Southwest Florida. Geol. Soc. Bull. 80 (1969) 485-508
96. Silenzi S., Antonioli F., and Chemello R. A new marker for sea surface temperature trend during the last centuries in temperate areas: Vermetid reef. Global Plan. Change 40 (2004) 105-114
97. Stiros S.C., Laborel J., Laborel-Deguen F., Papageorgiou S., Evin J., and Pirazzoli P.A. Seismic coastal uplift in a region of subsidence: Holocene raised shorelines of Samos Island, Aegean Sea, Greece. Mar. Geol. 170 (2000) 41-58
98. Templado J., Templado D., and Calvo M. The formation of the vermetid gastropod Dendropoma petraeum (Monterosato, 1884) on the coast of the Iberian Peninsula (Western Mediterranean). In: Giusti F., and Manganelli G. (Eds) (1992), Abstract 11° International Malacological Congress, Siena 514-515
99. ten Hove H.A., and van den Hurk P. A review of Recent and fossil serpulid "reefs"; Actuopalaeontology and the "Upper Malm" serpulid limestone in NW Germany. Geol. Mijnb. 72 (1993) 23-67
100. Thomas M.L.H., and Stevens J.A. Communities of constructional lips and cup reef rims in Bermuda. Coral Reefs 9 (1991) 225-230
101. Thommeret Y., Thommeret J., Laborel J., Montaggioni L.F., and Pirazzoli P.A. Late Holocene shoreline changes and seismotectonic displacements in western Crete (Greece). Z. Geomorphol. 40 (1981) 127-149
102. Tzur Y., and Safriel U.N. Vermetid Platforms as indicators of coastal movements. Israel J. Earth Sci. 27 (1978) 124-127
103. van Andel T.H., and Laborel J. Recent High Relative Sea-level Stand near Recife, Brazil. Science 145 (1964) 580-581
104. Verdinelli M.E., and Verdinelli M.A. Thanatocoenosis of Vermetids of the Genus Petaloconchus on the Island of Santa Catarina - Brazil. An. Acad. Bras. Ci. 68 (1996) 479-484
105. Vescogni A. Evoluzione delle biocostruzioni a vermetidi e loro utilizzo come "markers" paleobatimetrici e paleoclimatici. G. Geol. 62 (2000) 55-61
106. Wood R. Reef evolution (1999), Oxford University Press, Oxford
107. Zuschin M., and Piller W.E. Molluscan hard-substrate associations in the Northern Red Sea. Mar. Ecol. 18 (1997) 361-378
108. Zuschin M., and Stachowitsh M. The distribution of molluscan assemblages and their postmortem fate on coral reefs in the Gulf of Aqaba (northern Red Sea). Mar. Biol. 151 (2007) 2217-2230
109. Zuschin M., Hoehenegger J., and Steininger F.F. A comparison of living and dead molluscs on coral reef associated hard substrata in the northern Red Sea - implications for the fossil record. Palaeogeogr., Palaeoclimatol., Palaeoecol. 159 (2000) 167-190
110. Zuschin M., Hoehenegger J., and Steininger F.F. Molluscan assemblages on coral reefs and associated hard substrata in the northern Red Sea. Coral Reefs 20 (2001) 107-116
111. Zuschin M., Harzhauser M., and Mandic O. The stratigraphic and sedimentologic famework of fine-scale faunal replacement in the Middle Miocene of the Vienna Basin (Austria). Palaios 22 (2007) 285-295