메뉴 건너뛰기




Volumn 32, Issue 3, 2007, Pages 502-538

Phylogeny of Psephenidae (Coleoptera: Byrrhoidea) based on larval, pupal and adult characters

Author keywords

[No Author keywords available]

Indexed keywords

ADAPTIVE RADIATION; ADULT; BEETLE; CLADISTICS; LARVAL DEVELOPMENT; PARSIMONY ANALYSIS; PHYLOGENY; PUPA; TAXONOMY;

EID: 34447281268     PISSN: 03076970     EISSN: 13653113     Source Type: Journal    
DOI: 10.1111/j.1365-3113.2006.00374.x     Document Type: Article
Times cited : (32)

References (60)
  • 1
    • 0011609992 scopus 로고
    • Two new synonymies: Alabameubria Brown, a junior synonym of Dicranopselaphus and Alabameubria starki, a synonym of Dicranoselaphus variegatus (Coleoptera: Psephenidae)
    • Barr, C.B. Spangler, P.J 1994) Two new synonymies: Alabameubria Brown, a junior synonym of Dicranopselaphus and Alabameubria starki, a synonym of Dicranoselaphus variegatus (Coleoptera: Psephenidae). Entomological News, 105, 299 302.
    • (1994) Entomological News , vol.105 , pp. 299-302
    • Barr, C.B.1    Spangler, P.J.2
  • 4
    • 0000074009 scopus 로고
    • Phylogenetic analysis of Elateriformia (Coleoptera: Polyphaga) based on larval characters
    • Beutel, R.G 1995) Phylogenetic analysis of Elateriformia (Coleoptera: Polyphaga) based on larval characters. Journal of Zoological Systematics and Evolutionary Research, 33, 145 171.
    • (1995) Journal of Zoological Systematics and Evolutionary Research , vol.33 , pp. 145-171
    • Beutel, R.G.1
  • 5
    • 22944465259 scopus 로고
    • The larva of Eubrianax edwardsi (LeC.) (Coleoptera: Psephenidae)
    • Blackwelder, R.E 1930) The larva of Eubrianax edwardsi (LeC.) (Coleoptera: Psephenidae). Pan-Pacific Entomologist, 6, 139 141.
    • (1930) Pan-Pacific Entomologist , vol.6 , pp. 139-141
    • Blackwelder, R.E.1
  • 6
    • 34447280924 scopus 로고
    • The immature stages of Psephenoides gahani Champ. (Coleoptera: Dryopidae)
    • Böving, A.G 1926) The immature stages of Psephenoides gahani Champ. (Coleoptera: Dryopidae). Transactions of the Entomological Society of London, 74, 381 388.
    • (1926) Transactions of the Entomological Society of London , vol.74 , pp. 381-388
    • Böving, A.G.1
  • 7
    • 0008929980 scopus 로고
    • On the classification of beetles according to larval characters
    • Böving, A.G 1929) On the classification of beetles according to larval characters. Bulletin of the Brooklyn Entomological Society, 24, 55 97.
    • (1929) Bulletin of the Brooklyn Entomological Society , vol.24 , pp. 55-97
    • Böving, A.G.1
  • 8
    • 0001900143 scopus 로고
    • An illustrated synopsis of the principal larval forms of Coleoptera
    • Böving, A.G. Craighead, F.G 1931) An illustrated synopsis of the principal larval forms of Coleoptera. Entomologica Americana, 11, 1 351.
    • (1931) Entomologica Americana , vol.11 , pp. 1-351
    • Böving, A.G.1    Craighead, F.G.2
  • 9
    • 43949160320 scopus 로고
    • Branch support and tree stability
    • Bremer, K 1994) Branch support and tree stability. Cladistics, 10, 295 304.
    • (1994) Cladistics , vol.10 , pp. 295-304
    • Bremer, K.1
  • 10
    • 0038616516 scopus 로고
    • Ectopria leechi, a new false water penny from the United States (Coleoptera: Eubriidae)
    • Brigham, W.U 1981) Ectopria leechi, a new false water penny from the United States (Coleoptera: Eubriidae). Pan-Pacific Entomologist, 57, 313 320.
    • (1981) Pan-Pacific Entomologist , vol.57 , pp. 313-320
    • Brigham, W.U.1
  • 14
    • 0001196923 scopus 로고    scopus 로고
    • Systematics and bionomics of Cneoglossidae with a cladistic analysis of Byrrhoidea sensu Lawrence & Newton (1995) (Coleoptera, Elateriformia)
    • Costa, C., Vannin, S.A. Ide, S 1999) Systematics and bionomics of Cneoglossidae with a cladistic analysis of Byrrhoidea sensu Lawrence & Newton (1995) (Coleoptera, Elateriformia). Arquivos de Zoologia, 35, 231 300.
    • (1999) Arquivos de Zoologia , vol.35 , pp. 231-300
    • Costa, C.1    Vannin, S.A.2    Ide, S.3
  • 16
    • 0008070702 scopus 로고
    • Problems of phylogenetic relationships in Dryopoidea (Coleoptera)
    • Crowson, R.A 1978) Problems of phylogenetic relationships in Dryopoidea (Coleoptera). Entomologia Generalis, 4, 250 257.
    • (1978) Entomologia Generalis , vol.4 , pp. 250-257
    • Crowson, R.A.1
  • 17
    • 0040295343 scopus 로고
    • The taxonomic significance of the characters of immature insects
    • Emden, F.I 1957) The taxonomic significance of the characters of immature insects. Annual Review of Entomology, 2, 91 106.
    • (1957) Annual Review of Entomology , vol.2 , pp. 91-106
    • Emden, F.I.1
  • 18
    • 34447253640 scopus 로고
    • A new genus of Coleoptera of the family Psephenidae
    • Gahan, C.J 1914) A new genus of Coleoptera of the family Psephenidae. Entomologist, 47, 188 189.
    • (1914) Entomologist , vol.47 , pp. 188-189
    • Gahan, C.J.1
  • 19
    • 0003505452 scopus 로고
    • Published by the author. Instituto Miguel Lillo, Tucuman.
    • Goloboff, P 1993) NONA, version 1.8. (32 bit). Published by the author. Instituto Miguel Lillo, Tucuman.
    • (1993) NONA, Version 1.8. (32 Bit).
    • Goloboff, P.1
  • 20
    • 84977258371 scopus 로고
    • An inquiry into the natural classification of the Dryopoidea, based partly on a study of their internal anatomy (Col.)
    • Hinton, H.E 1939) An inquiry into the natural classification of the Dryopoidea, based partly on a study of their internal anatomy (Col.). Transactions of the Royal Entomological Society of London, 89, 133 184.
    • (1939) Transactions of the Royal Entomological Society of London , vol.89 , pp. 133-184
    • Hinton, H.E.1
  • 21
    • 0005672666 scopus 로고
    • On the respiratory adaptations, biology, and taxonomy of the Psephenidae, with notes on some related families (Coleoptera)
    • Hinton, H.E 1955) On the respiratory adaptations, biology, and taxonomy of the Psephenidae, with notes on some related families (Coleoptera). Proceedings of the Zoological Society of London, 135, 543 568.
    • (1955) Proceedings of the Zoological Society of London , vol.135 , pp. 543-568
    • Hinton, H.E.1
  • 23
    • 0011610613 scopus 로고
    • Description of Granuleubria, a new genus of Eubriinae from West and South Asia (Coleoptera: Psephenidae)
    • Jäch, M.A. Lee, C.-F. (1994) Description of Granuleubria, a new genus of Eubriinae from West and South Asia (Coleoptera: Psephenidae). Koleopterologische Rundschau, 64, 223 232.
    • (1994) Koleopterologische Rundschau , vol.64 , pp. 223-232
    • Jäch, M.A.1    Lee, C.-F.2
  • 24
    • 33646232227 scopus 로고    scopus 로고
    • On the taxonomy of Psephenoidinae: Revision of Afropsephenoides, and descriptions of two new species and one new genus (Coleoptera: Psephenidae)
    • Jeng, M.-L., Jäch, M.A., Satô, M. Yang, P.S 2006) On the taxonomy of Psephenoidinae: revision of Afropsephenoides, and descriptions of two new species and one new genus (Coleoptera: Psephenidae). Insect Systematics and Evolution, 37, 105 119.
    • (2006) Insect Systematics and Evolution , vol.37 , pp. 105-119
    • Jeng, M.-L.1    Jäch, M.A.2    Satô, M.3    Yang, P.S.4
  • 25
    • 34447255981 scopus 로고
    • Psephenus lecontei - The external anatomy of the larva
    • Kellicott, D.S 1883) Psephenus lecontei - the external anatomy of the larva. Canadian Entomologist, 15, 191 196.
    • (1883) Canadian Entomologist , vol.15 , pp. 191-196
    • Kellicott, D.S.1
  • 27
    • 1842680212 scopus 로고    scopus 로고
    • Relationships among coleopteran suborders and major endoneopteran lineages: Evidence from hind wing characters
    • Kukalová-Peck, J. Lawrence, J.F 2004) Relationships among coleopteran suborders and major endoneopteran lineages: evidence from hind wing characters. European Journal of Entomology, 101, 95 144.
    • (2004) European Journal of Entomology , vol.101 , pp. 95-144
    • Kukalová-Peck, J.1    Lawrence, J.F.2
  • 28
    • 84971072876 scopus 로고
    • Rhinorhipidae, a new beetle family from Australia, with comments on the phylogeny of the Elateriformia
    • Lawrence, J.F 1987) Rhinorhipidae, a new beetle family from Australia, with comments on the phylogeny of the Elateriformia. Invertebrate Taxonomy, 2, 1 53.
    • (1987) Invertebrate Taxonomy , vol.2 , pp. 1-53
    • Lawrence, J.F.1
  • 29
    • 0000240541 scopus 로고
    • Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names)
    • ed. by. J. Pakaluk. and. S.A.S. Łipiński. pp. Museum i Instytut Zoologii PAN, Warszawa.
    • Lawrence, J.F. Newton, A.F 1995) Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson, Vol. 2 (ed. by J. Pakaluk and S.A.S. Łipiński pp. 779 1092. Museum i Instytut Zoologii PAN, Warszawa.
    • (1995) Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson , vol.2 , pp. 779-1092
    • Lawrence, J.F.1    Newton, A.F.2
  • 30
    • 34447249395 scopus 로고    scopus 로고
    • Psephenidae: Revision of the genus Mataeopsephus Waterhouse
    • ed. by. M.A. Jäch. and. L. Ji. pp. Zoologisch-Botanische Gesellschaft in Österreich and Wiener Coleopterologenverein, Wien.
    • Lee, C.-F., Jäch, M.A. Satô, M 2003c) Psephenidae: revision of the genus Mataeopsephus Waterhouse. Water Beetles of China, Vol. 3 (ed. by M.A. Jäch and L. Ji pp. 481 517. Zoologisch-Botanische Gesellschaft in Österreich and Wiener Coleopterologenverein, Wien.
    • (2003) Water Beetles of China , vol.3 , pp. 481-517
    • Lee, C.-F.1    Jäch, M.A.2    Satô, M.3
  • 31
    • 0005672288 scopus 로고    scopus 로고
    • Psephenidae. II. Synopsis of Schinostethus Waterhouse, with descriptions of 14 new species (Coleoptera)
    • ed. by. M.A. Jäch. and. L. Ji. pp. Zoologisch-Botanische Gesellschaft in Österreich and Wiener Coleopterologenverein, Wien.
    • Lee, C.-F., Jäch, M.A. Yang, P.-S. (1998) Psephenidae. II. Synopsis of Schinostethus Waterhouse, with descriptions of 14 new species (Coleoptera). Water Beetles of China, Vol. 2 (ed. by M.A. Jäch and L. Ji pp. 303 326. Zoologisch-Botanische Gesellschaft in Österreich and Wiener Coleopterologenverein, Wien.
    • (1998) Water Beetles of China , vol.2 , pp. 303-326
    • Lee, C.-F.1    Jäch, M.A.2    Yang, P.-S.3
  • 32
    • 0037745797 scopus 로고    scopus 로고
    • Afrobrianax, a new genus of the Eubrianacinae, with notes on its phylogenetic position and description of one new species from South Africa (Coleoptera: Psephenidae)
    • Lee, C.-F., Philips, T.K. Yang, P.-S. (2003a) Afrobrianax, a new genus of the Eubrianacinae, with notes on its phylogenetic position and description of one new species from South Africa (Coleoptera: Psephenidae). African Entomology, 11, 1 8.
    • (2003) African Entomology , vol.11 , pp. 1-8
    • Lee, C.-F.1    Philips, T.K.2    Yang, P.-S.3
  • 33
    • 34447257778 scopus 로고    scopus 로고
    • Revision of Afroeubria Villiers (Coleoptera: Psephenidae: Eubriinae), including phylogenetic notes
    • Lee, C.-F., Philips, T.K. Yang, P.-S. (2003b) Revision of Afroeubria Villiers (Coleoptera: Psephenidae: Eubriinae), including phylogenetic notes. Annals of the Transvaal Museum, 40, 137 143.
    • (2003) Annals of the Transvaal Museum , vol.40 , pp. 137-143
    • Lee, C.-F.1    Philips, T.K.2    Yang, P.-S.3
  • 34
    • 0040601846 scopus 로고    scopus 로고
    • Nipponeubria yoshitomii Lee and Satô, a new species in a new genus of Eubriinae from Japan, with notes on the immature stages and description of the larva of Ectopria opaca (Kiesenwetter) (Coleoptera: Psephenidae)
    • Lee, C.-F. Satô, M 1996) Nipponeubria yoshitomii Lee and Satô, a new species in a new genus of Eubriinae from Japan, with notes on the immature stages and description of the larva of Ectopria opaca (Kiesenwetter) (Coleoptera: Psephenidae). Coleopterists Bulletin, 50, 122 134.
    • (1996) Coleopterists Bulletin , vol.50 , pp. 122-134
    • Lee, C.-F.1    Satô, M.2
  • 35
    • 22944438579 scopus 로고    scopus 로고
    • Revision of Eubrianacinae (Coleoptera, Psephenidae) I. Eubrianax Kiesenwetter and Heibrianax gen.n
    • Lee, C.-F., Satô, M. Yang, P.-S. (1999a) Revision of Eubrianacinae (Coleoptera, Psephenidae) I. Eubrianax Kiesenwetter and Heibrianax gen.n. Japanese Journal of Systematic Entomology, 5, 9 25.
    • (1999) Japanese Journal of Systematic Entomology , vol.5 , pp. 9-25
    • Lee, C.-F.1    Satô, M.2    Yang, P.-S.3
  • 36
    • 0037993165 scopus 로고    scopus 로고
    • Revision of Eubrianacinae (Coleoptera, Psephenidae) II. Mubrianx gen. nov
    • Lee, C.-F., Satô, M. Yang, P.-S. (1999b) Revision of Eubrianacinae (Coleoptera, Psephenidae) II. Mubrianx gen. nov. Elytra, 27, 429 438.
    • (1999) Elytra , vol.27 , pp. 429-438
    • Lee, C.-F.1    Satô, M.2    Yang, P.-S.3
  • 37
    • 0005681446 scopus 로고    scopus 로고
    • Revision of Eubrianacinae (Coleoptera, Psephenidae) III. Jinbrianax gen. nov
    • Lee, C.-F., Satô, M. Yang, P.-S. (1999c) Revision of Eubrianacinae (Coleoptera, Psephenidae) III. Jinbrianax gen. nov. Entomological Review of Japan, 54, 169 187.
    • (1999) Entomological Review of Japan , vol.54 , pp. 169-187
    • Lee, C.-F.1    Satô, M.2    Yang, P.-S.3
  • 38
    • 22944463954 scopus 로고    scopus 로고
    • Revision of Eubrianacinae (Coleoptera, Psephenidae) IV. Odontanax gen. nov
    • Lee, C.-F., Satô, M. Yang, P.-S. (2000a) Revision of Eubrianacinae (Coleoptera, Psephenidae) IV. Odontanax gen. nov. Japanese Journal of Systematic Entomology, 6, 151 170.
    • (2000) Japanese Journal of Systematic Entomology , vol.6 , pp. 151-170
    • Lee, C.-F.1    Satô, M.2    Yang, P.-S.3
  • 39
    • 22944491314 scopus 로고    scopus 로고
    • Revision of Eubrianacinae (Coleoptera, Psephenidae) V. Jaechanax gen. nov
    • Lee, C.-F., Satô, M. Yang, P.-S. (2000b) Revision of Eubrianacinae (Coleoptera, Psephenidae) V. Jaechanax gen. nov. Elytra, 28, 119 129.
    • (2000) Elytra , vol.28 , pp. 119-129
    • Lee, C.-F.1    Satô, M.2    Yang, P.-S.3
  • 40
    • 84981627428 scopus 로고
    • A revision of the genus Homoeogenus Waterhouse with notes on the immature stages of H. laurae sp.n. (Coleoptera: Psephenidae: Eubriinae)
    • Lee, C.-F. Yang, P.-S. (1993) A revision of the genus Homoeogenus Waterhouse with notes on the immature stages of H. laurae sp.n. (Coleoptera: Psephenidae: Eubriinae). Systematic Entomology, 18, 351 361.
    • (1993) Systematic Entomology , vol.18 , pp. 351-361
    • Lee, C.-F.1    Yang, P.-S.2
  • 41
    • 0009149164 scopus 로고
    • Psephenidae: 2. Notes on the genus Homoeogenus Waterhouse from China (Coleoptera)
    • ed. by. M.A. Jäch. and. L. Ji. pp. Zoologisch-Botanische Gesellschaft in Österreich and Wiener Coleopterologenverein, Wien.
    • Lee, C.-F. Yang, P.-S. (1995) Psephenidae: 2. Notes on the genus Homoeogenus Waterhouse from China (Coleoptera). Water Beetles of China, Vol. I (ed. by M.A. Jäch and L. Ji pp. 355 357. Zoologisch-Botanische Gesellschaft in Österreich and Wiener Coleopterologenverein, Wien.
    • (1995) Water Beetles of China , vol.1 , pp. 355-357
    • Lee, C.-F.1    Yang, P.-S.2
  • 42
    • 0030525155 scopus 로고    scopus 로고
    • Taxonomic revision of the Oriental species of Dicranopselaphus Guérin-Méneville (Coleoptera: Psephenidae: Eubriinae)
    • Lee, C.-F. Yang, P.-S. (1996) Taxonomic revision of the Oriental species of Dicranopselaphus Guérin-Méneville (Coleoptera: Psephenidae: Eubriinae). Entomologica Scandinavica, 27, 169 196.
    • (1996) Entomologica Scandinavica , vol.27 , pp. 169-196
    • Lee, C.-F.1    Yang, P.-S.2
  • 43
    • 0005734286 scopus 로고    scopus 로고
    • Notes on the Granuleubria Jäch and Lee, with descriptions of four new species and the immature stages of G. fontana new species
    • Lee, C.-F. Yang, P.-S. (1999) Notes on the Granuleubria Jäch and Lee, with descriptions of four new species and the immature stages of G. fontana new species. Coleopterists Bulletin, 53, 259 274.
    • (1999) Coleopterists Bulletin , vol.53 , pp. 259-274
    • Lee, C.-F.1    Yang, P.-S.2
  • 44
    • 0036063148 scopus 로고    scopus 로고
    • Notes on the larva of the genus Microeubria Lee and Yang, with description of a new species from Borneo (Coleoptera: Psephenidae: Eubriinae)
    • Lee, C.-F. Yang, P.-S. (2002) Notes on the larva of the genus Microeubria Lee and Yang, with description of a new species from Borneo (Coleoptera: Psephenidae: Eubriinae). Coleopterists Bulletin, 56, 246 252.
    • (2002) Coleopterists Bulletin , vol.56 , pp. 246-252
    • Lee, C.-F.1    Yang, P.-S.2
  • 45
    • 0027136435 scopus 로고
    • Revision of the genus Schinosethus Waterhouse with notes on the immature stages and ecology of S. satoi, n. sp
    • Lee, C.-F., Yang, P.-S. Brown, H.P 1993) Revision of the genus Schinosethus Waterhouse with notes on the immature stages and ecology of S. satoi, n. sp. Annals of the Entomological Society of America, 86, 683 693.
    • (1993) Annals of the Entomological Society of America , vol.86 , pp. 683-693
    • Lee, C.-F.1    Yang, P.-S.2    Brown, H.P.3
  • 46
    • 0005707397 scopus 로고    scopus 로고
    • The East Asian species of the genus Macroeubria Pic (Coleoptera, Psephenidae, Eubriinae)
    • Lee, C.-F., Yang, P.-S. Satô, M 1997) The East Asian species of the genus Macroeubria Pic (Coleoptera, Psephenidae, Eubriinae). Japanese Journal of Systematic Entomology, 3, 129 160.
    • (1997) Japanese Journal of Systematic Entomology , vol.3 , pp. 129-160
    • Lee, C.-F.1    Yang, P.-S.2    Satô, M.3
  • 47
    • 0038669607 scopus 로고    scopus 로고
    • Phylogeny of the genera of Eubrianacinae and descriptions of additional members of Eubrianax (Coleoptera: Psephenidae)
    • Lee, C.-F., Yang, P.-S. Satô, M 2001) Phylogeny of the genera of Eubrianacinae and descriptions of additional members of Eubrianax (Coleoptera: Psephenidae). Annals of the Entomological Society of America, 94, 347 362.
    • (2001) Annals of the Entomological Society of America , vol.94 , pp. 347-362
    • Lee, C.-F.1    Yang, P.-S.2    Satô, M.3
  • 48
    • 0034404568 scopus 로고    scopus 로고
    • Phylogenetic systematics of the major lineages of the pannote mayflies (Empheroptera: Pannota)
    • McCafferty, W.P. Wang, T.-Q. (2000) Phylogenetic systematics of the major lineages of the pannote mayflies (Empheroptera: Pannota). Transactions of the American Entomological Society, 126, 9 101.
    • (2000) Transactions of the American Entomological Society , vol.126 , pp. 9-101
    • McCafferty, W.P.1    Wang, T.-Q.2
  • 49
    • 84984483735 scopus 로고
    • The behaviour and form of Psephenus herricki (DeKay) (Coleoptera: Psephenidae) in relation to water flow
    • McShaffrey, D. McCafferty, W.P 1987) The behaviour and form of Psephenus herricki (DeKay) (Coleoptera: Psephenidae) in relation to water flow. Freshwater Biology, 18, 319 324.
    • (1987) Freshwater Biology , vol.18 , pp. 319-324
    • McShaffrey, D.1    McCafferty, W.P.2
  • 51
  • 55
  • 57
    • 34447272996 scopus 로고
    • Family Ptilodactylidae
    • no. 20. Coleopterists' Association of Japan, Department of Zoology, National Science Museum, Tokyo (. in Japanese).
    • Satô, M 1983) Family Ptilodactylidae. Checklist of Coleoptera of Japan, no. 20. Coleopterists' Association of Japan, Department of Zoology, National Science Museum, Tokyo (in Japanese).
    • (1983) Checklist of Coleoptera of Japan
    • Satô, M.1
  • 58
    • 34447255299 scopus 로고
    • Taxonomic notes on the aquatic Coleoptera of Japan III
    • in Japanese).
    • Satô, M 1985) Taxonomic notes on the aquatic Coleoptera of Japan III. Coleopterists' News, 69, 1 5 (in Japanese).
    • (1985) Coleopterists' News , vol.69 , pp. 1-5
    • Satô, M.1
  • 59
    • 0006125604 scopus 로고
    • Boundary layer control by water pennies (Coleoptera: Psephenidae)
    • Smith, J.A. Dartnall, A.J 1980) Boundary layer control by water pennies (Coleoptera: Psephenidae). Aquatic Insects, 2, 65 72.
    • (1980) Aquatic Insects , vol.2 , pp. 65-72
    • Smith, J.A.1    Dartnall, A.J.2
  • 60
    • 0042746746 scopus 로고
    • The aquatic beetle subfamily Larainae (Coleoptera: Elmidae) in Mexico, Central America, and the West Indies
    • Spangler, P.J. Santiago-Fragoso, S 1992) The aquatic beetle subfamily Larainae (Coleoptera: Elmidae) in Mexico, Central America, and the West Indies. Smithsonian Contributions to Zoology, 528, 1 74.
    • (1992) Smithsonian Contributions to Zoology , vol.528 , pp. 1-74
    • Spangler, P.J.1    Santiago-Fragoso, S.2


* 이 정보는 Elsevier사의 SCOPUS DB에서 KISTI가 분석하여 추출한 것입니다.