Reassessment of the Aetosaur 'Desmatosuchus' chamaensis with a reanalysis of the phylogeny of the Aetosauria (Archosauria: Pseudosuchia)

Journal of Systematic Palaeontology

Volumn 5, Issue 1, 2007, Pages 41-68

  Parker, William G ^a  

^a NATIONAL PARK SERVICE   (United States)

Author keywords

Aetosauria; Chinle Formation; Heliocanthus; Late Triassic; New Mexico; Phylogeny; Vertebrate palaeontology

Indexed keywords

FOSSIL RECORD; MORPHOLOGY; NEW GENUS; ORNAMENTATION; PHYLOGENETICS; PHYLOGENY; RELATEDNESS; REPTILE; TRIASSIC;

NEW MEXICO; NORTH AMERICA; UNITED STATES;

ARCHOSAURIA; VERTEBRATA;

EID: 33846895444     PISSN: 14772019     EISSN: 14780941     Source Type: Journal    
DOI: 10.1017/S1477201906001994     Document Type: Article

References (94)

1. Agassiz, L. 1844. Monographie des poisons fossils du Vieux Grés Rouge ou Systéme Dévonien (Old Red Sandstone) des Iles Britanniques ed de Russie. Jent et Gassman: Neuchâtel, 171 pp.
2. Akers, J. P., Cooley, M. E. & Repenning, C. A. 1958. Moenkopi and Chinle Formations of Black Mesa and adjacent areas. New Mexico Geological Society Guidebook 9: 88-94.
3. Benton, M. J. 1999. Scleromochlus taylori and the origin of dinosaurs and pterosaurs. Philosophical Transactions of the Royal Society of Londoon, Series B 354: 1423-1446.
4. - 2004. Origin and relationships of Dinosauria. Pp. 7-19 in Weishampel, D. B., Dodson, P. & Osmólska, H. (eds) The Dinosauria, second edition. University of California Press: Berkeley.
5. Bonaparte, J. F. 1971. El Mesozoico de America del Sur y sus tetrapodos. Opera Lillona 22: 1-183.
6. Camp, C. L. & Welles, S. P. 1956. Triassic dicynodont reptiles: part I, the North American genus Placerias. Memoirs of the University of California 13: 255-304.
7. Carpenter, K. & Parrish, J. M. 1985. Late Triassic vertebrates from Revuelto Creek, Quay County, New Mexico. New Mexico Geological Society Guidebook 36: 197-198.
8. Casamiquela, R. M. 1961. Dos nuevos estagonolepoideos Argentinos (de Ischigualasto, San Juan). Revista Asocíacion Geológia de Argentina 16: 143-203.
9. - 1967. Materiales adicionales y reinterpretación de Aetosauroides scagliai (de Ischigulasto, San Juan). Revista del Museo de la Plata, Paleontologia Serie 5 33: 173-196.
10. Case, E. C. 1920. Preliminary description of a new suborder of phytosaurian reptiles with a description of a new species of Phytosaurus. Journal of Geology 28: 524-535.
11. - 1922. New reptiles and stegocephalians from the Upper Triassic of western Texas. Carnegie Institution Publication 321: 1-84.
12. - 1932. A perfectly preserved segment of the armour of a phytosaur, with associated vertebrae. Contributions from the Museum of Paleontology, University of Michigan 4: 57-80.
13. Cope, E. D. 1869. Synopsis of the extinct Batrachia, Reptilia, and Aves of North America. Transactions of the American Philosophical Society, new series 14: 1-252.
14. - 1875. Report on the geology of that part of northwestern New Mexico examined during the field-season of 1874. In Annual Report upon the geographical explorations west of the 100th Meridian [Wheeler Survey], Appendix LL, Annual Report Chief of Engineers for 1875, pp. 61-97 of separate issue, 981-1017 of full report.
15. - 1887. A contribution to the history of the Vertebrata of the Trias of North America. Proceedings of the American Philosophical Society 24: 209-228.
16. - 1892. A contribution to the vertebrate paleontology of Texas. Proceedings of the American Philosophical Society 30: 123-131.
17. de Queiroz, K. & Gauthier, J. 1992. Phylogenetic taxonomy. Annual Review of Ecology and Systematics 23: 449-480.
18. Desojo, J. B. 2002. New data on the Late Triassic aetosaur Neoaetosauroides Bonaparte, 1969 from the Los Colorados Formation, west-central Argentina. Journal of Vertebrate Paleontology 22: 49A.
19. - 2003. Redescripción del aetosaurio Chilenosuchus forttae Casamiquela (Diapsida: Arcosauria): presencia de Triásico continental en el norte de Chile. Revista Geológica de Chile 30: 53-63.
20. - 2004. Los aetosaurias (Aniota, Diapsida) de America Del Sur: sus relaciones y aportes a la biogeografia y biostratigrafia del Triasico continental. PhD Thesis: Universidad de Buenos Aires, Buenos Aires, 176 pp.
21. - & Báez, A. M. 2005. El esqueleto postcraneano de Neoaetosauroides (Archosauria: Aetosauria) del Triásico Superior del centro-oeste de Argentina. Ameghiniana 42: 115-126.
22. - & Heckert, A. B. 2004. New information on the braincase and mandible of Coahomasuchus (Archosauria: Aetosauria) from the Otischalkian (Carnian) of Texas. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 2004: 605-616.
23. Elder, R. L. 1978. Paleontology and paleoecology of the Dockum Group, Upper Triassic, Howard County, Texas. MSc Thesis: University of Texas, Austin, 205 pp.
24. Fiorillo, A. R., Padian, K. & Musikasinthorn, C. 2000. Taphonomy and depositional setting of the Placerias quarry (Chinle Formation: Late Triassic, Arizona). Palaios 15: 373-386.
25. Fraas, O. 1877. Aëtosaurus ferratus Fr. Die gepanzerte Vogel-Esche aus dem Stubensandstein bei Stuttgart. Württembergische naturwissenschaftliche Jahreshefte 33: 1-22.
26. Gauthier, J. A. 1986. Saurischian monophyly and the origin of birds. Memoirs, California Academy of Sciences 8: 1-55.
27. Gower, D. J. & Walker, A. D. 2002. New data on the braincase of the aetosaurian archosaur (Reptilia: Diapsida) Stagonolepis robertsoni Agassiz. Zoological Journal of the Linnean Society 136: 7-23.
28. - & Wilkinson, M. 1996. Is there any consensus on basal archosaur phylogeny? Proceedings of the Royal Society of London, Series B 263: 1399-1406.
29. Gregory, J. T. 1953. Typothorax and Desmatosuchus. Postilla 16: 1-27.
30. - & Westphal, F. 1969. Remarks on the phytosaur genera of the European Trias. Journal of Paleontology 43: 1296-1298.
31. Harris, S. R., Gower, D. J. & Wilkinson, M. 2003a. Intraorganismal homology, character construction, and the phylogeny of aetosaurian archosaurs (Reptilia, Diapsida). Systematic Biology 52: 239-252.
32. -, - & - 2003b. Phylogenetic methods and aetosaur relationships: a rejoinder. Systematic Biology 52: 851-852.
33. Heckert, A. B. 1997. The tetrapod fauna of the Upper Triassic Lower Chinle Group (Adamanian: Latest Carnian) of the Zuni Mountains, west-central New Mexico. New Mexico Museum of Natural History and Science Bulletin 11: 29-39.
34. - & Jenkins, H. S. 2005. The microvertebrate fauna of the Upper Triassic (Revueltian) Snyder Quarry, North-central New Mexico. New Mexico Geological Society Guidebook 56: 319-334.
35. - & Lucas, S. G. 1999. A new aetosaur from the Upper Triassic of Texas and the phylogeny of aetosaurs. Journal of Vertebrate Paleontology 19: 50-68.
36. - & - 2000. Taxonomy, phylogeny, biostratigraphy, biochronology, paleobiogeography, and evolution of the Late Triassic Aetosauria (Archosauria: Crurotarsi). Zentralblatt für Geologie und Paläontologie, Teil I, Heft 11-12: 1539-1587.
37. - & - 2002a. South American occurrences of the Adamanian (Late Triassic, Latest Carnian) index taxon Stagonolepis (Archosauria: Aetosauria) and their biochronological significance. Journal of Paleontology 76: 852-863.
38. - & - 2002b. Acaenasuchus geoffreyi (Archosauria: Aetosauria) from the upper Triassic Chinle Group: juvenile of Desmatosuchus haplocerus. New Mexico Museum of Natural History and Science Bulletin 21: 205-214.
39. - & - 2003. Clarifying aetosaur phylogeny requires more fossils, not more trees: reply to intraorganisimal homology, character construction, and the phylogeny of aetosaurian archosaurs (Reptilia, Diapsida). Systematic Biology 52: 253-255.
40. - & Zeigler, K. E. 2002. The Late Triassic Snyder Quarry: a brief history of discovery and excavation. New Mexico Museum of Natural History Bulletin 24: 5-13.
41. -, Hunt, A. P. & Lucas, S. G. 1996. Redescription of Redondasuchus reseri, a late Triassic aetosaur (Reptilia: Archosauria) from New Mexico (U.S.A.) and the biochronology and phylogeny of aetosaurs. Geobios 29: 619-632.
42. -, Zeigler, K. E. & Lucas, S. G. 2003. Aetosaurs (Archosauria: Aetosauria) from the Upper Triassic (Revueltian) Snyder Quarry, New Mexico. New Mexico Museum of Natural History and Science Bulletin 24: 115-126.
43. -, Lucas, S. G. & Hunt, A. P. 2005a. Triassic vertebrates in Arizona. New Mexico Museum of Natural History and Science Bulletin 29: 16-44.
44. -, -, Sullivan, R. M., Hunt, A. P. & Speilmann, J. A. 2005b. The vertebrate fauna of the Upper Triassic (Revueltian: mid-Norian) Painted Desert Member (Petrified Forest Formation: Chinle Group) in the Chama Basin, northern New Mexico. New Mexico Geological Society Guidebook 56: 302-318.
45. Huene, F. von, 1920. Osteologie von Aëtosaurus ferratus O. Fraas. Acta Zoologica 1: 465-491.
46. -, 1942. Die fossilen Reptilien des südamerikanischen Gonwanalandes. Ergebnisse dur Sauriergrabung in Südbrasilien 1928/1929. C.H. Beck: München, pp. 161-332.
47. Hunt, A. P. 2001. The vertebrate fauna, biostratigraphy and biochronology of the type Revueltian faunachron, Bull Canyon Formation (Upper Triassic), east-central New Mexico. New Mexico Geological Society Guidebook 52: 123-152.
48. - & Lucas, S. G. 1991. A new aetosaur from the Redonda Formation (Late Triassic: middle Norian) of east-central New Mexico, U.S.A. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 12: 728-736.
49. -, - & Reser, P. K. 1993. A complete skeleton of the stagonolepidid Typothorax coccinarum from the Upper Triassic Bull Canyon Formation of east-central New Mexico, U.S.A. New Mexico Museum of Natural History and Science Bulletin 3: 209-212.
50. Huxley, T. H. 1859. On the Stagonolepis Robertsoni (Agassiz) of the Elgin Sandstone; and on the recently discovered footmarks in the sandstones of Cummingstone. Proceedings of the Geological Society (London) 15: 440-460.
51. ICZN (International Commission on Zoological Nomenclature). 1999. International Code of Zoological Nomenclature: (4th edition). International Trust for Zoological Nomenclature: London, 306 pp.
52. Jenkins, F. A. Jr., Shubin, N. H., Amaral, W. W., Gatesy, S. M., Schaff, C. R., Clemmensen, L. B., Downs, W. R., Davidson, A. R., Bonde, N. & Osbæck, F. 1994. Late Triassic continental vertebrates and depositional environments of the Fleming Fjord Formation, Jameson Land, East Greenland. Meddelelser om Grønland Geoscience 32: 1-25.
53. Kischlat, E. E. 2000. Tecodôncios: a aurora dos arcossáurios Triássico. Pp. 273-316 in Holz, M. & De Ros, L. F. (eds) Paleontologia do Rio Grande do Sul. Edição CIGO/UFRGS: Port Allegre, 398p.
54. Krebs, B. 1976. Pseudosuchia. Pp. 40-98 in Kuhn, O. (ed.) Handbuch der Paläoherpetologie, Volume 13. Gustav Fischer Verlag: Stuttgart.
55. Long, R. A. & Ballew, K. L. 1985. Aetosaur dermal armour from the late Triassic of southwestern North America, with special reference to material from the Chinle Formation of Petrified Forest National Park. Museum of Northern Arizona Bulletin 47: 45-68.
56. - & Murry, P. A. 1995. Late Triassic (Carnian and Norian) tetrapods from the southwestern United States. New Mexico Museum of Natural History and Science Bulletin 4: 1-254.
57. Lucas, S. G. 1993. The Chinle Group: revised stratigraphy and biochronology of Upper Triassic nonmarine strata in the western United States. Museum of Northern Arizona Bulletin 59: 27-50.
58. - 1998. Global Triassic tetrapod biostratigraphy and biochronology. Palaeogeography, Palaeoclimatology, Palaeoecology 143: 347-384.
59. - & Hunt, A. P. 1993. Field guide to nonmarine Triassic strata of the southern Colorado Plateau, New Mexico and Arizona. New Mexico Museum of Natural History and Science Bulletin 3: G1-G58.
60. -, Heckert, A. B. & Hunt, A. P. 1995. Unusual aetosaur armour from the Upper Triassic of west Texas, U.S.A. Paläontologische Zeitschrift 69: 467-473.
61. -, - & - 2002. A new species of the aetosaur Typothorax (Archosauria: Aetosauria) from the upper Triassic of east-central New Mexico. New Mexico Museum of Natural Science and History Bulletin 21: 221-233.
62. -, Zeigler, K. E., Heckert, A. B. & Hunt, A. P. 2005. Review of Upper Triassic stratigraphy and biostratigraphy in the Chama Basin, northern New Mexico. New Mexico Geological Society Guidebook 56: 170-181.
63. Lydekker, R. 1887. The fossil Vertebrata of India. Records of the Geological Survey of India 20: 51-80.
64. Marsh, O. C. 1884. The classification and affinities of dinosaurian reptiles. Nature 31: 68-69.
65. - 1896. A new belodont reptile (Stegomus) from the Connecticut River sandstone. American Journal of Science 4: 59.
66. Martz, J. W. 2002. The morphology and ontogeny of Typothorax coccinarum (Archosauria, Aetosauria) from the Upper Triassic of the American Southwest. MSc thesis: Texas Tech University, Lubbock, 279 pp.
67. - & Small, B. J. (2006). Tecovasuchus chatterjeei, a new aetosaur (Archosauria: Aetosauria) from the Tecovas Formation (Upper Triassic, Carnian) of Texas. Journal of Vertebrate Paleontology 26: 308-320.
68. -, Mueller, B. & Small, B. J. 2003. Two new aetosaurs (Archosauria, Stagonolepididiae) from the Upper Triassic of Texas and Colorado, and problems in aetosaur identification and taxonomy. Journal of Vertebrate Paleontology 23: 76A.
69. Meyer, H. von, 1861. Reptilien aus dem Stubensandstein des oberen Keupers. Palaeontographica 7: 253-346.
70. Padian, K., Horner, J. & Dhaliwal, J. 2004. Species recognition as the principle cause of bizarre structures in archosaurs. Journal of Vertebrate Paleontology 24: 100A.
71. Parker, W. G. 2003. Description of a new specimen of Desmatosuchus haplocerus from the Late Triassic of Northern Arizona. MSc Thesis: Northern Arizona University, Flagstaff, 315 pp.
72. - (2006). Faunal review of the Upper Triassic Chinle Formation of Arizona. Mesa Southwest Museum Bulletin 11: 34-54.
73. - & Irmis, R. B. 2005. Advances in Late Triassic vertebrate paleontology based on new material from Petrified Forest National Park, Arizona. New Mexico Museum of Natural History and Science Bulletin 29: 45-58.
74. -, Irmis, R. B., Nesbitt, S. N., Martz, J. W. & Browne, L. S. 2005. The Pseudosuchian Revueltosaurus callenderi and its implications for the diversity of early ornithischian dinosaurs. Proceedings of the Royal Society of London Series B 272: 963-969.
75. Parrish, J. M. 1993. Phylogeny of the Crocodylotarsi, with reference to archosaurian and crurotarsan monophyly. Journal of Vertebrate Paleontology 13: 287-308.
76. - 1994. Cranial osteology of Longosuchus meadei and the phylogeny and distribution of the Aetosauria. Journal of Vertebrate Paleontology 14: 196-209.
77. - & Carpenter, K. 1986. A new vertebrate fauna from the Dockum Formation (Late Triassic) of eastern New Mexico. Pp. 151-160 in Padian, K. (ed.) The Beginning of the Age of Dinosaurs. Cambridge University Press: Cambridge.
78. Sawin, H. J. 1947. The Pseudosuchian reptile Typothorax meadei. Journal of Paleontology 21: 201-238.
79. Sereno, P. C. 1991. Basal archosaurs: phylogenetic relationships and functional implications. Society of Vertebrate Paleontology Memoir 2: 1-53.
80. Small, B. J. 1985. The Triassic thecodontian reptile Desmatosuchus: osteology and relationships. MSc thesis: Texas Tech University, Lubbock, 83 pp.
81. - 2002. Cranial anatomy of Desmatosuchus haplocerus (Reptilia: Archosauria: Aetosauria). Zoological Journal of the Linnean Society 136: 97-111.
82. Sues, H.-D. 1992. A remarkable new armoured archosaur from the Upper Triassic of Virginia. Journal of Vertebrate Paleontology 12: 142-149.
83. Swofford, D. L. 2003. PAUP*. Phylogenetic Analysis Using Parsimony (*and Other Methods). Version 4. Sinauer Associates: Sunderland, Massachusetts.
84. Walker, A. D. 1961. Triassic reptiles from the Elgin area: Stagonolepis, Dasygnathus and their allies. Philosophical Transactions of the Royal Society of London, Series B 248: 103-204.
85. Weems, R. E. 1980. An unusual newly discovered archosaur from the Upper Triassic of Virginia, U.S.A. Transactions of the American Philosophical Society 70: 1-53.
86. Wild, R. 1989. Aëtosaurus (Reptilia: Thecodontia) from the Upper Triassic (Norian) of Cene near Bergamo, Italy, with a revision of the genus. Revista del Museo Civico di Scienze Naturali "Enrico Caffe" 14: 1-24.
87. Woody, D. T. 2003. Revised geological assessment of the Sonsela Member, Chinle Formation, Petrified Forest National Park, Arizona. MSc Thesis: Northern Arizona University, Flagstaff.
88. Zeigler, K. E. 2003. Taphonomic analysis of the Snyder Quarry: a fire-related Upper Triassic vertebrate fossil assemblage from North-Central New Mexico. New Mexico Museum of Natural History and Science Bulletin 24: 49-62.
89. -, Heckert, A. B. & Lucas, S. G. 2000. The aetosaur Desmatosuchus from the Upper Triassic Snyder quarry, northern New Mexico, and its biochronological significance. New Mexico Geology 22: 46.
90. -, - & - 2003a [for 2002]. A new species of Desmatosuchus (Archosauria: Aetosauria) from the upper Triassic of the Chama Basin, north-central New Mexico. New Mexico Museum of Natural History and Science Bulletin 21: 215-219.
91. -, - & - 2003b. The Upper Triassic (Norian: Revueltian) Snyder Quarry, Chama Basin, North-central New Mexico: an overview. New Mexico Museum of Natural History and Science Bulletin 24: 71-79.
92. -, - & - 2003c. The vertebrate fauna of theUpper Triassic (Revueltian) Snyder Quarry. New Mexico Museum of Natural History and Science Bulletin 24: 71-79.
93. -, - & - 2005. Taphonomic analysis of a fire-related Upper Triassic vertebrate fossil assemblage from north-central New Mexico. New Mexico Geological Society Guidebook 56: 341-354.
94. Zittel, K. A. 1887-90. Handbuch der Palaeontologie. 1. Abteilung: Palaeozoologie, 3. - XII + 900 S., 719 Abb. München & Leipzig.