A new protocetid whale (Cetacea: Archaeoceti) from the late middle Eocene of South Carolina

American Museum Novitates

Volumn , Issue 3480, 2005, Pages 1-65

  Geisler, Jonathan H ^a   Sanders, Albert E ^b   Luo, Zhe Xi ^c  

^a GEORGIA SOUTHERN UNIVERSITY   (United States)

^b Charleston Museum   (United States)

^c SECTION OF VERTEBRATE PALEONTOLOGY   (United States)

Author keywords

[No Author keywords available]

Indexed keywords

EID: 33746237861     PISSN: 00030082     EISSN: None     Source Type: Book Series    
DOI: 10.1206/0003-0082(2005)480[0001:ANPWCA]2.0.CO;2     Document Type: Article

References (120)

1. Albright, L.B. 1996. A protocetid cetacean from the Eocene of South Carolina. Journal of Paleontology 70: 519-523.
2. Bajpai, S., and J.G.M. Thewissen. 1998. Middle Eocene cetaceans from the Harudi and Subathu Formations of India. In J.G.M. Thewissen (editor), The emergence of whales, evolutionary patterns in the origin of Cetacea: 213-233. New York: Plenum Press.
3. Bajpai, S., and J.G.M. Thewissen. 2000. A new, diminuitive whale from Kachchh (Gujarat, India) and its implications for locomotor evolution of cetaceans. Current Science (New Delhi) 79: 1478-1482.
4. Bajpai, S., J.G.M. Thewissen, and A. Sahni. 1996. Indocetus (Cetacea, Mammalia) endocasts from Kachchh (India). Journal of Vertebrate Paleontology 16: 582-584.
5. Barnes, L.G., and E.D. Mitchell. 1978. Cetacea. In V.J. Maglio and H.B.S. Cooke (editors), Evolution of African mammals: 582-602. Cambridge, MA: Harvard University Press.
6. Barnes. L.G. 1984. Whales, dolphins, and porpoises: origin and evolution of Cetacea. In T.W. Broadhead (editor), Mammals. Notes for a short course organized by P.D. Gingerich and C.E. Badgley: 139-154. University of Tennessee Department of Geological Sciences. Studies in Geology 8(1-4).
7. Baum, G.R., J.S. Collins, R.M. Jones, B.A. Madlinger, and R.J. Powell. 1980. Correlation of the Eocene strata of the Carolinas. South Carolina Geology 24: 19-27.
8. Berggren, W.A., D.V. Kent, C.C. Swisher III, and M-P. Aubry. 1995. A revised Cenozoic geochronology and chronostratigraphy. In W.A. Berggren, D.V. Kent, and J.A. Hardenbol (editors), Geochronology, time scales and global stratigraphic correlation: 129-212. SEPM Society for Sedimentary Geology Special Publication No. 54.
9. Blow, W.C., and R.B. Manning. 1996. Preliminary descriptions of 25 new decapod crustaceans from the middle Eocene of the Carolinas, U.S.A. Tulane Studies in Geology and Paleontology 29: 1-26.
10. Breathnach, A.S. 1955. Observations on endocranial casts of recent and fossil cetaceans. Journal of Anatomy 89: 532-546.
11. Bremer, K. 1988. The limits of amino acid sequence data in angiosperm phylogenetic reconstruction. Evolution 42: 795-803.
12. Bremer, K. 1994. Branch support and tree stability. Cladistics 10: 295-304.
13. Buchholtz, E.A. 1998. Implications of vertebral morphology for locomotor evolution in early Cetacea. In J.G.M. Thewissen (editor), The emergence of whales, evolutionary patterns in the origin of Cetacea: 325-351. New York: Plenum Press.
14. Carte, A., A. MacAlister, and W.H. Flower. 1868. On the anatomy of Balaenoptera rostrata. Philosophical Transactions of the Royal Society of London 158: 201-262.
15. Cifelli, R.L. 1982. The petrosal structure of Hyopsodus with respect to that of some other ungulates, and its phylogenetic implications. Journal of Paleontology 56(3): 795-805.
16. Cooke, C.W. 1936. Geology of the coastal plain of South Carolina. U.S. Geological Survey Bulletin 867: 1-196.
17. Coombs, M.C., and W.P. Coombs, Jr. 1982. Anatomy of the ear region of four Eocene artiodactyls: Gobiohyus,? Helohyus, Diacodexis and Homacodon. Journal of Vertebrate Paleontology, 2: 219-236.
18. Cronin, T.R. 1999. Principles of paleoclimatology. New York: Columbia University Press.
19. Dart, R.A. 1923. The brain of the Zeuglodontidae (Cetacea). Proceedings of the Zoological Society of London 1923: 615-648.
20. Dennstedt, A. 1904. Die Sinus durae matris der Haussäugetiere. Anatomische Hefte 25(75): 1-96.
21. Dooley, A.C., Jr., N.C. Fraser, and Z.-X. Luo. 2004. The earliest member of the rorqual-gray whale clade (Mammalia, Cetacea). Journal of Vertebrate Paleontology 24: 453-463.
22. Dyce, K.M., W.O. Sack, and C.J.G. Wensing. 1996. Textbook of veterinary anatomy, 2nd ed. Philadelphia: WB Saunders.
23. Edwards, L.E., L.M. Bybell, G.S. Gohn, and N.O. Frederiksen. 1997. Paleontology and physical stratigraphy of the USGS-Pregnall no. 1 core (DOR-208), Dorchester County, South Carolina. USGS Open-file Report 97-145: 1-35.
24. Feldmann, R.M., K.L. Bice, C.S. Hopkins, E.W. Salva, and K. Pickford. 1998. Decapod crustaceans from the Eocene Castle Hayne Limestone, North Carolina: paleoceanographic implications. Journal of Paleontology, The Paleontological Society Memoir, 48: 1-28.
25. Fish, F.E. 1998. Biochemical perspective on the origin of cetacean flukes. In J.G.M. Thewissen (editor), The emergence of whales, evolutionary patterns in the origin of Cetacea: 303-324. New York: Plenum Press.
26. Fordyce, R.E. 1994. Waipatia maerewhenua, new genus and new species (Waipatiidae, new family), an archaic late Oligocene dolphin (Cetacea: Odontoceti: Platanistoidea) from New Zealand. In A. Berta and T. Deméré (editors), Contributions in marine mammal paleontology honoring Frank C. Whitmore, Jr.: 147-176. Proceedings of the San Diego Society of Natural History 29.
27. Fraas, E. 1904. Neue zeuglodonten aus dem unteren Mitteleocän vom Mokattam bei Cairo. Geologische und Palaeontologische Abhandlungen, Neue Folge 6(3): 199-220.
28. Fraser, F.C.., and P.E. Purves. 1960. Hearing in cetaceans: evolution of the acessory air sacs and the structure and function of the outer and middle ear in recent cetaceans. Bulletin of the British Museum (Natural History), Zoology 7: 1-140.
29. Gatesy, J. 1997. More DNA support for a Cetacea/Hippopotamidae clade: the blood-clotting protein gamma-fibrinogen. Molecular Biology and Evolution 14: 537-543.
30. Gatesy, J., C. Hayashi, A. Cronin, and P. Arctander. 1996. Evidence from milk casein genes that cetaceans are close relatives of hippopotamid artiodactyls. Molecular Biology and Evolution 13: 954-963.
31. Gatesy, J., M. Milinkovitch, V. Waddell, and M. Stanhope. 1999. Stability of cladistic relationships between Cetacea and higher-level artiodactyl taxa. Systematic Biology 48: 6-20.
32. Geisler, J. 2001. New morphological evidence for the phylogeny of Artiodactyla, Cetacea, and Mesonychidae. American Museum Novitates 3344: 1-53.
33. Geisler, J.H., and Z.-X. Luo. 1996. The petrosal and inner ear of Herpetocetus sp. (Mammalia: Cetacea) and their implications for the phylogeny and hearing of archaic mysticetes. Journal of Paleontology 70: 1045-1066.
34. Geisler, J.H., and Z.-X. Luo. 1998. Relationships of Cetacea to terrestrial ungulates and the evolution of cranial vasculature in Cete. In J.G.M. Thewissen (editor), The emergence of whales, evolutionary patterns in the origin of Cetacea: 163-212. New York: Plenum Press.
35. Geisler, J.H., A.E. Sanders, and Z.-X. Luo. 1996. A new protocetid cetacean from the Eocene of South Carolina, U.S.A.; phylogenetic and biogeographic implications. In J.E. Repetski (editor), Sixth North American Paleontological Convention abstracts of papers: 139. Paleontological Society Special Papers 8.
36. Geisler, J.H., and M.D. Uhen. 2003. Morphological support for a close relationship between hippos and whales. Journal of Vertebrate Paleontology 23: 991-996.
37. Getty, R. 1975. The anatomy of the domestic animals, 5th ed. Philadelphia: W.B. Saunders.
38. Gingerich, P.D. 1992. Marine mammals (Cetacea and Sirenia) from the Eocene of Gebel Mokattum and Fayum, Egypt: stratigraphy, age, and paleoenvironments. University of Michigan Papers of Paleontology 30: 1-84.
39. Gingerich, P.D. 2003. Land-to-sea transition of early whales: evolution of Eocene Archaeoceti (Cetacea) in relation to skeletal proportions and locomotion of living semiaquatic mammals. Paleobiology 29: 429-454.
40. Gingerich, P.D., M. Arif, M.A. Bhatti, H.A. Raza, and S.M. Raza. 1995a. Protosiren and Babiacetus (Mammalia, Sirenia and Cetacea) from the middle Eocene Drazida Formation, Sulaiman Range, Punjab (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 29: 331-357.
41. Gingerich, P.D., M. Arif, and W.C. Clyde. 1995b. New archaeocetes (Mammalia, Cetacea) from the middle Eocene Domanda Formation of the Sulaiman Range, Punjab (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 29: 291-330.
42. Gingerich, P.D., M. Arif, M.A. Bhatti, M. Anwar, and W.J. Sanders. 1997. Basilosaurus drazindai and Basiloterus hussaini, new Archaeoceti (Mammalia, Cetacea) from the middle Eocene Drazinda Formation, with a revised interpretation of ages of whale-bearing strata in the Kirthar Group of the Sulaiman Range, Punjab (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 30: 55-81.
43. Gingerich, P.D., S.M. Raza, M. Arif, M. Anwar, and X. Zhou. 1993. Partial skeletons of Indocetus ramani (Mammalia, Cetacea) from the lower middle Eocene Domanda Shale in the Sulaiman Range of Punjab (Pakistan). Contributions from the Museum of Paleontology, University of Michigan, 28: 393-416.
44. Gingerich, P.D., S.M. Raza, M. Arif, M. Anwar, and X. Zhou. 1994. New whale from the Eocene of Pakistan and the origin of cetacean swimming. Nature 368: 844-847.
45. Gingerich, P.D., and D.E. Russell. 1981. Pakicetus inachus a new archaeocete (Mammalia, Cetacea) from the early-middle Eocene Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 25: 235-246.
46. Gingerich, P.D., and D.E. Russell. 1990. Dentition of early Eocene Pakicetus (Mammalia, Cetacea). Contributions from the Museum of Paleontology, University of Michigan 28: 1-20.
47. Gingerich, P.D., D.E. Russell, and S.M.I. Shah. 1983. Origin of whales in epicontinental remnant seas: new evidence from the early Eocene of Pakistan. Science 220: 403-406.
48. Gingerich, P.D., B.H. Smith, and E.L. Simons. 1990. Hind limbs of Eocene Basilosaurus: evidence of feet in whales. Science 249: 154-157.
49. Gingerich, P.D., M. ul-Haq, I.S. Zalmout, I.H. Khan, and M.S. Malkani. 2001a. Origin of whales from early artiodactyls: hands and feet of Eocene Protocetidae from Pakistan. Science 293: 2239-2242.
50. Gingerich, P.D., M. ul-Haq, I.H. Khan, and I.S. Zalmout. 2001b. Eocene stratigraphy and archaeocete whales (Mammalia, Cetacea) of Drug Lahar in the eastern Sulaiman Range, Balochistan (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 30: 269-319.
51. Harris, W.B., V.A. Zullo, and R.A. Laws. 1993. Coastal onlap stratigraphy of the onshore Paleogene, southeastern Atlantic Coastal Plain, U.S.A. In H.W. Posamentier, C.P. Summerhayes, B.U. Haq, and G.P. Allen (editors), Sequence stratigraphy and facies associations: 537-561. International Association of Sedimentologists, Special Publication 18.
52. Hendy, M.D., and D. Penny. 1982. Branch and bound algorithms to determine minimal evolutionary trees. Mathematical Biosciences 59: 277-290.
53. Howell, A.B. 1927. Contribution to the anatomy of the Chinese finless porpoise, Neomeris phocaenoides. Proceedings of the United States National Museum 70(13): 1-43.
54. Hulbert, R.C., Jr. 1998. Postcranial osteology of the North American Middle Eocene protocetid Georgiacetus. In J.G.M. Thewissen (editor), The emergence of whales, evolutionary patterns in the origin of Cetacea: 235-267. New York: Plenum Press.
55. Hulbert, R.C., Jr., R.M. Petkewich, G.A. Bishop, D. Bukry, and D.P. Aleshire. 1998. A new middle Eocene protocetid whale (Mammalia: Cetacea: Archaeoceti) and associated biota from Georgia. Journal of Paleontology 72: 905-925.
56. Kasuya, T. 1973. Systematic consideration of recent toothed whales based on the morphology of tympano-periotic bone. Scientific Reports of the Whale Research Institute 25: 1-103.
57. Kellogg, R. 1936. A review of the Archaeoceti. Carnegie Institute of Washington Publication 482.
58. Köhler, R., and R.E. Fordyce. 1997. An archaeocete whale (Cetacea: Archaeoceti) from the Eocene Waihao Greensand, New Zealand. Journal of Vertebrate Paleontology 17: 574-583.
59. Kumar, K., and A. Sahni. 1986. Remingtonocetus harudiensis, new combination, a middle Eocene archaeocete (Mammalia, Cetacea) from western Kutch, India. Journal of Vertebrate Paleontology 6: 326-349.
60. Luo, Z.-X. 1998. Homology and transformation of cetacean ectotypmpanic structures. In J.G.M. Thewissen (editor), The emergence of whales, evolutionary patterns in the origin of Cetacea: 269-301. New York: Plenum Press.
61. Luo, Z.-X., and E.R. Eastman. 1995. Petrosal and inner ear of a squalodontoid whale: implications for the evolution of hearing in odontocetes. Journal of Vertebrate Paleontology 15: 431-442.
62. Luo, Z.-X., and P.D. Gingerich. 1999. Terrestrial Mesonychia to aquatic Cetacea: transformation of the basicranium and evolution of hearing in whales. University of Michigan Papers in Paleontology 31: 1-98.
63. Luo, Z.-X., and K. Marsh. 1996. The petrosal (periotic) and inner ear of a Pliocene kogiine whale (Kogiinae, Odontoceti): implications on relationships and hearing evolution of toothed whales. Journal of Vertebrate Paleontology 16: 328-348.
64. Lyell, C. 1845. On the White Limestone of South Carolina and Georgia, and the Eocene strata of other parts of the U.S., with appendix, on the corals, by Mr. Lonsdale. Proceedings of the Geological Society of London 4(1843-1845): 563-576.
65. McLeod, S.A., and L.G. Barnes. 1996. The systematic position of Pappocetus lugardi and a new taxon from North America (Archaeoceti: Protocetidae). In J.E. Repetski (editor), Sixth North American Paleontological Convention abstracts of papers: 270. Paleontological Society Special Papers 8.
66. MacPhee, R.D.E. 1981. Auditory regions of primates and eutherian insectivores. Contributions to Primatology 18: 1-282.
67. MacPhee, R.D.E. 1994. Morphology, adaptations, and relationships of Plesiorycteropus, and a diagnosis of a new order of eutherian mammals. Bulletin of the American Museum of Natural History 220: 1-214.
68. Nikaido, M., A.P. Rooney, and N. Okada. 1999. Phylogenetic relationships among cetartiodactyls based on insertions of short and long interspersed elements: hippopotamuses are the closest extant relatives of whales. Proceedings of the National Academy of Sciences U.S.A. 96: 10261-10266.
69. Norris, K.S. 1980. Peripheral sound processing in odontocetes. In R.-G. Busnel and J.F. Fish (editors), Animal sonar systems: 495-509. New York: Plenum Press.
70. Novacek, M.J. 1986. The skull of leptictid insectivorans and the higher-level classification of eutherian mammals. Bulletin of the American Museum of Natural History 183: 1-111.
71. Nummela, S., J.G.M. Thewissen, S. Bajpai, S.T. Hussain, and K. Kumar. 2004. Eocene evolution of whale hearing. Nature 430: 776-778.
72. O'Leary, M.A., and J.H. Geisler. 1999. The position of Cetacea within Mammalia: phylogenetic analysis of morphological data from extinct and extant taxa. Systematic Biology 48: 455-490.
73. O'Leary, M.A., and K.D. Rose. 1995. Postcranial skeleton of the early Eocene mesonychid Pachyaena (Mammalia: Mesonychia). Journal of Vertebrate Paleontology 15: 401-430.
74. Padget, D.H. 1957. The development of the cranial venous system in man, from the viewpoint of comparative anatomy. Contributions to Embryology, Carnegie Institution 36: 81-153.
75. Pompeckj, J.F. 1922. Das Ohrskelett von Zeuglodon. Senckenbergiana 4: 44-100.
76. Prothero, D.R., E.M. Manning, and M. Fischer. 1988. The phylogeny of the ungulates. In M.J. Benton (editor), The phylogeny and classification of the tetrapods, mammals: 201-234. Systematic Association Special Volume 35B, vol. 2.Oxford: Clarendon Press.
77. Radinsky, L.B. 1965. Evolution of the tapiroid skeleton from Heptodon to Tapirus. Bulletin of the Museum of Comparative Zoology 134(3): 69-106.
78. Reid, M.S., W.R. Aucott, R.W. Lee, and R.A. Renken. 1986. Hydrologic and geologic analysis of a well in Dorchester County, South Carolina. U.S. Geological Survey Water-Resources Investigations Report 86-4161.
79. Rose, K.D. 1982. Skeleton of Diacodexis, oldest known artiodactyl. Science 216: 621-623.
80. Rose, K.D. 1985. Comparative osteology of North American dichobunid artiodactyls. Journal of Paleontology 59: 1203-1226.
81. Sanders, A.E., and L.G. Barnes. 2003 (2002). Paleontology of the late Oligocene Chandler Bridge and Ashley formations of South Carolina, 3: Eomysticetidae, a new family of primitive mysticetes (Mammalia: Cetacea). In R. Emry (editor), Cenezoic mammals of land and sea: tributes to the career of Clayton E. Ray. Smithsonian Contributions to Paleobiology 93: 313-356.
82. Sanders, A.E., and M.P. Katuna. 2000. Proposal for revision of the stratigraphie nomenclature of the Santee Limestone (Eocene) of South Carolina. Abstracts with Programs, Geological Society of America, 49th Annual Meeting, South-eastern Section, March 2000, 32(2): A-71.
83. Sanders, J.E., A.H. Anderson, Jr., and R. Carola. 1976. Physical geology. New York: Harper and Row.
84. Schulte, H. von. 1916. Anatomy of a fetus of Balaenoptera borealis. Memoirs of the American Museum of Natural History 1: 389-502.
85. Schulte, H. von, and M. de Forest Smith. 1918. The external characters, skeletal muscles, and peripheral nerves of Kogia breviceps (Blainville). Bulletin of the American Museum of Natural History 38: 7-72.
86. Shimamura, M., H. Yasue, K. Ohshima, H. Abe, H. Kato, T. Kishiro, M. Goto, I. Munechika, and N. Okada. 1997. Molecular evidence from retroposons that whales form a clade within even-toed ungulates. Nature 388: 666-670.
87. Sisson, S. 1921. The anatomy of the domesticated animals. Philadelphia: W.B. Saunders.
88. Slijper, E.J. 1936. Die cetacean vergleichend-anatomisch und systematisch. Capita Zoologica 7: 1-590.
89. Sloan, E. 1908. Catalogue of the mineral localities of South Carolina. South Carolina Geological Survey Bulletin 2, series 4: 1-505.
90. Smith, A.G., D.G. Smith, and B.M. Funnell. 1994. Atlas of Mesozoic and Cenozoic coastlines. Cambridge: Cambridge University Press.
91. Sorenson, M.D. 1996. TreeRot. University of Michigan, Ann Arbor. Can be downloaded from http://people.bu.edu/msoren/TreeRot.html.
92. Spoor, F., S. Bajpai, S.T. Hussain, K. Kumar, and J.G.M. Thewissen. 2002. Vestibular evidence for the evolution of aquatic behaviour in early cetaceans. Nature 417: 163-166.
93. Stromer, E. 1903. Zeuglodon-reste aus dem Oberen Mittleocän des Fayum. Beiträge zur Paläntologie und Geologie Österreich-Ungarns und des Orients 15: 65-100.
94. Stromer, E. 1908. Die Archaeoceti des ägyptiscen Eozäns. Beiträge zur Paläontologie und Geologie Österreich-Ungarns und des Orients 21: 106-178.
95. Swofford, D.L. 2002. PAUP*, Phylogenetic analysis using parsimony (* and other methods). Sinauer Associates, vers. 4.0b10. Available at http://paup.csit.fsu.edu/.
96. Thewissen, J.G.M. 1994. Phylogenetic aspects of cetacean origins: a morphological perspective. Journal of Mammalian Evolution 2: 157-184.
97. Thewissen, J.G.M., and D.P. Domning. 1992. The role of phenacodontids in the origin of the modern orders of ungulate mammals. Journal of Vertebrate Paleontology 12(4): 494-504.
98. Thewissen, J.G.M., and S.T. Hussain. 1993. Origin of underwater hearing in whales. Nature 361:444-445.
99. Thewissen, J.G.M., and S.T. Hussain. 2000. Attockicetus praecursor, a new remingtonocetid cetacean from marine Eocene sediments of Pakistan. Journal of Mammalian Evolution 7: 133-146.
100. Thewissen, J.G.M., S.T. Hussain, and M. Arif. 1994. Fossil evidence for the origin of aquatic locomotion in archaeocete whales. Science 263: 210-212.
101. Thewissen, J.G.M., S.I. Madar, and S.T. Hussain. 1996. Ambulocetus natans, an Eocene cetacean (Mammalia) from Pakistan. Courier Forschungsinstitut Senckenberg 191: 1-86.
102. Thewissen, J.G.M., D.E. Russell, P.D. Gingerich, and S.T. Hussain. 1983. A new dichobunid artiodactyl (Mammalia) from the Eocene of north-west Pakistan. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen, Series B, 86(2): 153-180.
103. Thewissen, J.G.M., E.M. Williams, L.J. Roe, and S.T. Hussain. 2001a. Skeletons of terrestrial cetaceans and the relationship of whales to artiodactyls. Nature 413: 277-281.
104. Uhen, M.D. 1998a. Middle to late Eocene basilosaurines and dorudontines. In J.G.M. Thewissen (editor), The emergence of whales, evolutionary patterns in the origin of Cetacea: 29-61.New York: Plenum Press.
105. Uhen, M.D. 1998b. New protocetid (Mammalia: Cetacea) from the late middle Eocene Cook Mountain Formation of Louisiana. Journal of Vertebrate Paleontology 18(3): 664-668.
106. Uhen, M.D. 1999. New species of protocetid archaeocete whale, Eocetus wardii (Mammalia: Cetacea) from the middle Eocene of North Carolina. Journal of Paleontology 73(3): 512-528.
107. Uhen, M.D. 2001. New material of Eocetus wardii (Mammalia: Cetacea) from the middle Eocene of North Carolina. Southeastern Geology 40: 135-148.
108. Uhen, M.D., and G. Tichy. 2000. A new basilosaurid archaeocete from Austria. Journal of Vertebrate Paleontology 20(suppl.): 74A-75A.
109. Vogl, A.W., and H.D. Fisher. 1981. Arterial circulation of the spinal cord and brain in the Monodontidae (Order Cetacea). Journal of Morphology 170: 171-180.
110. Vogl, A.W., and H.D. Fisher. 1981. The internal carotid artery does not directly supply the brain in the Monodontidae (Order Cetacea). Journal of Morphology 170: 207-214.
111. Ward, L.W., B.W. Blackwelder, G.S. Gohn, and R.Z. Poore. 1979. Stratigraphic revision of Eocene, Oligocene, and lower Miocene formations of South Carolina. Geologic Notes 23: 2-23.
112. Weems, R.W., J.M. Self-Trail, and L.E. Edwards. 2004. Supergroup stratigraphy of the Atlantic and Gulf coastal plains (Middle? Jurassic through Holocene, eastern North America). Southeastern Geology 42: 191-216.
113. Wible, J.R. 1984. The ontogeny and phylogeny of the mammalian cranial arterial system. Ph.D. dissertation, Duke University, Durham, North Carolina, 705 pp.
114. Wible, J.R. 1986. Transformations in the extracranial course of the internal carotid artery in mammalian phylogeny. Journal of Vertebrate Paleontology 6: 313-325.
115. Wible, J.R. 1987. The eutherian stapedial artery: character analysis and implications for superordinal relationships. Zoological Journal of the Linnean Society 91: 107-135.
116. Wible, J.R. 1990. Petrosals of late Cretaceous marsupials from North America, and a cladistic analysis of the petrosal in then an mammals. Journal of Vertebrate Paleontology 10: 183-205.
117. Wilkinson, M. 1992. Ordered versus unordered characters. Cladistics 8: 375-385.
118. Woodburne, M.O. 1968. The cranial myology and osteology of Dicotyles tajacu, the collared peccary, and its bearing on classification. Memoirs of the Southern California Academy of Sciences 7: 1-48.
119. Zhou, X., W.J. Sanders, and P.D. Gingerich. 1992. Functional and behavioral implications of vertebral structure in Pachyaena ossifraga (Mammalia, Mesonychia). Contributions from the Museum of Paleontology, University of Michigan 28: 289-319.
120. Zhou, X., R. Zhai, and P.D. Gingerich. 1995. Skull of a new mesonychid (Mammalia, Mesonychia) from the late Paleocene of China. Journal of Vertebrate Paleontology 15: 387-400.