메뉴 건너뛰기




Volumn 18, Issue 3, 2004, Pages 321-348

The basal phylogeny of Scarabaeoidea (Insecta:Coleoptera) inferred from larval morphology

Author keywords

Bolboceratidae; Ceratocanthidae; Geotrupidae; Glaphyridae; Hybosoridae; Lucanidae; Passalidae; Pleocomidae; Scarabaeidae; Trogidae

Indexed keywords

PHYLOGENY;

EID: 3142623179     PISSN: 14455226     EISSN: None     Source Type: Journal    
DOI: 10.1071/IS03013     Document Type: Article
Times cited : (65)

References (137)
  • 1
    • 0000342477 scopus 로고    scopus 로고
    • A study of the larva of Carabdytes upin Balke, Hendrich and Wewalka (Coleoptera: Adephaga: Dytiscidae), with comments on the phylogeny of the Colymbetinae
    • Alarie, Y., and Balke, M. (1999). A study of the larva of Carabdytes upin Balke, Hendrich and Wewalka (Coleoptera: Adephaga: Dytiscidae), with comments on the phylogeny of the Colymbetinae. The Coleopterists Bulletin 53, 146-154.
    • (1999) The Coleopterists Bulletin , vol.53 , pp. 146-154
    • Alarie, Y.1    Balke, M.2
  • 2
    • 84981853002 scopus 로고
    • Revision of the Australian species of Phaenognatha Hope (Coleoptera: Scarabaeidae: Aclopinae)
    • Allsopp, P. G. (1981). Revision of the Australian species of Phaenognatha Hope (Coleoptera: Scarabaeidae: Aclopinae). Journal of the Australian Entomological Society 20, 185-195.
    • (1981) Journal of the Australian Entomological Society , vol.20 , pp. 185-195
    • Allsopp, P.G.1
  • 3
    • 3142548701 scopus 로고
    • Neophaenognatha, a new genus for the Neotropical species of Phaenognatha Hope (Scarabaeidae: Aclopinae) with the description of N. capella n. sp., and designation of lectotypes
    • Allsopp, P. G. (1983). Neophaenognatha, a new genus for the Neotropical species of Phaenognatha Hope (Scarabaeidae: Aclopinae) with the description of N. capella n. sp., and designation of lectotypes. The Coleopterists Bulletin 37, 208-211.
    • (1983) The Coleopterists Bulletin , vol.37 , pp. 208-211
    • Allsopp, P.G.1
  • 4
    • 0242541878 scopus 로고
    • Checklist of the Hybosorinae (Coleoptera: Scarabaeidae)
    • Allsopp, P. G. (1984). Checklist of the Hybosorinae (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 38, 105-117.
    • (1984) The Coleopterists Bulletin , vol.38 , pp. 105-117
    • Allsopp, P.G.1
  • 5
    • 3142519494 scopus 로고
    • Il Pachypus caesus Erichson 1840: Brevi note sulla biologia e la distribuzione in Sicilia (Coleoptera: Scarabaeoidea)
    • Arnone, M., and Sparacio, I. (1990). Il Pachypus caesus Erichson 1840: brevi note sulla biologia e la distribuzione in Sicilia (Coleoptera: Scarabaeoidea). Naturalista Siciliano 14, 63-71.
    • (1990) Naturalista Siciliano , vol.14 , pp. 63-71
    • Arnone, M.1    Sparacio, I.2
  • 6
    • 0001678678 scopus 로고
    • Larval chaetotaxy of Aleocharinae (Staphylinoidea) based on a description of Atheta coriarica Kraatz
    • Ashe, J. S., and Watrous, L. E. (1984). Larval chaetotaxy of Aleocharinae (Staphylinoidea) based on a description of Atheta coriarica Kraatz. The Coleopterists Bulletin 38, 165-179.
    • (1984) The Coleopterists Bulletin , vol.38 , pp. 165-179
    • Ashe, J.S.1    Watrous, L.E.2
  • 7
    • 0011604533 scopus 로고
    • Larval taxonomy of the Trogidae in North America with notes on biologies and life histories (Coleoptera: Scarabaeidae)
    • Baker, C. W. (1968). Larval taxonomy of the Trogidae in North America with notes on biologies and life histories (Coleoptera: Scarabaeidae). United States National Museum Bulletin 279, 1-79.
    • (1968) United States National Museum Bulletin , vol.279 , pp. 1-79
    • Baker, C.W.1
  • 8
    • 0011599295 scopus 로고    scopus 로고
    • A new genus of Ceratocanthidae from the Oriental Region (Coleoptera: Scarabaeoidea)
    • Ballerio, A. (1999). A new genus of Ceratocanthidae from the Oriental Region (Coleoptera: Scarabaeoidea). Elytron 13, 149-164.
    • (1999) Elytron , vol.13 , pp. 149-164
    • Ballerio, A.1
  • 9
    • 0001429555 scopus 로고    scopus 로고
    • A new genus and species of Ceratocanthidae from Tanzania (Coleoptera: Scarabaeoidea)
    • Ballerio, A. (2000a). A new genus and species of Ceratocanthidae from Tanzania (Coleoptera: Scarabaeoidea). African Zoology 35, 131-137.
    • (2000) African Zoology , vol.35 , pp. 131-137
    • Ballerio, A.1
  • 10
    • 0034345260 scopus 로고    scopus 로고
    • Revision of the genus Ebbrittoniella Matrtínez (Coleoptera: Scarabaeoidea: Ceratocanthidae)
    • Ballerio, A. (2000b). Revision of the genus Ebbrittoniella Matrtínez (Coleoptera: Scarabaeoidea: Ceratocanthidae). Revue Suisse de Zoologie 107, 259-275.
    • (2000) Revue Suisse de Zoologie , vol.107 , pp. 259-275
    • Ballerio, A.1
  • 11
    • 0345181875 scopus 로고
    • Révision du genre Pygopleurus Motschulsky (Coleoptera: Scarabaeoidea: Glaphyridae)
    • Baraud, J. (1989). Révision du genre Pygopleurus Motschulsky (Coleoptera: Scarabaeoidea: Glaphyridae). Annales de la Société Entomologique de France (N.S.) 25, 331-375.
    • (1989) Annales de la Société Entomologique de France (N.S.) , vol.25 , pp. 331-375
    • Baraud, J.1
  • 12
    • 3142626193 scopus 로고
    • Révision du genre Hybalus Brullé, 1834 (Coleoptera: Scarabaeoidea: Orphninae)
    • Baraud, J. (1991). Révision du genre Hybalus Brullé, 1834 (Coleoptera: Scarabaeoidea: Orphninae). Annales de la Société Entomologique de France (N.S.) 27, 3-28.
    • (1991) Annales de la Société Entomologique de France (N.S.) , vol.27 , pp. 3-28
    • Baraud, J.1
  • 13
    • 0011609754 scopus 로고
    • Chaetonyx robustus ssp. liguricus Mariani, 1946: Descrizione del terzo stadio larvale e note sulla morfologia preimmaginale della famiglia (Coleoptera: Scarabaeoidea: Orphnidae)
    • Barbero, E., and Palestrini, C. (1993). Chaetonyx robustus ssp. liguricus Mariani, 1946: descrizione del terzo stadio larvale e note sulla morfologia preimmaginale della famiglia (Coleoptera: Scarabaeoidea: Orphnidae). Bollettino della Società Entomologica Italiana 125, 143-149.
    • (1993) Bollettino Della Società Entomologica Italiana , vol.125 , pp. 143-149
    • Barbero, E.1    Palestrini, C.2
  • 14
    • 0034480303 scopus 로고    scopus 로고
    • Phylogenetic relationships of the suborders of Coleoptera (Insecta)
    • doi:10.1006/CLAD.1999.0124
    • Beutel, R. G., and Haas, F. (2000). Phylogenetic relationships of the suborders of Coleoptera (Insecta). Cladistics 16, 103-141. doi:10.1006/CLAD. 1999.0124
    • (2000) Cladistics , vol.16 , pp. 103-141
    • Beutel, R.G.1    Haas, F.2
  • 15
    • 0000391301 scopus 로고
    • Notation of primary setae and pores on larvae of Carabidae (Coleoptera: Adephaga)
    • Bousquet, Y., and Goulet, H. (1984). Notation of primary setae and pores on larvae of Carabidae (Coleoptera: Adephaga). Canadian Journal of Zoology 62, 573-588.
    • (1984) Canadian Journal of Zoology , vol.62 , pp. 573-588
    • Bousquet, Y.1    Goulet, H.2
  • 16
    • 0008929980 scopus 로고
    • On the classification of beetles according to larval characters
    • Böving, A. G. (1929). On the classification of beetles according to larval characters. Bulletin of the Brooklyn Entomological Society 24, 55-80.
    • (1929) Bulletin of the Brooklyn Entomological Society , vol.24 , pp. 55-80
    • Böving, A.G.1
  • 17
    • 0001708459 scopus 로고
    • Description of the larva of Plectris aliena Chapin and explanation of new terms applied to the epipharynx and raster
    • Böving, A. G. (1936). Description of the larva of Plectris aliena Chapin and explanation of new terms applied to the epipharynx and raster. Proceedings of the Entomological Society of Washington 38, 169-185.
    • (1936) Proceedings of the Entomological Society of Washington , vol.38 , pp. 169-185
    • Böving, A.G.1
  • 18
    • 0001900143 scopus 로고
    • An illustrated synopsis of the principal larval forms of the order Coleoptera
    • Böving, A. G., and Craighead, F. C. (1931). An illustrated synopsis of the principal larval forms of the order Coleoptera. Entomologica Americana (N.S.) 54, 1-351.
    • (1931) Entomologica Americana (N.S.) , vol.54 , pp. 1-351
    • Böving, A.G.1    Craighead, F.C.2
  • 19
    • 84981655114 scopus 로고
    • Phylogeny of the families of Scarabaeoidea based on characters of the hindwing articulation, hindwing base and wing venation
    • Browne, J., and Scholtz, C. H. (1995). Phylogeny of the families of Scarabaeoidea based on characters of the hindwing articulation, hindwing base and wing venation. Systematic Entomology 20, 145-173.
    • (1995) Systematic Entomology , vol.20 , pp. 145-173
    • Browne, J.1    Scholtz, C.H.2
  • 20
    • 0001273309 scopus 로고    scopus 로고
    • Evolution of the scarab hindwing articulation and wing base: A contribution towards the phylogeny of the Scarabaeidae. (Coleoptera: Scarabaeoidea)
    • doi:10.1046/J.1365-3113. 1998.00059.X
    • Browne, J., and Scholtz, C. H. (1998). Evolution of the scarab hindwing articulation and wing base: a contribution towards the phylogeny of the Scarabaeidae. (Coleoptera: Scarabaeoidea). Systematic Entomology 23, 307-326. doi:10.1046/J.1365-3113. 1998.00059.X
    • (1998) Systematic Entomology , vol.23 , pp. 307-326
    • Browne, J.1    Scholtz, C.H.2
  • 21
    • 0002793480 scopus 로고    scopus 로고
    • A phylogeny of the families of Scarabaeoidea (Coleoptera)
    • doi:10.1046/J.1365-3113.1999.00067.X
    • Browne, J., and Scholtz, C. H. (1999). A phylogeny of the families of Scarabaeoidea (Coleoptera). Systematic Entomology 24, 51-84. doi:10.1046/J.1365-3113.1999.00067.X
    • (1999) Systematic Entomology , vol.24 , pp. 51-84
    • Browne, J.1    Scholtz, C.H.2
  • 22
    • 85056066716 scopus 로고    scopus 로고
    • Glaphyridae MacLeay 1819
    • Eds R. H Arnett, M. C. Thomas, P. E. Skelley, and J. H. Frank. CRC Press: Boca Raton
    • Carlson, D. C. (2002). Glaphyridae MacLeay 1819. In 'American Beetles. Vol. 2'. (Eds R. H Arnett, M. C. Thomas, P. E. Skelley, and J. H. Frank.) pp. 37-38. (CRC Press: Boca Raton.)
    • (2002) American Beetles , vol.2 , pp. 37-38
    • Carlson, D.C.1
  • 23
    • 0011606223 scopus 로고
    • A new genus of Ochodaeinae and a description of the larva of Pseudochodaeus estriatus (Schaeffer)
    • Carlson, D. C., and Ritcher, P. O. (1974). A new genus of Ochodaeinae and a description of the larva of Pseudochodaeus estriatus (Schaeffer). The Pan-Pacific Entomologist 50, 99-110.
    • (1974) The Pan-Pacific Entomologist , vol.50 , pp. 99-110
    • Carlson, D.C.1    Ritcher, P.O.2
  • 24
    • 0001111484 scopus 로고
    • The phylogenetic significance of ommatidium structure in the compound eyes of Polyphagan beetles
    • Caveney, S. (1986). The phylogenetic significance of ommatidium structure in the compound eyes of Polyphagan beetles. Canadian Journal of Zoology 64, 1787-1819.
    • (1986) Canadian Journal of Zoology , vol.64 , pp. 1787-1819
    • Caveney, S.1
  • 25
    • 3142524926 scopus 로고
    • The nomenclature and taxonomy of the genera of the scarabaeid subfamily Glaphyrinae
    • Chapin, E. A. (1938). The nomenclature and taxonomy of the genera of the scarabaeid subfamily Glaphyrinae. Proceedings of the Biological Society of Washington 51, 79-86.
    • (1938) Proceedings of the Biological Society of Washington , vol.51 , pp. 79-86
    • Chapin, E.A.1
  • 26
    • 0038408097 scopus 로고    scopus 로고
    • Eulichadidae: Description of the larva of Eulichas dudgeoni Jäch, with comparative notes on larvae of other Eulichas spp., and on Stenocolus scutellaris LeConte (Coleoptera)
    • Eds M. A. Jäch and L. Ji. Zoologisch-Botanische Gesellschaft in Österreich und Wiener Coleopterologenverein: Wien, Austria
    • Costa, C., and Vanin, S. A. (1998). Eulichadidae: description of the larva of Eulichas dudgeoni Jäch, with comparative notes on larvae of other Eulichas spp., and on Stenocolus scutellaris LeConte (Coleoptera). In 'Water Beetles of China. Vol. 2'. (Eds M. A. Jäch and L. Ji.) pp. 327-336. (Zoologisch-Botanische Gesellschaft in Österreich und Wiener Coleopterologenverein: Wien, Austria.)
    • (1998) Water Beetles of China , vol.2 , pp. 327-336
    • Costa, C.1    Vanin, S.A.2
  • 27
    • 3142523073 scopus 로고
    • Contributi alla conoscenza dei Coleotteri Scarabeidi. I. Il genere Pachypus Serville (Coleoptera: Scarabaeidae: Pachypodinae)
    • Crovetti, A. (1969). Contributi alla conoscenza dei Coleotteri Scarabeidi. I. Il genere Pachypus Serville (Coleoptera: Scarabaeidae: Pachypodinae). Bolletino di Zoologia Agraria e di Bachicoltura, Serie II, 9 (1968-1969), 133-188.
    • (1969) Bolletino di Zoologia Agraria e di Bachicoltura, Serie II , vol.9 , Issue.1968-1969 , pp. 133-188
    • Crovetti, A.1
  • 29
    • 0000698003 scopus 로고
    • The phylogeny of Coleoptera
    • doi:10.1146/ANNUREV.EN.05. 010160.000551
    • Crowson, R. A. (1960). The phylogeny of Coleoptera. Annual Review of Entomology 5, 111-134. doi:10.1146/ANNUREV.EN.05. 010160.000551
    • (1960) Annual Review of Entomology , vol.5 , pp. 111-134
    • Crowson, R.A.1
  • 32
    • 0006976836 scopus 로고    scopus 로고
    • Morphology and chaetotaxy of the first-instar larva of Hydraena (Phothydraena) hernandoi Fresneda and Lagar (Hydraenidae)
    • Delgado, J. A., and Soler, A. G. (1996). Morphology and chaetotaxy of the first-instar larva of Hydraena (Phothydraena) hernandoi Fresneda and Lagar (Hydraenidae). Kolepterologische Rundschau 66, 147-154.
    • (1996) Kolepterologische Rundschau , vol.66 , pp. 147-154
    • Delgado, J.A.1    Soler, A.G.2
  • 33
    • 0006993875 scopus 로고    scopus 로고
    • Morphology and chaetotaxy of larval Hydraenidae (Coleoptera) 1: Genus Limnebius Leach, 1815 based on a description of Limnebius cordobanus d'Orchymont
    • Delgado, J. A., and Soler, A. G. (1997). Morphology and chaetotaxy of larval Hydraenidae (Coleoptera) 1: genus Limnebius Leach, 1815 based on a description of Limnebius cordobanus d'Orchymont. Aquatic Insects 19, 37-49.
    • (1997) Aquatic Insects , vol.19 , pp. 37-49
    • Delgado, J.A.1    Soler, A.G.2
  • 35
    • 0002664906 scopus 로고
    • Phylogentic significance of the structure of the external male genitalia in Scarabaeoidea
    • D'Hotman, D., and Scholtz, C. H. (1990a). Phylogentic significance of the structure of the external male genitalia in Scarabaeoidea. Entomology Memoirs 77, 1-51.
    • (1990) Entomology Memoirs , vol.77 , pp. 1-51
    • D'Hotman, D.1    Scholtz, C.H.2
  • 36
    • 0001846110 scopus 로고
    • Comparative morphology of the male genitalia of derived groups of Scarabaeoidea (Coleoptera)
    • D'Hotman, D., and Scholtz, C. H. (1990b). Comparative morphology of the male genitalia of derived groups of Scarabaeoidea (Coleoptera). Elytron 4, 3-39.
    • (1990) Elytron , vol.4 , pp. 3-39
    • D'Hotman, D.1    Scholtz, C.H.2
  • 37
    • 84981562873 scopus 로고
    • Taxonomic review of immature dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae)
    • Edmonds, W. D., and Halffter, G. (1978). Taxonomic review of immature dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae). Systematic Entomology 3, 307-331.
    • (1978) Systematic Entomology , vol.3 , pp. 307-331
    • Edmonds, W.D.1    Halffter, G.2
  • 39
    • 0003786054 scopus 로고
    • Published by the author: Port Jefferson Station, New York, USA
    • Farris, J. S. (1988). 'Hennig86, version 1.5.' (Published by the author: Port Jefferson Station, New York, USA.)
    • (1988) Hennig86, Version 1.5
    • Farris, J.S.1
  • 40
    • 0002344962 scopus 로고
    • Os Escarabídeos de Africa (Sul do Sȧara). I
    • Ferreira, M. C. (1972)[1968-69]. Os Escarabídeos de Africa (Sul do Sȧara). I. Revista de Entomologia de Moçambique 11, 5-1008.
    • (1968) Revista de Entomologia de Moçambique , vol.11 , pp. 5-1008
    • Ferreira, M.C.1
  • 41
    • 0036922211 scopus 로고    scopus 로고
    • Revision of Sceliages Westwood, a millipede-eating genus of southern African dung beetles (Coleoptera: Scarabaeidae)
    • doi:10.1071/IT01025
    • Forgie, S. A., Grebennikov, V. V., and Scholtz, C. H. (2002). Revision of Sceliages Westwood, a millipede-eating genus of southern African dung beetles (Coleoptera: Scarabaeidae). Invertebrate Systematics 16, 931-955. doi:10.1071/IT01025
    • (2002) Invertebrate Systematics , vol.16 , pp. 931-955
    • Forgie, S.A.1    Grebennikov, V.V.2    Scholtz, C.H.3
  • 42
    • 0003505458 scopus 로고
    • Published by author: Instituto Miguel Lillo, Tucumán, Argentina
    • Goloboff, P. A. (1993). 'Nona, version 1.6.' (Published by author: Instituto Miguel Lillo, Tucumán, Argentina.)
    • (1993) Nona, Version 1.6
    • Goloboff, P.A.1
  • 43
    • 3042821661 scopus 로고    scopus 로고
    • Review of larval morphology of beetles of the suborder Archostemata (Insecta: Coleoptera), including first-instar chaetotaxy
    • Grebennikov, V. V. (2004a). Review of larval morphology of beetles of the suborder Archostemata (Insecta: Coleoptera), including first-instar chaetotaxy. European Journal of Entomology 101, 273-292.
    • (2004) European Journal of Entomology , vol.101 , pp. 273-292
    • Grebennikov, V.V.1
  • 44
    • 3142586750 scopus 로고    scopus 로고
    • Grub-like larvae of Neuroptera (Insecta): A morphological review of the families Ithonidae and Polystoechotidae and a description of Oliarces clara
    • in press
    • Grebennikov, V. V. (2004b). Grub-like larvae of Neuroptera (Insecta): a morphological review of the families Ithonidae and Polystoechotidae and a description of Oliarces clara. European Journal of Entomology 101, (in press).
    • (2004) European Journal of Entomology , vol.101
    • Grebennikov, V.V.1
  • 45
    • 0038753904 scopus 로고    scopus 로고
    • Morphology of the minute larva of Ptinella tenella, with special reference to effects of miniaturisation and the systematic position of Ptiliidae (Coleoptera: Staphylinoidea)
    • doi:10.1016/S1467-8039(02)00022-1
    • Grebennikov, V. V., and Beutel, R. G. (2002). Morphology of the minute larva of Ptinella tenella, with special reference to effects of miniaturisation and the systematic position of Ptiliidae (Coleoptera: Staphylinoidea). Arthropod Structure and Development 31, 157-172. doi:10.1016/S1467-8039(02)00022-1
    • (2002) Arthropod Structure and Development , vol.31 , pp. 157-172
    • Grebennikov, V.V.1    Beutel, R.G.2
  • 46
    • 3042769636 scopus 로고    scopus 로고
    • Phylogenetic analysis of Trechitae (Coleoptera: Carabidae) based on larval morphology, with description of first-instar Phrypeus and a key to genera
    • in press
    • Grebennikov, V. V., and Maddison, D. R. (2004). Phylogenetic analysis of Trechitae (Coleoptera: Carabidae) based on larval morphology, with description of first-instar Phrypeus and a key to genera. Systematic Entomology 29, (in press).
    • (2004) Systematic Entomology , vol.29
    • Grebennikov, V.V.1    Maddison, D.R.2
  • 47
    • 0037688970 scopus 로고    scopus 로고
    • Larvae of Dascillidae (Coleoptera): Morphological study and discussion on their proposed relationships to Scarabaeoidea and Eulichadidae
    • Grebennikov, V. V., and Scholtz, C. H. (2003). Larvae of Dascillidae (Coleoptera): morphological study and discussion on their proposed relationships to Scarabaeoidea and Eulichadidae. Insect Systematics and Evolution 34, 29-39.
    • (2003) Insect Systematics and Evolution , vol.34 , pp. 29-39
    • Grebennikov, V.V.1    Scholtz, C.H.2
  • 48
    • 18744382446 scopus 로고    scopus 로고
    • Larva and pupa of Cyphopisthes descarpentriesi Paulian (Coleoptera: Scarabaeoidea: Ceratocanthidae) and their phylogenetic implications
    • doi:10.1046/ J.1440-6055.2002.00307.X
    • Grebennikov, V. V., Ballerio, A., and Scholtz, C. H. (2002). Larva and pupa of Cyphopisthes descarpentriesi Paulian (Coleoptera: Scarabaeoidea: Ceratocanthidae) and their phylogenetic implications. Australian Journal of Entomology 41, 367-374. doi:10.1046/ J.1440-6055.2002.00307.X
    • (2002) Australian Journal of Entomology , vol.41 , pp. 367-374
    • Grebennikov, V.V.1    Ballerio, A.2    Scholtz, C.H.3
  • 49
    • 3142631677 scopus 로고    scopus 로고
    • Larvae of Ceratocanthidae and Hybosoridae (Coleoptera: Scarabaeoidea): Study of morphology, phylogenetic analysis and evidence of paraphyly of Hybosoridae
    • in press
    • Grebennikov, V. V., Ballerio, A., Ocampo, F. C., and Scholtz, C. H. (2004). Larvae of Ceratocanthidae and Hybosoridae (Coleoptera: Scarabaeoidea): study of morphology, phylogenetic analysis and evidence of paraphyly of Hybosoridae. Systematic Entomology 29, (in press).
    • (2004) Systematic Entomology , vol.29
    • Grebennikov, V.V.1    Ballerio, A.2    Ocampo, F.C.3    Scholtz, C.H.4
  • 50
    • 0033813585 scopus 로고    scopus 로고
    • Systematic revision of endemic southern African genera of Bolboceratinae (Coleoptera: Scarabaeoidea: Bolboceratidae)
    • doi:10.1080/00222930050020122
    • Gussmann, S. M. V., and Scholtz, C. H. (2000). Systematic revision of endemic southern African genera of Bolboceratinae (Coleoptera: Scarabaeoidea: Bolboceratidae). Journal of Natural History 34, 1045-1123. doi:10.1080/ 00222930050020122
    • (2000) Journal of Natural History , vol.34 , pp. 1045-1123
    • Gussmann, S.M.V.1    Scholtz, C.H.2
  • 51
    • 0034938586 scopus 로고    scopus 로고
    • Systematic revision of the Afrotropical genus Bolbaffer Vulcano, Martínez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with description of eight new species
    • doi:10.1080/002229301300323910
    • Gussmann, S. M. V., and Scholtz, C. H. (2001). Systematic revision of the Afrotropical genus Bolbaffer Vulcano, Martínez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with description of eight new species. Journal of Natural History 35, 1013-1084. doi:10.1080/ 002229301300323910
    • (2001) Journal of Natural History , vol.35 , pp. 1013-1084
    • Gussmann, S.M.V.1    Scholtz, C.H.2
  • 52
    • 0001901748 scopus 로고
    • Evolution and classification of the Hydrophiloidea - A systematic review
    • Eds J. Pakaluk and S. A. Ślipiński Muzeum 1 Instytut Zoologii PAN: Warsawa, Poland
    • Hansen, M. (1995). Evolution and classification of the Hydrophiloidea - a systematic review. In 'Biology, Phylogeny, and Classification of Coleoptera. Papers Celebrating the 80th Birthday of Roy A. Crowson. Vol. 1'. (Eds J. Pakaluk and S. A. Ślipiński.) pp. 321-353. (Muzeum 1 Instytut Zoologii PAN: Warsawa, Poland.)
    • (1995) Biology, Phylogeny, and Classification of Coleoptera. Papers Celebrating the 80th Birthday of Roy A. Crowson , vol.1 , pp. 321-353
    • Hansen, M.1
  • 54
    • 0004288169 scopus 로고
    • Princeton University Press: Princeton, NJ, USA
    • Hanski, I., and Cambefort, Y. (Eds) (1991). 'Dung Beetle Ecology.' (Princeton University Press: Princeton, NJ, USA.)
    • (1991) Dung Beetle Ecology
    • Hanski, I.1    Cambefort, Y.2
  • 55
    • 0344367509 scopus 로고
    • Descriptions of the larvae and pupae of some North American Mantispinae (Neuroptera: Mantispidae) and development of a system of larval chaetotaxy for Neuroptera
    • Hoffman, K. M., and Brushwein, J. R. (1992). Descriptions of the larvae and pupae of some North American Mantispinae (Neuroptera: Mantispidae) and development of a system of larval chaetotaxy for Neuroptera. Transactions of the American Entomological Society 118, 159-196.
    • (1992) Transactions of the American Entomological Society , vol.118 , pp. 159-196
    • Hoffman, K.M.1    Brushwein, J.R.2
  • 56
    • 0039515221 scopus 로고
    • The systematic position of the genus Diphyllostoma Fall (Coleoptera: Scarabaeoidea)
    • Holloway, B. A. (1972). The systematic position of the genus Diphyllostoma Fall (Coleoptera: Scarabaeoidea). New Zealand Journal of Science 15, 31-38.
    • (1972) New Zealand Journal of Science , vol.15 , pp. 31-38
    • Holloway, B.A.1
  • 57
    • 3142553947 scopus 로고
    • Coleoptera: Scarabaeoidea
    • ABRS and CSIRO Publishing: Canberra, Australia
    • Houston, W. W. K. (Ed.) (1992). Coleoptera: Scarabaeoidea. In 'Zoological Catalogue of Australia. Vol. 9'. (ABRS and CSIRO Publishing: Canberra, Australia.)
    • (1992) Zoological Catalogue of Australia , vol.9
    • Houston, W.W.K.1
  • 58
    • 84918519489 scopus 로고    scopus 로고
    • Pleocomidae LeConte 1861
    • Eds R. H. Arnett, M. C. Thomas, P. E. Skelley, and J. H. Frank. CRC Press: Boca Raton, FL, USA
    • Hovore, F. T. (2002). Pleocomidae LeConte 1861. In 'American Beetles. Vol. 2'. (Eds R. H. Arnett, M. C. Thomas, P. E. Skelley, and J. H. Frank.) pp. 20-22. (CRC Press: Boca Raton, FL, USA.)
    • (2002) American Beetles , vol.2 , pp. 20-22
    • Hovore, F.T.1
  • 59
    • 0001246839 scopus 로고
    • Biology and taxonomy of North American beetles of the subfamily Geotrupinae with revisions of the genera Bolbocerosoma, Eucanthus, Geotrupes and Peltotrupes (Scarabaeidae)
    • Howden, H. F. (1955). Biology and taxonomy of North American beetles of the subfamily Geotrupinae with revisions of the genera Bolbocerosoma, Eucanthus, Geotrupes and Peltotrupes (Scarabaeidae). Proceedings of the United States National Museum 104, 151-319.
    • (1955) Proceedings of the United States National Museum , vol.104 , pp. 151-319
    • Howden, H.F.1
  • 60
    • 0000546533 scopus 로고
    • Larval and adult characters of Frickius Germain, its relationship to the Geotrupini, and a phylogeny of some major taxa in the Scarabaeoidea (Insecta: Coleoptera)
    • Howden, H. F. (1982). Larval and adult characters of Frickius Germain, its relationship to the Geotrupini, and a phylogeny of some major taxa in the Scarabaeoidea (Insecta: Coleoptera). Canadian Journal of Zoology 60, 2713-2724.
    • (1982) Canadian Journal of Zoology , vol.60 , pp. 2713-2724
    • Howden, H.F.1
  • 61
    • 84971060145 scopus 로고
    • A revision of the Australian beetle genera Bolboleaus Howden and Cooper, Blackbolbus Howden and Cooper, and Bolborhachium Boucomont (Scarabaeidae: Geotrupinae)
    • Howden, H. F. (1985). A revision of the Australian beetle genera Bolboleaus Howden and Cooper, Blackbolbus Howden and Cooper, and Bolborhachium Boucomont (Scarabaeidae: Geotrupinae). Australian Journal of Zoology Supplementary Series No. 111, 1-179.
    • (1985) Australian Journal of Zoology Supplementary Series No. 111 , vol.111 , pp. 1-179
    • Howden, H.F.1
  • 62
    • 84971098602 scopus 로고
    • The Geotrupinae of New Guinea (Coleoptera: Scarabaeidae)
    • Howden, H. F. (1989). The Geotrupinae of New Guinea (Coleoptera: Scarabaeidae). Invertebrate Taxonomy 3, 261-289.
    • (1989) Invertebrate Taxonomy , vol.3 , pp. 261-289
    • Howden, H.F.1
  • 63
    • 84970567237 scopus 로고
    • A revision of the Australian beetle genera Eucanthus Westwood, Bolbobaineus Howden, Cooper, Australobolbus Howden, Cooper and Gilletinus Boucomont (Scarabaeidae: Geotrupinae)
    • Howden, H. F. (1992). A revision of the Australian beetle genera Eucanthus Westwood, Bolbobaineus Howden, Cooper, Australobolbus Howden, Cooper and Gilletinus Boucomont (Scarabaeidae: Geotrupinae). Invertebrate Taxonomy 6, 605-717.
    • (1992) Invertebrate Taxonomy , vol.6 , pp. 605-717
    • Howden, H.F.1
  • 64
    • 0242541877 scopus 로고    scopus 로고
    • A new tribe of Hybosorinae with a description of a new species of Callosides Howden (Coleoptera: Scarabaeidae)
    • Howden, H. F. (2001). A new tribe of Hybosorinae with a description of a new species of Callosides Howden (Coleoptera: Scarabaeidae). The Coleopterists Bulletin 55, 199-204.
    • (2001) The Coleopterists Bulletin , vol.55 , pp. 199-204
    • Howden, H.F.1
  • 65
    • 0007958076 scopus 로고
    • The generic classification of the Bolboceratini of the Australian Region, with description of four new genera (Scarabaeidae: Geotrupinae)
    • Howden, H. F., and Cooper, J. B. (1977). The generic classification of the Bolboceratini of the Australian Region, with description of four new genera (Scarabaeidae: Geotrupinae). Australian Journal of Entomology (Suppl.) 50, 1-179.
    • (1977) Australian Journal of Entomology , vol.50 , Issue.SUPPL. , pp. 1-179
    • Howden, H.F.1    Cooper, J.B.2
  • 66
    • 0034365779 scopus 로고    scopus 로고
    • Tribes of New World Ceratocanthinae, with keys to genera and descriptions of new species (Coleoptera: Scarabaeidae)
    • Howden, H. F., and Gill, B. D. (2000). Tribes of New World Ceratocanthinae, with keys to genera and descriptions of new species (Coleoptera: Scarabaeidae). Sociobiology 35, 283-329.
    • (2000) Sociobiology , vol.35 , pp. 283-329
    • Howden, H.F.1    Gill, B.D.2
  • 67
    • 1542644182 scopus 로고
    • The new tribe Athyreini and its included genera (Coleoptera: Scarabaeidae: Geotrupini)
    • Howden, H. F., and Martínez, A. (1963). The new tribe Athyreini and its included genera (Coleoptera: Scarabaeidae: Geotrupini). The Canadian Entomologist 95, 345-352.
    • (1963) The Canadian Entomologist , vol.95 , pp. 345-352
    • Howden, H.F.1    Martínez, A.2
  • 68
    • 0038924373 scopus 로고
    • A review of the New World genus Athyreus Macleay (Scarabaeidae, Geotrupinae, Athyreini)
    • Howden, H. F., and Martínez, A. (1978). A review of the New World genus Athyreus Macleay (Scarabaeidae, Geotrupinae, Athyreini). Contributions of the American Entomological Institute 15, 1-70.
    • (1978) Contributions of the American Entomological Institute , vol.15 , pp. 1-70
    • Howden, H.F.1    Martínez, A.2
  • 69
    • 0040106865 scopus 로고
    • Adult habits, larval morphology, and phylogenetic placement of Taurocerastes patagonicus Philippi (Scarabaeidae: Geotrupinae)
    • Howden, H. F., and Peck, S. B. (1987). Adult habits, larval morphology, and phylogenetic placement of Taurocerastes patagonicus Philippi (Scarabaeidae: Geotrupinae). Canadian Journal of Zoology 65, 329-332.
    • (1987) Canadian Journal of Zoology , vol.65 , pp. 329-332
    • Howden, H.F.1    Peck, S.B.2
  • 70
    • 0010704639 scopus 로고    scopus 로고
    • Phylogenetic analysis of the subtribe Rutelina and revision of the Rutela generic groups (Coleoptera: Scarabaeidae: Rutelinae: Rutelini)
    • Jameson, M. L. (1998). Phylogenetic analysis of the subtribe Rutelina and revision of the Rutela generic groups (Coleoptera: Scarabaeidae: Rutelinae: Rutelini). Bulletin of the University of Nebraska State Museum 14, 1-184.
    • (1998) Bulletin of the University of Nebraska State Museum , vol.14 , pp. 1-184
    • Jameson, M.L.1
  • 71
    • 0034001151 scopus 로고    scopus 로고
    • Synopsis of the Mexican and Guatemalan genera Rutelisca Bates and Metapachylus Bates (Coleoptera: Scarabaeidae: Rutelinae) with comments on classification of the subtribe Rutelina
    • Jameson, M. L. (2000)[1999]. Synopsis of the Mexican and Guatemalan genera Rutelisca Bates and Metapachylus Bates (Coleoptera: Scarabaeidae: Rutelinae) with comments on classification of the subtribe Rutelina. Proceedings of the Washington Entomological Society 102, 332-349.
    • (1999) Proceedings of the Washington Entomological Society , vol.102 , pp. 332-349
    • Jameson, M.L.1
  • 72
    • 3142515900 scopus 로고    scopus 로고
    • Diphyllostomatidae Holloway 1972
    • Eds R. H. Arnett, M. C. Thomas, P. E. Skelley, and J. H. Frank CRC Press: Boca Raton, FL, USA
    • Jameson, M. L., and Ratcliffe, B. C. (2002). Diphyllostomatidae Holloway 1972. In 'American Beetles. Vol. 2'. (Eds R. H. Arnett, M. C. Thomas, P. E. Skelley, and J. H. Frank.) pp. 10-11. (CRC Press: Boca Raton, FL, USA.)
    • (2002) American Beetles , vol.2 , pp. 10-11
    • Jameson, M.L.1    Ratcliffe, B.C.2
  • 73
    • 0001818334 scopus 로고
    • A synopsis of the Neotropical genus Calomacraspis Bates with a key to larvae of the American genera of Rutelini (Coleoptera: Scarabaeidae: Rutelinae)
    • Jameson, M. L., Ratcliffe, B. C., and Morón, M. A. (1994). A synopsis of the Neotropical genus Calomacraspis Bates with a key to larvae of the American genera of Rutelini (Coleoptera: Scarabaeidae: Rutelinae). Annals of the Entomological Society of America 87, 43-58.
    • (1994) Annals of the Entomological Society of America , vol.87 , pp. 43-58
    • Jameson, M.L.1    Ratcliffe, B.C.2    Morón, M.A.3
  • 74
    • 0005707401 scopus 로고
    • Morphologie abdominale des coléoptères et systématique de l'orde
    • Jeannel, R., and Paulian, R. (1944). Morphologie abdominale des coléoptères et systématique de l'orde. Revue Française d'Entomologie 11, 65-110.
    • (1944) Revue Française D'Entomologie , vol.11 , pp. 65-110
    • Jeannel, R.1    Paulian, R.2
  • 75
    • 1042274805 scopus 로고    scopus 로고
    • Morphology and phylogeny of the larval stages of the tribe Agathidiini (Coleoptera: Leiodidae: Leiodinae)
    • Warszawa.
    • Kilian, A. (1998). Morphology and phylogeny of the larval stages of the tribe Agathidiini (Coleoptera: Leiodidae: Leiodinae). Annales Zoologici 48, 125-220[Warszawa].
    • (1998) Annales Zoologici , vol.48 , pp. 125-220
    • Kilian, A.1
  • 76
    • 0000266346 scopus 로고
    • Chaetotaxy of larval Histeridae (Coleoptera:Hydrophiloidea) based on a description of Onthophilus nodatus LeConte
    • Kovarik, P. W., and Passoa, S. (1993). Chaetotaxy of larval Histeridae (Coleoptera:Hydrophiloidea) based on a description of Onthophilus nodatus LeConte. Annals of the Entomological Society of America 86, 560-576.
    • (1993) Annals of the Entomological Society of America , vol.86 , pp. 560-576
    • Kovarik, P.W.1    Passoa, S.2
  • 77
    • 0012849943 scopus 로고    scopus 로고
    • New and otherwise noteworthy Lethrus species from central Asia and Afghanistan (Coleoptera: Geotrupidae)
    • Král, D., and Olexa, A. (1996). New and otherwise noteworthy Lethrus species from central Asia and Afghanistan (Coleoptera: Geotrupidae). Folia Heyrovskyana 4, 49-65.
    • (1996) Folia Heyrovskyana , vol.4 , pp. 49-65
    • Král, D.1    Olexa, A.2
  • 78
    • 26744435992 scopus 로고    scopus 로고
    • Revision of the genera Odontotrypes and Phelotrupes (Coleoptera: Geotrupidae)
    • Král, D., Maly, V., and Schneider, J. (2001). Revision of the genera Odontotrypes and Phelotrupes (Coleoptera: Geotrupidae). Folia Heyrovskyana Supplementum 8, 1-178.
    • (2001) Folia Heyrovskyana Supplementum , vol.8 , pp. 1-178
    • Král, D.1    Maly, V.2    Schneider, J.3
  • 79
    • 0012634729 scopus 로고
    • A new key to the suprageneric taxa in the beetle family Cetoniidae, with annotated list of the known genera
    • Krikken, J. (1984). A new key to the suprageneric taxa in the beetle family Cetoniidae, with annotated list of the known genera. Zoologische Verhandelingen 210, 1-75.
    • (1984) Zoologische Verhandelingen , vol.210 , pp. 1-75
    • Krikken, J.1
  • 80
    • 0038494767 scopus 로고
    • Revision of some hybosorine genera from the Indo-Malayan subregion: Microphaeochroops, Microphaeolodes, Mimocoelodes, Pantolasius and Phaeochridius (Coleoptera: Scarabaeidae: Hybosorinae)
    • Kuijten, P. J. (1985). Revision of some hybosorine genera from the Indo-Malayan subregion: Microphaeochroops, Microphaeolodes, Mimocoelodes, Pantolasius and Phaeochridius (Coleoptera: Scarabaeidae: Hybosorinae). Zoologische Verhandelingen 222, 1-39.
    • (1985) Zoologische Verhandelingen , vol.222 , pp. 1-39
    • Kuijten, P.J.1
  • 81
    • 3142627971 scopus 로고
    • Revision of the African and Madagascan species of Phaeochrous Castelnau, 1840 (Coleoptera: Scarabaeidae: Hybosorinae)
    • Kuijten, P. J. (1986). Revision of the African and Madagascan species of Phaeochrous Castelnau, 1840 (Coleoptera: Scarabaeidae: Hybosorinae). Zoologische Wetenschappen 249, 1-50.
    • (1986) Zoologische Wetenschappen , vol.249 , pp. 1-50
    • Kuijten, P.J.1
  • 83
    • 1842680212 scopus 로고    scopus 로고
    • Relationships among coleopteran suborders and major endoneopteran lineages: Evidence from hind wing characters
    • Kukalová-Peck, J., and Lawrence, J. F. (2004). Relationships among coleopteran suborders and major endoneopteran lineages: Evidence from hind wing characters. European Journal of Entomology 101, 95-144.
    • (2004) European Journal of Entomology , vol.101 , pp. 95-144
    • Kukalová-Peck, J.1    Lawrence, J.F.2
  • 85
    • 0000792752 scopus 로고
    • Order Coleoptera (general discussion, family key, various family treatments)
    • Ed. F. W. Stehr. Kendall/Hunt Publishing Co.: Dubuque, IA, USA
    • Lawrence, J. F. (1991). Order Coleoptera (general discussion, family key, various family treatments). In 'Immature Insects. Vol. 2'. (Ed. F. W. Stehr.) pp. 144-658. (Kendall/Hunt Publishing Co.: Dubuque, IA, USA.)
    • (1991) Immature Insects , vol.2 , pp. 144-658
    • Lawrence, J.F.1
  • 87
    • 0020374020 scopus 로고
    • Evolution and classification of beetles
    • doi:10.1146/ANNUREV.ES.13.110182.001401
    • Lawrence, J. F., and Newton, A. F. (1982). Evolution and classification of beetles. Annual Review of Ecology and Systematics 13, 261-290. doi:10.1146/ANNUREV.ES.13.110182.001401
    • (1982) Annual Review of Ecology and Systematics , vol.13 , pp. 261-290
    • Lawrence, J.F.1    Newton, A.F.2
  • 88
    • 0000240541 scopus 로고
    • Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names)
    • Eds J. Pakaluk and S. A. Ślipiński. Muzeum I Instytut Zoologii PAN: Warsawa, Poland
    • Lawrence, J. F., and Newton, A. F. (1995). Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). In 'Biology, Phylogeny, and Classification of Coleoptera. Papers Celebrating the 80th Birthday of Roy A. Crowson. Vol. 2'. (Eds J. Pakaluk and S. A. Ślipiński.) pp. 779-1006. (Muzeum I Instytut Zoologii PAN: Warsawa, Poland.)
    • (1995) Biology, Phylogeny, and Classification of Coleoptera. Papers Celebrating the 80th Birthday of Roy A. Crowson , vol.2 , pp. 779-1006
    • Lawrence, J.F.1    Newton, A.F.2
  • 90
    • 3142517649 scopus 로고    scopus 로고
    • Eine neue Amphicoma-Art aus Jordanien
    • Mitter, H. (1996). Eine neue Amphicoma-Art aus Jordanien. Koleopterologische Rundschau 66, 155-156.
    • (1996) Koleopterologische Rundschau , vol.66 , pp. 155-156
    • Mitter, H.1
  • 91
    • 0032390022 scopus 로고    scopus 로고
    • Analyse phylogénétique et paraphylie des Coprini et Dichotomiini (Coleoptera: Scarabaeidae). Scénario biogéographique
    • Montreuil, O. (1998). Analyse phylogénétique et paraphylie des Coprini et Dichotomiini (Coleoptera: Scarabaeidae). Scénario biogéographique. Annales de la Societe Entomologique de France 34, 135-148.
    • (1998) Annales de la Societe Entomologique de France , vol.34 , pp. 135-148
    • Montreuil, O.1
  • 93
    • 0000305562 scopus 로고    scopus 로고
    • New data on the systematics of the scarab beetles of the subfamily Hybosorinae (Coleoptera: Scarabaeidae): The establishment of the new tribe comprising four monotypic South American genera, and description of some new taxa from the Lower Cretaceous of Transbaikalia
    • Nikolajev, G. V. (1999). New data on the systematics of the scarab beetles of the subfamily Hybosorinae (Coleoptera: Scarabaeidae): the establishment of the new tribe comprising four monotypic South American genera, and description of some new taxa from the Lower Cretaceous of Transbaikalia. TETHYS Entomological Research 1, 173-182.
    • (1999) TETHYS Entomological Research , vol.1 , pp. 173-182
    • Nikolajev, G.V.1
  • 94
    • 0003558470 scopus 로고    scopus 로고
    • Published by the author: Ithaca, NY, USA
    • Nixon, K. C. (2002). 'WinClada version 1.00.08.' (Published by the author: Ithaca, NY, USA.)
    • (2002) WinClada Version 1.00.08
    • Nixon, K.C.1
  • 95
    • 0036304989 scopus 로고    scopus 로고
    • Hybosorids of the United States and expanding distribution of the introduced species Hybosorus illigeri (Coleoptera: Scarabaeoidea: Hybosoridae)
    • Ocampo, F. C. (2002). Hybosorids of the United States and expanding distribution of the introduced species Hybosorus illigeri (Coleoptera: Scarabaeoidea: Hybosoridae). Annals of the Entomological Society of America 95, 316-322.
    • (2002) Annals of the Entomological Society of America , vol.95 , pp. 316-322
    • Ocampo, F.C.1
  • 96
    • 3142627970 scopus 로고
    • Un nouveau Coleoptere Scarabeide de Madagascar type d'une nouvelle sous-famille
    • Paulian, R. (1958). Un nouveau Coleoptere Scarabeide de Madagascar type d'une nouvelle sous-famille. Le Naturaliste Malgache 10, 39-46.
    • (1958) Le Naturaliste Malgache , vol.10 , pp. 39-46
    • Paulian, R.1
  • 97
    • 3142593833 scopus 로고
    • Les Ochodaeidae (Coleoptera: Scarabaeoidea) de Madagascar
    • Paulian, R. (1976). Les Ochodaeidae (Coleoptera: Scarabaeoidea) de Madagascar. Nouvelle Revue d'Entomologie 6, 139-152.
    • (1976) Nouvelle Revue D'Entomologie , vol.6 , pp. 139-152
    • Paulian, R.1
  • 98
    • 0041137201 scopus 로고
    • Révision des Ceratocanthidae (Coleoptera: Scarabaeidae). I. Les formes africaines
    • Paulian, R. (1977). Révision des Ceratocanthidae (Coleoptera: Scarabaeidae). I. Les formes africaines. Revue de Zoologie Africaine 91, 253-316.
    • (1977) Revue de Zoologie Africaine , vol.91 , pp. 253-316
    • Paulian, R.1
  • 99
    • 33645825294 scopus 로고
    • Belohinidae. Insectes Coléoptères
    • Paulian, R. (1979a). Belohinidae. Insectes Coléoptères. Faune de Madagascar 50, 11-49.
    • (1979) Faune de Madagascar , vol.50 , pp. 11-49
    • Paulian, R.1
  • 100
    • 3142626192 scopus 로고
    • Ceratocanthidae. Insectes Coléoptères
    • Paulian, R. (1979b). Ceratocanthidae. Insectes Coléoptères. Faune de Madagascar 50, 51-56.
    • (1979) Faune de Madagascar , vol.50 , pp. 51-56
    • Paulian, R.1
  • 103
    • 1842487573 scopus 로고    scopus 로고
    • A phylogenetic analysis of dung beetles (Scarabaeinae: Scarabaeidae): Unrolling an evolutionary history
    • doi:10.1071/IS03030
    • Philips, T. K., Pretorius, E., and Scholtz, C. H. (2004). A phylogenetic analysis of dung beetles (Scarabaeinae: Scarabaeidae): unrolling an evolutionary history. Invertebrate Systematics 18, 53-88. doi:10.1071/IS03030
    • (2004) Invertebrate Systematics , vol.18 , pp. 53-88
    • Philips, T.K.1    Pretorius, E.2    Scholtz, C.H.3
  • 104
    • 0036399402 scopus 로고    scopus 로고
    • A phylogenetic analysis of the Eucraniini (Coleoptera: Scarabaeidae: Scarabaeinae)
    • Philips, T. K., Scholtz, C. H., and Ocampo, F. C. (2002). A phylogenetic analysis of the Eucraniini (Coleoptera: Scarabaeidae: Scarabaeinae). Insect Systematics & Evolution 33, 241-252.
    • (2002) Insect Systematics & Evolution , vol.33 , pp. 241-252
    • Philips, T.K.1    Scholtz, C.H.2    Ocampo, F.C.3
  • 105
    • 85056069141 scopus 로고    scopus 로고
    • Scarabaeidae Latreille 1802
    • Eds R. H. Arnett, M. C. Thomas, P. E. Skelley, and J. H. Frank. CRC Press: Boca Raton, FL, USA
    • Ratcliffe, B. C., Jameson, M. L., and Smith, A. B. T. (2002). Scarabaeidae Latreille 1802. In 'American Beetles. Vol. 2'. (Eds R. H. Arnett, M. C. Thomas, P. E. Skelley, and J. H. Frank.) pp. 39-81. (CRC Press: Boca Raton, FL, USA.)
    • (2002) American Beetles , vol.2 , pp. 39-81
    • Ratcliffe, B.C.1    Jameson, M.L.2    Smith, A.B.T.3
  • 106
    • 0002859196 scopus 로고
    • Coleoptera Passalidae: Morfología y división en grandes grupos géneros americanos
    • Reyes-Castillo, P. (1970). Coleoptera Passalidae: morfología y división en grandes grupos géneros americanos. Folia Entomologica Mexicana 20-22, 1-236.
    • (1970) Folia Entomologica Mexicana , vol.20-22 , pp. 1-236
    • Reyes-Castillo, P.1
  • 108
    • 0010600981 scopus 로고
    • White grubs and their allies. A study of North American Scarabaeidae larvae
    • Oregon State University Press: Corvallis, OR, USA
    • Ritcher, P. O. (1966). 'White grubs and their allies. A study of North American Scarabaeidae larvae.' Oregon State Monographs, Studies in Entomology. (Oregon State University Press: Corvallis, OR, USA.)
    • (1966) Oregon State Monographs, Studies in Entomology
    • Ritcher, P.O.1
  • 109
    • 0038922769 scopus 로고
    • Morphology of the posterior procoxal bridges in Scarabaeoidea (Coleoptera)
    • Ritcher, P. O. (1969). Morphology of the posterior procoxal bridges in Scarabaeoidea (Coleoptera). The Coleopterists Bulletin 23, 89-92.
    • (1969) The Coleopterists Bulletin , vol.23 , pp. 89-92
    • Ritcher, P.O.1
  • 111
    • 0003264295 scopus 로고
    • A review of the genus Glaresis Erichson (Coleoptera: Trogidae) of sub-Saharan Africa
    • Scholtz, C. H. (1983). A review of the genus Glaresis Erichson (Coleoptera: Trogidae) of sub-Saharan Africa. Journal of the Entomological Society of Southern Africa 46, 209-225.
    • (1983) Journal of the Entomological Society of Southern Africa , vol.46 , pp. 209-225
    • Scholtz, C.H.1
  • 112
    • 84981608323 scopus 로고
    • Phylogeny and Systematics of Trogidae (Coleoptera: Scarabaeoidea)
    • Scholtz, C. H. (1986). Phylogeny and Systematics of Trogidae (Coleoptera: Scarabaeoidea). Systematic Entomology 11, 355-363.
    • (1986) Systematic Entomology , vol.11 , pp. 355-363
    • Scholtz, C.H.1
  • 113
    • 0000011307 scopus 로고
    • Phylogenetic trends in the Scarabaeoidea (Coleoptera)
    • Scholtz, C. H. (1990). Phylogenetic trends in the Scarabaeoidea (Coleoptera). Journal of Natural History 24, 1027-1066.
    • (1990) Journal of Natural History , vol.24 , pp. 1027-1066
    • Scholtz, C.H.1
  • 114
    • 0029808286 scopus 로고    scopus 로고
    • Polyphyly in the Geotrupidae (Coleoptera: Scarabaeoidea): A case for a new family
    • Scholtz, C. H., and Browne, D. J. (1996). Polyphyly in the Geotrupidae (Coleoptera: Scarabaeoidea): a case for a new family. Journal of Natural History 30, 597-614.
    • (1996) Journal of Natural History , vol.30 , pp. 597-614
    • Scholtz, C.H.1    Browne, D.J.2
  • 116
    • 3142521297 scopus 로고    scopus 로고
    • Scarabaeoidea
    • in press (Eds R. G. Beutel and R. A. B. Leschen.) (Gruyter: Berlin, Germany.)
    • Scholtz, C. H., and Grebennikov, V. V. (in press). Scarabaeoidea. In 'Handbook of Zoology. Vol. 3'. (Eds R. G. Beutel and R. A. B. Leschen.) (Gruyter: Berlin, Germany.)
    • Handbook of Zoology , vol.3
    • Scholtz, C.H.1    Grebennikov, V.V.2
  • 117
    • 84981584599 scopus 로고
    • Description of a Polynoncus Burmeister larva, with implications for phylogeny of the Trogidae (Coleoptera: Scarabaeoidea)
    • Scholtz, C. H., and Peck, S. B. (1990). Description of a Polynoncus Burmeister larva, with implications for phylogeny of the Trogidae (Coleoptera: Scarabaeoidea). Systematic Entomology 15, 383-389.
    • (1990) Systematic Entomology , vol.15 , pp. 383-389
    • Scholtz, C.H.1    Peck, S.B.2
  • 118
    • 0041982442 scopus 로고
    • Systematic position of the South American genus Cryptogenius Westwood (Coleoptera: Scarabaeoidea: Trogidae and Hybosoridae)
    • Scholtz, C. H., D'Hotman, D., and Nel, A. (1987a). Systematic position of the South American genus Cryptogenius Westwood (Coleoptera: Scarabaeoidea: Trogidae and Hybosoridae). The Coleopterists Bulletin 41, 193-199.
    • (1987) The Coleopterists Bulletin , vol.41 , pp. 193-199
    • Scholtz, C.H.1    D'Hotman, D.2    Nel, A.3
  • 119
    • 84981689189 scopus 로고
    • Glaresidae, a new family of Scarabaeoidea (Coleoptera) to accommodate the genus Glaresis Erichson
    • Scholtz, C. H., D'Hotman, D., and Nel, A. (1987b). Glaresidae, a new family of Scarabaeoidea (Coleoptera) to accommodate the genus Glaresis Erichson. Systematic Entomology 12, 343-354.
    • (1987) Systematic Entomology , vol.12 , pp. 343-354
    • Scholtz, C.H.1    D'Hotman, D.2    Nel, A.3
  • 121
    • 84981569430 scopus 로고
    • Glaresidae, archaeopteryx of the Scarabaeoidea (Coleoptera)
    • Scholtz, C. H., Browne, J., and Kukalová-Peck, J. (1994). Glaresidae, archaeopteryx of the Scarabaeoidea (Coleoptera). Systematic Entomology 19, 259-277.
    • (1994) Systematic Entomology , vol.19 , pp. 259-277
    • Scholtz, C.H.1    Browne, J.2    Kukalová-Peck, J.3
  • 122
    • 3142517650 scopus 로고
    • Passalidae: State of larval taxonomy with description of New World species
    • Schuster, J. C. (1992). Passalidae: state of larval taxonomy with description of New World species. The Florida Entomologist 75, 357-369.
    • (1992) The Florida Entomologist , vol.75 , pp. 357-369
    • Schuster, J.C.1
  • 123
    • 3142523071 scopus 로고
    • New larval descriptions from Taiwan, Philippine Islands, Brunei and Ivory Coast
    • Schuster, J. C., and Reyes-Castillo, P. (1990). New larval descriptions from Taiwan, Philippine Islands, Brunei and Ivory Coast. Florida Entomologists 73, 267-273.
    • (1990) Florida Entomologists , vol.73 , pp. 267-273
    • Schuster, J.C.1    Reyes-Castillo, P.2
  • 124
    • 0000808867 scopus 로고
    • Social behavior in Passalid beetles (Coleoptera: Passalidae): Cooperative brood care
    • Schuster, J. C., and Schuster, L. B. (1985). Social behavior in Passalid beetles (Coleoptera: Passalidae): cooperative brood care. The Florida Entomologist 68, 266-272.
    • (1985) The Florida Entomologist , vol.68 , pp. 266-272
    • Schuster, J.C.1    Schuster, L.B.2
  • 125
    • 0036745592 scopus 로고    scopus 로고
    • Revision of the southern South American endemic genus Aulacopalpus Guérin-Méneville with phylogenetic and biogeographic analysis of the subtribe Brachysternina (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini)
    • Smith, A. B. T. (2002). Revision of the southern South American endemic genus Aulacopalpus Guérin-Méneville with phylogenetic and biogeographic analysis of the subtribe Brachysternina (Coleoptera: Scarabaeidae: Rutelinae: Anoplognathini). The Coleopterists Bulletin 56, 379-437.
    • (2002) The Coleopterists Bulletin , vol.56 , pp. 379-437
    • Smith, A.B.T.1
  • 126
    • 3142557555 scopus 로고    scopus 로고
    • Revision of the relict South African genus Arrowinus, description of its larva, and phylogenetic assessment of Arrowinini trib. n. (Coleoptera: Staphylinidae: Staphylininae)
    • in press
    • Solodovnikov, A. Yu., and Newton, A. F. (2004). Revision of the relict South African genus Arrowinus, description of its larva, and phylogenetic assessment of Arrowinini trib. n. (Coleoptera: Staphylinidae: Staphylininae). Systematic Entomology 29, (in press).
    • (2004) Systematic Entomology , vol.29
    • Yu, S.A.1    Newton, A.F.2
  • 127
    • 0012714502 scopus 로고    scopus 로고
    • Lomanoxia Martínez, 1951, and a new tribe Lomanoxiini with notes on comparative morphology (Coleoptera: Scarabaeidae: Aphodiinae)
    • Stebnicka, Z. (1999). Lomanoxia Martínez, 1951, and a new tribe Lomanoxiini with notes on comparative morphology (Coleoptera: Scarabaeidae: Aphodiinae). Acta Zoologica Cracoviensia 42, 279-286.
    • (1999) Acta Zoologica Cracoviensia , vol.42 , pp. 279-286
    • Stebnicka, Z.1
  • 128
    • 0009361231 scopus 로고    scopus 로고
    • Australian genera and species in the tribes Odontolochini, Psammodiini, Rhyparini, Stereomerini and part of the Eupariini (Coleoptera: Scarabaeoidea: Aphodiina)
    • Stebnicka, Z., and Howden, H. F. (1996). Australian genera and species in the tribes Odontolochini, Psammodiini, Rhyparini, Stereomerini and part of the Eupariini (Coleoptera: Scarabaeoidea: Aphodiina). Invertebrate Taxonomy 10, 97-170.
    • (1996) Invertebrate Taxonomy , vol.10 , pp. 97-170
    • Stebnicka, Z.1    Howden, H.F.2
  • 129
    • 0041799042 scopus 로고
    • Termitophilous scarabs of the tribe Corythoderini: A taxonomic review (Coleoptera: Aphodiinae)
    • Tangelder, I. R. M., and Krikken, J. (1982). Termitophilous scarabs of the tribe Corythoderini: a taxonomic review (Coleoptera: Aphodiinae). Zoologische Verhandelingen 194, 1-194.
    • (1982) Zoologische Verhandelingen , vol.194 , pp. 1-194
    • Tangelder, I.R.M.1    Krikken, J.2
  • 130
    • 0033675206 scopus 로고    scopus 로고
    • Glypholoma larvae at last: Phylogenetic implications for basal Staphylinidae? (Coleoptera: Staphylinidae: Glypholomatinae)
    • doi:10.1071/IT00019
    • Thayer, M. K. (2000). Glypholoma larvae at last: phylogenetic implications for basal Staphylinidae? (Coleoptera: Staphylinidae: Glypholomatinae). Invertebrate Taxonomy 14, 741-754. doi:10.1071/IT00019
    • (2000) Invertebrate Taxonomy , vol.14 , pp. 741-754
    • Thayer, M.K.1
  • 131
    • 0032364025 scopus 로고    scopus 로고
    • Description de la larve de Bolbelasmus bocchus (Erichson) et position systématique du genre (Coleoptera: Geotrupidae: Bolboceratinae)
    • Verdú, J. R., Galante, E., and Lumaret, J.-P. (1998). Description de la larve de Bolbelasmus bocchus (Erichson) et position systématique du genre (Coleoptera: Geotrupidae: Bolboceratinae). Annnales de la Société Entomologique de France (N.S.) 34, 245-251.
    • (1998) Annnales de la Société Entomologique de France (N.S.) , vol.34 , pp. 245-251
    • Verdú, J.R.1    Galante, E.2    Lumaret, J.-P.3
  • 133
    • 0038069683 scopus 로고
    • Beiträge zur Kenntnis der Coleopteren Larven besonderer Berücksichtigung der Clavicornia
    • Verhoeff, K. W. (1923a). Beiträge zur Kenntnis der Coleopteren Larven besonderer Berücksichtigung der Clavicornia. Archiv für Naturgeschichte 89, 1-109.
    • (1923) Archiv für Naturgeschichte , vol.89 , pp. 1-109
    • Verhoeff, K.W.1
  • 134
    • 0037732095 scopus 로고
    • Über Larventypen der Coleopteren und die Auflösung der alten Malacodermata
    • Verhoeff, K. W. (1923b). Über Larventypen der Coleopteren und die Auflösung der alten Malacodermata. Zeitschrift für Wissenschaftliche Insektenbiologie 17, 115-125.
    • (1923) Zeitschrift für Wissenschaftliche Insektenbiologie , vol.17 , pp. 115-125
    • Verhoeff, K.W.1
  • 135
    • 0006117975 scopus 로고
    • Morphology and ontogeny of postembryonic larval Agathidium and Anisotoma (Coleoptera: Leiodidae)
    • Wheeler, Q. D. (1990). Morphology and ontogeny of postembryonic larval Agathidium and Anisotoma (Coleoptera: Leiodidae). American Museum Novitates 2986, 1-46.
    • (1990) American Museum Novitates , vol.2986 , pp. 1-46
    • Wheeler, Q.D.1
  • 136
    • 0024943843 scopus 로고
    • Euchirinae (Coleoptera: Scarabaeidae) of the world: Distribution and taxonomy
    • Young, R. M. (1989). Euchirinae (Coleoptera: Scarabaeidae) of the world: distribution and taxonomy. The Coleopterists Bulletin 43, 205-236.
    • (1989) The Coleopterists Bulletin , vol.43 , pp. 205-236
    • Young, R.M.1
  • 137
    • 0000631199 scopus 로고
    • Sistematica generica dei Geotrupinae (Coleoptera, Scarabaeoidea: Geotrupidae), filogenesi della sottofamiglia e considerazioni biogeografiche
    • Turin.
    • Zunino, M. (1984). Sistematica generica dei Geotrupinae (Coleoptera, Scarabaeoidea: Geotrupidae), filogenesi della sottofamiglia e considerazioni biogeografiche. Museo Regionale di Scienze Naturali Bollettino 2, 9-162[Turin].
    • (1984) Museo Regionale di Scienze Naturali Bollettino , vol.2 , pp. 9-162
    • Zunino, M.1


* 이 정보는 Elsevier사의 SCOPUS DB에서 KISTI가 분석하여 추출한 것입니다.