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note
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This work was funded in part by a NERC LOIS Special Topic allocation (GST/02/ 736) and as part of the JONUS (Joint Nutrient Study) programme funded by the UK Ministry of Agriculture, Fisheries and Food, Department of the Environment, Transport and the Regions and the Environment Agency. The views expressed are those of the authors and do not reflect policies of the funding departments. We thank the anonymous referees of this paper for their useful and constructive comments which significantly improved the final paper. We are grateful for technical support at UEA, especially from Penny King and Tristam Hardman, who assisted with CHNS elemental analysis. Also many thanks to Philip Judge who helped produce the colour figures. John Ridgway publishes by permission of the Director, British Geological Survey. This paper is LOIS publication number 718.
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46
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85037968696
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First submitted 25 May 1999. Accepted for publication after revision 17 November 1999.
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