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Volumn 38, Issue 2, 2000, Pages 73-79

Sex chromatin and sex chromosome systems in nonditrysian Lepidoptera (Insecta)

Author keywords

Amphiesmenoptera; Dacnonypha; Exoporia; Incurvariina; Karyotype; Lepidoptera; Nepti culina; Phylogeny; Sex chromatin; Sex chromosome; Tischeriidae; Tischeriina; Trichoptera; Zeugloptera

Indexed keywords

TISCHERIA EKEBLADELLA;

EID: 0034210517     PISSN: 09475745     EISSN: None     Source Type: Journal    
DOI: 10.1046/j.1439-0469.2000.382130.x     Document Type: Article
Times cited : (61)

References (49)
  • 1
    • 84998293228 scopus 로고
    • Karyotypes of some Lepidoptera. Chromosomes and changes in their holokinetic organisation as revealed by new cytological techniques
    • Bigger, T. R. L., 1975: Karyotypes of some Lepidoptera. Chromosomes and changes in their holokinetic organisation as revealed by new cytological techniques. Cytologia 40, 713-726.
    • (1975) Cytologia , vol.40 , pp. 713-726
    • Bigger, T.R.L.1
  • 2
    • 0017743179 scopus 로고
    • Linkage and cytogenetic studies in the swallowtail butterflies Papilio polyxenes Fab. & Papilio machaon L. & their hybrids
    • Clarke, C.; Mittwoch, U.; Traut, W., 1977: Linkage and cytogenetic studies in the swallowtail butterflies Papilio polyxenes Fab. & Papilio machaon L. & their hybrids. Proc. R. Soc. London: Series B 198, 385-399.
    • (1977) Proc. R. Soc. London: Series B , vol.198 , pp. 385-399
    • Clarke, C.1    Mittwoch, U.2    Traut, W.3
  • 3
    • 0000097167 scopus 로고
    • Evolution and classification of the Lepidoptera
    • Common, I. F. B., 1975: Evolution and classification of the Lepidoptera. Annu. Rev. Ent. 20, 183-203.
    • (1975) Annu. Rev. Ent. , vol.20 , pp. 183-203
    • Common, I.F.B.1
  • 4
    • 0002687625 scopus 로고
    • A new family of monotrysian moths from austral South America (Lepidoptera: Palaephatidae), with a phylogenetic review of the Monotrysia
    • Davis, D. R., 1986: A new family of monotrysian moths from austral South America (Lepidoptera: Palaephatidae), with a phylogenetic review of the Monotrysia. Smithson Contr. Zool. 434, I-IV, 1-202.
    • (1986) Smithson Contr. Zool. , vol.434 , Issue.1-4 , pp. 1-202
    • Davis, D.R.1
  • 5
    • 0017082659 scopus 로고
    • Sex chromatin and chromosome numbers in Lepidoptera
    • Ennis, T. E., 1976: Sex chromatin and chromosome numbers in Lepidoptera. Can. J. Genet. Cytol. 18, 119-130.
    • (1976) Can. J. Genet. Cytol. , vol.18 , pp. 119-130
    • Ennis, T.E.1
  • 6
    • 0008602396 scopus 로고
    • Study of mitotic chromosome of ghost moth
    • in Chinese, with English summary
    • Fayao, L.; Farong, S.; Darong, Y.; Yongchen, L.; Yaoxiong, Y., 1988: Study of mitotic chromosome of ghost moth. Zoological Res. 9, 15-20 (in Chinese, with English summary).
    • (1988) Zoological Res. , vol.9 , pp. 15-20
    • Fayao, L.1    Farong, S.2    Darong, Y.3    Yongchen, L.4    Yaoxiong, Y.5
  • 7
    • 0002333268 scopus 로고    scopus 로고
    • Analysis of morphological and behavioural evidence for the phylogeny and higher classification of Trichoptera (Insecta)
    • Frania, H. E.; Wiggins, G. B., 1997: Analysis of morphological and behavioural evidence for the phylogeny and higher classification of Trichoptera (Insecta). Royal Ontario Museum. Life Sci. Contributions 160, 1-67.
    • (1997) Royal Ontario Museum. Life Sci. Contributions , vol.160 , pp. 1-67
    • Frania, H.E.1    Wiggins, G.B.2
  • 8
    • 0008531013 scopus 로고
    • Karyotype and meiotic mechanism in Muga silkmoths, Antheraea compta Roth. and A. assamensis (Helf.) (Lepidoptera: Saturniidae)
    • Gupta, M. L.; Narang, R. C., 1981: Karyotype and meiotic mechanism in Muga silkmoths, Antheraea compta Roth. and A. assamensis (Helf.) (Lepidoptera: Saturniidae). Genetica 57, 21-27.
    • (1981) Genetica , vol.57 , pp. 21-27
    • Gupta, M.L.1    Narang, R.C.2
  • 9
    • 0017726907 scopus 로고
    • Cytogenetical studies on the chromosomes of silk gland cells of the silkworm with special reference to the structure and behaviour of the sex chromosomes
    • Ito, S., 1977: Cytogenetical studies on the chromosomes of silk gland cells of the silkworm with special reference to the structure and behaviour of the sex chromosomes. Japan. J. Genetics 52, 327-340.
    • (1977) Japan. J. Genetics , vol.52 , pp. 327-340
    • Ito, S.1
  • 10
    • 0008572579 scopus 로고
    • Comparative analysis of the wing articulation in archaic Lepidoptera
    • in Russian, with English summary
    • Ivanov, V. D., 1994: Comparative analysis of the wing articulation in archaic Lepidoptera. Entomologicheskoe Obozrenie 73, 569-590 (in Russian, with English summary).
    • (1994) Entomologicheskoe Obozrenie , vol.73 , pp. 569-590
    • Ivanov, V.D.1
  • 11
    • 85017073916 scopus 로고
    • The chromosome in the primitive or microlepidopterous moth-groups. I
    • Kawazoe, A., 1987a: The chromosome in the primitive or microlepidopterous moth-groups. I. Proc. Japan Acad. Series B 63, 25-28.
    • (1987) Proc. Japan Acad. Series B , vol.63 , pp. 25-28
    • Kawazoe, A.1
  • 12
    • 85017073916 scopus 로고
    • The chromosome in the primitive or microlepidopterous moth-groups. II
    • Kawazoe, A.; 1987b: The chromosome in the primitive or microlepidopterous moth-groups. II. Proc. Japan Acad. Series B 63, 87-90.
    • (1987) Proc. Japan Acad. Series B , vol.63 , pp. 87-90
    • Kawazoe, A.1
  • 13
    • 85017073916 scopus 로고
    • The chromosome in the primitive or microlepidopterous moth-groups. I
    • Kawazoe, A.; 1987c: The chromosome in the primitive or microlepidopterous moth-groups. I. Proc. Japan Acad. Series B 63, 193-196.
    • (1987) Proc. Japan Acad. Series B , vol.63 , pp. 193-196
    • Kawazoe, A.1
  • 14
    • 85017073916 scopus 로고
    • The chromosome in the primitive or microlepidopterous moth-groups. IV
    • Kawazoe, A.; 1987d: The chromosome in the primitive or microlepidopterous moth-groups. IV. Proc. Japan Acad. Series B 63, 257-260.
    • (1987) Proc. Japan Acad. Series B , vol.63 , pp. 257-260
    • Kawazoe, A.1
  • 15
    • 0008533283 scopus 로고
    • Ecology, case structure, larval morphology and chromosomes of the caddis-fly, Allogamus auricollis (Pictet, 1834), with a discussion on the variation of recombination indices in the Stenophylacini (Trichoptera, Integripalpia: Limnephilidae)
    • Kiauta, B.; Kiauta, M. A. J. E., 1979: Ecology, case structure, larval morphology and chromosomes of the caddis-fly, Allogamus auricollis (Pictet, 1834), with a discussion on the variation of recombination indices in the Stenophylacini (Trichoptera, Integripalpia: Limnephilidae). Genetica 50, 119-126.
    • (1979) Genetica , vol.50 , pp. 119-126
    • Kiauta, B.1    Kiauta, M.A.J.E.2
  • 16
    • 0014453505 scopus 로고
    • The chromosomes of the caddis-fly, gliphotaelius pellucidus (Retzius, 1783) (Trichoptera, Limnephilidae: Limnephilinae)
    • Kiauta, B.; Lankhorst, L., 1969: The chromosomes of the caddis-fly, Gliphotaelius pellucidus (Retzius, 1783) (Trichoptera, Limnephilidae: Limnephilinae). Genetica 40, 1-6.
    • (1969) Genetica , vol.40 , pp. 1-6
    • Kiauta, B.1    Lankhorst, L.2
  • 17
    • 0008567336 scopus 로고
    • Digametie beim Weibchen der Trichoptere Limnophilus decipiens Kol
    • Klingstedt, H., 1931: Digametie beim Weibchen der Trichoptere Limnophilus decipiens Kol. Acta Zool. Fenn. 10, 1-69.
    • (1931) Acta Zool. Fenn. , vol.10 , pp. 1-69
    • Klingstedt, H.1
  • 18
    • 0008567609 scopus 로고
    • Imaginal morphology and the phylogeny of the moth families Opostegidae, Nepticulidae and Tischeriidae (Lepidoptera, Nepticulomorpha)
    • in Russian, with English summary
    • Kozlov, M. V., 1987: Imaginal morphology and the phylogeny of the moth families Opostegidae, Nepticulidae and Tischeriidae (Lepidoptera, Nepticulomorpha). Zool. Zhurn. 66, 847-859 (in Russian, with English summary).
    • (1987) Zool. Zhurn. , vol.66 , pp. 847-859
    • Kozlov, M.V.1
  • 19
    • 0001166072 scopus 로고
    • Studies on the morphology and systematics of primitive Lepidoptera (Insecta)
    • Kristensen, N. P., 1984: Studies on the morphology and systematics of primitive Lepidoptera (Insecta). Steenstrupia 10, 141-191.
    • (1984) Steenstrupia , vol.10 , pp. 141-191
    • Kristensen, N.P.1
  • 20
    • 0000209567 scopus 로고    scopus 로고
    • Early evolution of the Lepidoptera+Trichoptera lineage: Phylogeny and the ecological scenario
    • Grandcolas, P. (ed.), The Origin of Biodiversity in Insects: Phylogenetic Tests of Evolutionary Scenarios. Paris: Éditions du Muséum
    • Kristensen, N. P., 1997: Early evolution of the Lepidoptera+Trichoptera lineage: phylogeny and the ecological scenario. In: Grandcolas, P. (ed.), The Origin of Biodiversity in Insects: Phylogenetic Tests of Evolutionary Scenarios. Mém. Mus. Natn. Hist. Nat. 173. Paris: Éditions du Muséum. pp. 253-271.
    • (1997) Mém. Mus. Natn. Hist. Nat. , vol.173 , pp. 253-271
    • Kristensen, N.P.1
  • 21
    • 0008533423 scopus 로고    scopus 로고
    • Phylogeny and palaeontology
    • Kristensen, N. P. (ed.), Lepidoptera: Moths and Butterflies 1. Berlin, New York: Walter de Gruyter
    • Kristensen, N. P.; Skalski, A. W., 1998: Phylogeny and palaeontology. In: Kristensen, N. P. (ed.), Lepidoptera: Moths and Butterflies 1. Handbuch der Zoologie/Handbook of Zoology IV. Berlin, New York: Walter de Gruyter. pp. 35, 77-25.
    • (1998) Handbuch der Zoologie/Handbook of Zoology IV , pp. 35
    • Kristensen, N.P.1    Skalski, A.W.2
  • 22
    • 0001355637 scopus 로고    scopus 로고
    • Karyotypes, sex chromosome systems, and male meiosis in Finnish psyllids (Homoptera: Psylloidea)
    • Kuznetsova, V. G.; Nokkala, S.; Maryanska-Nadachowska, A., 1997: Karyotypes, sex chromosome systems, and male meiosis in Finnish psyllids (Homoptera: Psylloidea). Folia biologica (Krakow) 45, 143-152.
    • (1997) Folia Biologica (Krakow) , vol.45 , pp. 143-152
    • Kuznetsova, V.G.1    Nokkala, S.2    Maryanska-Nadachowska, A.3
  • 23
    • 0000308927 scopus 로고
    • The system and evolution of infraorders in the lepidoptera (Micropterigomorpha-Papilionomorpha) treated on the base of functional morphology of the genitalia
    • in Russian, with English summary
    • Kuznetzov, V. I.; Stekolnikov, A. A., 1978: The system and evolution of infraorders in the Lepidoptera (Micropterigomorpha-Papilionomorpha) treated on the base of functional morphology of the genitalia. Entomol. Obozr. 57, 870-890 (in Russian, with English summary).
    • (1978) Entomol. Obozr. , vol.57 , pp. 870-890
    • Kuznetzov, V.I.1    Stekolnikov, A.A.2
  • 24
    • 0008569477 scopus 로고
    • Karyotype of Tischeria. Ekebladella (Lepidoptera, Nepticulomorpha, Tischeriidae)
    • Tobias, V.I.; Lvovsky, A.L. (eds), Leningrad: Nauka. in Russian
    • Lukhtanov, V. A.; Puplesiene, J., 1990: Karyotype of Tischeria. Ekebladella (Lepidoptera, Nepticulomorpha, Tischeriidae). In: Tobias, V.I.; Lvovsky, A.L. (eds), Advantages of Entomology in USSR: Insects - Hymenoptera and Lepidoptera. Leningrad: Nauka. pp. 181-182 (in Russian).
    • (1990) Advantages of Entomology in USSR: Insects - Hymenoptera and Lepidoptera , pp. 181-182
    • Lukhtanov, V.A.1    Puplesiene, J.2
  • 25
    • 0008603072 scopus 로고    scopus 로고
    • Karyological peculiarities and main features of karyotype evolution in lepidopterans of the nepticuloid, tischerioid, gelechioid and tineoid complexes (Lepidoptera: Nepticuloidea, Tischerioidea, Gelechioidea s.l., Psychoidea-Gracillarioidea (Lepidoptera)
    • in Russian, with English summary
    • Lukhtanov, V. A.; Puplesiene, J., 1996: Karyological peculiarities and main features of karyotype evolution in lepidopterans of the nepticuloid, tischerioid, gelechioid and tineoid complexes (Lepidoptera: Nepticuloidea, Tischerioidea, Gelechioidea s.l., Psychoidea-Gracillarioidea (Lepidoptera). - Entomologicheskoe Obozrenie 75, 310-323 (in Russian, with English summary).
    • (1996) Entomologicheskoe Obozrenie , vol.75 , pp. 310-323
    • Lukhtanov, V.A.1    Puplesiene, J.2
  • 26
    • 85017039158 scopus 로고
    • Notes on the W-chromosome of the butterfly, Graphium sarpedon (Papilionidae: Lepidoptera)
    • Maeki, K., 1981: Notes on the W-Chromosome of the butterfly, Graphium sarpedon (Papilionidae: Lepidoptera). Proc. Japan Acad. Series B 57, 371-373.
    • (1981) Proc. Japan Acad. Series B , vol.57 , pp. 371-373
    • Maeki, K.1
  • 27
    • 0000962119 scopus 로고    scopus 로고
    • Absence of sex chromatin corresponds with a sex-chromosome univalent in females of Trichoptera
    • Marec, F. & Novák, K.; 1998: Absence of sex chromatin corresponds with a sex-chromosome univalent in females of Trichoptera. Eur. J. Entomol. 95, 197-209.
    • (1998) Eur. J. Entomol. , vol.95 , pp. 197-209
    • Marec, F.1    Novák, K.2
  • 28
    • 0000266040 scopus 로고
    • Contribution a l'analyse phylogénétique des néolépidoptères (Lepidoptera, Glossata)
    • Minet, J.; 1984: Contribution a l'analyse phylogénétique des néolépidoptères (Lepidoptera, Glossata). Nouv. Rev. Ent. (N.S.) 1, 139-149.
    • (1984) Nouv. Rev. Ent. (N.S.) , vol.1 , pp. 139-149
    • Minet, J.1
  • 29
    • 0002263677 scopus 로고
    • Primitive (non-ditrysian) Lepidoptera of the Andes: Diversity, distribution, biology & phylogenetic relationships
    • Lamas, G. (ed.), Second Symposium on Neotropical Lepidoptera, Arequipa
    • Nielsen, E. S., 1983: Primitive (non-ditrysian) Lepidoptera of the Andes: diversity, distribution, biology & phylogenetic relationships. In: Lamas, G. (ed.), Second Symposium on Neotropical Lepidoptera, Arequipa, Peru. J. Res. Lepid. (Suppl. 1), 1-16.
    • (1983) Peru. J. Res. Lepid. , Issue.SUPPL. 1 , pp. 1-16
    • Nielsen, E.S.1
  • 30
    • 0024467720 scopus 로고
    • Phylogeny of major lepidopteran groups
    • Fernholm, B.; Bremer, K.; Jörnvall, H. (eds), Amsterdam: Elsevier Sci. Publishers (Biomedic. Division)
    • Nielsen, E. S., 1989: Phylogeny of major lepidopteran groups. In: Fernholm, B.; Bremer, K.; Jörnvall, H. (eds), The Hierarchy of the Life. Amsterdam: Elsevier Sci. Publishers (Biomedic. Division), pp. 281-294.
    • (1989) The Hierarchy of the Life , pp. 281-294
    • Nielsen, E.S.1
  • 31
    • 0030318559 scopus 로고    scopus 로고
    • The Australian moth family Lophocoronidae and the basal phylogeny of the Lepidoptera-Glossata
    • Nielsen, E. S.; Kristensen, N. P., 1996: The Australian moth family Lophocoronidae and the basal phylogeny of the Lepidoptera-Glossata. Invertebr. Taxon. 10, 1199-1302.
    • (1996) Invertebr. Taxon. , vol.10 , pp. 1199-1302
    • Nielsen, E.S.1    Kristensen, N.P.2
  • 32
    • 84986409865 scopus 로고
    • Unusual sex chromosome inheritance in six species of small ermine moths (Yponomeuta, Yponomeutidae, Lepidoptera)
    • Nilsson, N.-O.; Löfstedt, C.; Därving, L., 1988: Unusual sex chromosome inheritance in six species of small ermine moths (Yponomeuta, Yponomeutidae, Lepidoptera). Hereditas 108, 259-265.
    • (1988) Hereditas , vol.108 , pp. 259-265
    • Nilsson, N.-O.1    Löfstedt, C.2    Därving, L.3
  • 33
    • 0008567787 scopus 로고
    • Karyologische Untersuchungen an Trichopteren
    • Pl. 3
    • Pchakadze, G. M., 1930: Karyologische Untersuchungen an Trichopteren. Arch. Russ. Anat. Histol. Embryol. 9, 227-237, 311-321. Pl. 3.
    • (1930) Arch. Russ. Anat. Histol. Embryol. , vol.9 , Issue.227-237 , pp. 311-321
    • Pchakadze, G.M.1
  • 34
    • 0008532669 scopus 로고
    • Somatic and meiotic analysis of Danaus limniace Cramer (Lepidoptera, Nymphalidae)
    • Rishi, S.; Rishi, K. K., 1985: Somatic and meiotic analysis of Danaus limniace Cramer (Lepidoptera, Nymphalidae). Chromosome Inf. Serv. 39, 22-23.
    • (1985) Chromosome Inf. Serv. , vol.39 , pp. 22-23
    • Rishi, S.1    Rishi, K.K.2
  • 36
    • 0003332515 scopus 로고
    • Das Verhalten der Geschlechschromosomen bei Lepidopteren, Nebst einem Beitrag zur Kenntnis der Eireifung, Samenreifung und Befruchtung
    • Seiler, J., 1914: Das Verhalten der Geschlechschromosomen bei Lepidopteren, Nebst einem Beitrag zur Kenntnis der Eireifung, Samenreifung und Befruchtung. Arch. Zellforsch. 13, 159-269.
    • (1914) Arch. Zellforsch. , vol.13 , pp. 159-269
    • Seiler, J.1
  • 37
    • 0008568573 scopus 로고
    • Die Zytologie und Geschlechsbestimmung der diploid parthenogenetischen Solenobia triquetrella
    • Seiler, J., 1953: Die Zytologie und Geschlechsbestimmung der diploid parthenogenetischen Solenobia triquetrella. Arch. Julius Klaus Stift. Vererb. Forsch. 24, 124-154.
    • (1953) Arch. Julius Klaus Stift. Vererb. Forsch. , vol.24 , pp. 124-154
    • Seiler, J.1
  • 38
    • 85012558281 scopus 로고    scopus 로고
    • Contribution to the knowledge of central Asiatic Microlepidoptera with descriptions of new species
    • Sruoga, V.; Puplesiene, J., 1998: Contribution to the knowledge of Central Asiatic Microlepidoptera with descriptions of new species. Acta Zool. Lituaniaca. Entomologia 8, 38-48.
    • (1998) Acta Zool. Lituaniaca. Entomologia , vol.8 , pp. 38-48
    • Sruoga, V.1    Puplesiene, J.2
  • 39
    • 0000086982 scopus 로고
    • Achiasmatische Oogenese bei Trichopteren
    • Suomalainen, E., 1966: Achiasmatische Oogenese bei Trichopteren. Chromosoma 18, 201-207.
    • (1966) Chromosoma , vol.18 , pp. 201-207
    • Suomalainen, E.1
  • 40
    • 0001062460 scopus 로고
    • On the sex chromosome trivalent in some Lepidoptera females
    • Suomalainen, E., 1969a: On the sex chromosome trivalent in some Lepidoptera females. Chromosoma 28, 298-308.
    • (1969) Chromosoma , vol.28 , pp. 298-308
    • Suomalainen, E.1
  • 41
    • 0001185040 scopus 로고
    • Chromosome evolution in the Lepidoptera
    • Suomalainen, E., 1969b: Chromosome evolution in the Lepidoptera. Chromosome Today 2, 132-138.
    • (1969) Chromosome Today , vol.2 , pp. 132-138
    • Suomalainen, E.1
  • 42
    • 0015162308 scopus 로고
    • Unequal sex chromosomes in a moth, Lozotaenia forsterana F. (Lepidoptera: Tortricidae)
    • Suomalainen, E., 1971: Unequal sex chromosomes in a moth, Lozotaenia forsterana F. (Lepidoptera: Tortricidae). Hereditas 68, 313-316.
    • (1971) Hereditas , vol.68 , pp. 313-316
    • Suomalainen, E.1
  • 44
    • 0002429089 scopus 로고
    • A genetic linkage study of W-chromosome-autosome fusions, breakage, and kinetic organization of chromosomes in Ephestia (Lepidoptera)
    • Traut, W., 1986: A genetic linkage study of W-chromosome-autosome fusions, breakage, and kinetic organization of chromosomes in Ephestia (Lepidoptera). Genetica 69, 69-79.
    • (1986) Genetica , vol.69 , pp. 69-79
    • Traut, W.1
  • 45
    • 0030167526 scopus 로고    scopus 로고
    • Sex chromatin in Lepidoptera
    • Traut, W.; Marec, F., 1996: Sex chromatin in Lepidoptera. Q Rev. Biol. 71, 239-256.
    • (1996) Q Rev. Biol. , vol.71 , pp. 239-256
    • Traut, W.1    Marec, F.2
  • 46
    • 0030768204 scopus 로고    scopus 로고
    • Sex chromosome differentiation in some species of Lepidoptera
    • Traut, W.; Marec, F., 1997: Sex chromosome differentiation in some species of Lepidoptera. Chromosome Res. 5, 283-291.
    • (1997) Chromosome Res. , vol.5 , pp. 283-291
    • Traut, W.1    Marec, F.2
  • 47
    • 0014350255 scopus 로고
    • Geschlechtschromatin bei Lepidopteren
    • Traut, W.; Mosbacher, G. C., 1968: Geschlechtschromatin bei Lepidopteren. Chromosoma 25, 343-356.
    • (1968) Chromosoma , vol.25 , pp. 343-356
    • Traut, W.1    Mosbacher, G.C.2
  • 48
    • 0032768893 scopus 로고    scopus 로고
    • Molecular differentiation of sex chromosomes probed by comparative
    • Traut, W.; Sahara, K.; Otto, T. D.; Marec, F., 1999: Molecular differentiation of sex chromosomes probed by comparative. Chromosoma 108, 173-180.
    • (1999) Chromosoma , vol.108 , pp. 173-180
    • Traut, W.1    Sahara, K.2    Otto, T.D.3    Marec, F.4
  • 49
    • 0008534219 scopus 로고
    • Goldtschmidt's theory of sex determination in Lymantria
    • Winge, Ö., 1937: Goldtschmidt's theory of sex determination in Lymantria. J. Genet. 36, 81-89.
    • (1937) J. Genet. , vol.36 , pp. 81-89
    • Winge, O.1


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