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We thank J. Denhart, J. N. Haas, S. Hick, R. L. Jefferies, J. H. McAndrews, W. McKenna, L. C. K. Shane, and L. J. Wang for field, laboratory, and general assistance, and L. C. Cwynar, E. Ito. K. Kelts, J. H. McAndrews, D. M. Peteet, L. C. K. Shane, T. Webb III, and H. E. Wright Jr. for discussion and comments. Supported by the Natural Sciences and Engineering Research Council of Canada, Royal Ontario Museum, Ontario Ministry of Education and Training (Ontario Graduate Scholarship), and University of Toronto (Open Fellowships and Department of Botany).
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